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THOMAS LINCOLN 
CASEY 


LIBRARY 
1925 


TRANSACTIONS 


OF THE 


ENTOMOLOGICAL SOCIETY 


OF 


LONDON. 


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THE 


TRANSACTIONS 


OF THE 


ENTOMOLOGICAL SOCIETY 


OF 


LONDON 


FOR THE YEAR 


1394. 


LONDON: 


PRINTED FOR THE SOCIETY BY SIMMONS AND BOTTEN, LIMITED, 
44, SHOE LANE, LONDON, E.C. 
SOLD AT THE SOCIETY’S ROOMS, 11, CHANDOS STREET, 
CAVENDISH SQUARE, W., 
AND BY LONGMANS, GREEN, AND CO., 
PATERNOSTER ROW, E.C.; AND NEW YORK. 


1894, 


Milner 
Of a De 


Oren 
ENTOMOLOGICAL SOCIETY OF LONDON. 


FOUNDED, 1833. 


INCORPORATED BY ROYAL CHARTER, 1885. 


OFFICERS and COUNCIL for the Session 1894-95, 


President, 
HENRY JOHN ELWHS, F.LS., F.Z.S. 


Vice- Presidents. 
Ture Rr. Hon. Lorp WALSINGHAM, M.A., LL.D., F.RB.S. 
Proressor EDWARD B. POULTON, M.A., F.R.S., F.LS. 
CoLonEL CHARLES SWINHOE, M.A., F.L.S., F.Z.S. 


Oreasurer. 
ROBERT McLACHLAN, F.B.S., F.LS. 


Secretaries. 


HERBERT GOSS, F.L:S. 
Tue Rey. Canon FOWLER, M.A., F.LS. 


Librarian. 
GEORGE C. CHAMPION, F.Z.S. 


Council. 


WALTER F. H. BLANDFORD, M.A., F.Z.S. 
GEORGE CHARLES CHAMPION, F.Z.S. 
HENRY JOHN ELWES, F.LS., F.Z.8. 

THE Rev. Canon FOWLER, M.A., F.LS. 
CHARLES JOSEPH GAHAN, M.A., F.ES. 
HERBERT GOSS, F.L.S., F.G.S. 

ROBERT McLACHLAN, F.R.5., F.L.S. 
FREDERIC MERRIFIELD, F.E.S. 

Pror. EDWARD B. POULTON, M.A., F.R.S. 
CoLoNEL CHARLES SWINHOE, M.A., F.L.S. 
GEORGE HENRY VERRALL, F.E:S. 
JAMES J. WALKER, R.N., F.L.S. 

Lorp WALSINGHAM, M.A., LL.D., F.R.S. 


Resident Librarian. 
W. R. HALL. 


THE 
TRANSACTIONS 
OF THE 
ENTOMOLOGICAL SOCIETY OF LONDON. 
1834—1894. 


The Transactions can now be obtained by Fellows 
at the following reduced prices :— 


PUBLIC. FELLOWS. 
First Series, 4 volumes (1834—1849) ............ Price £413 0 £310 OU 
Second Series, 5 volumes (1850—1861)......... coh DD aplonid 
Third Series, 5 volumes (1862—1869).........-+6 LUO 4.10 0 
‘ty he Transactions for the year 1868 O00 
55 aA 1869 iS ed i 
op 05 Toe Bis aye (U 
‘ : 1 5 0 | 
3 ; 120 
4 : 116 0 
3) ” u 12 0 
55 33 1) 52i0o- 3.0 0 
e . 112 0 | 
2? ” 1. 49 
if ss 100 015 0 
5 35 To 20 016 6 
if i 019 0 014 3 
” ” 116 0 Ae RO 
* ¥, 110 0 7226 
a er oR Ds 
ss - LOB isct oe 1.80 1 10 
ie i 7S a aa 1 60 019 6 
‘ “ Tee ea eae ae ; hed yt 019 6 
x x iT) ee . 146 019 0 
ie a TS an 115 0 163 
fs CRO ia tines sees 116 6 17 6 
¥ a TS00 1s sciotess 119 0 110 0 
Me 5 RSD A Penoseos 116 0 yD 
a i SOR ee: ; 1 Sane 1.29 
se sf Ce LG ae 1 ome 019 8 
paniepskeclese TOG. ib 24 ih 


First Series, vol. v., is out of print. First Series, vols. i.—iv., and 
Second Series, vol. iv., cannot be sold separately. 

The other volumes may be obtained separately, also the following :— 
Pascoe's ‘ Longicornia Malayand’ ..s.eesereee £212 0 £1 19/0 
Baly’s ‘ Phytophaga Malayana, Pt.1., Aposta- 

SUCOTOY vwsaah acoctajvsorseccoonssteseanaenssebensstiece 016 0 012 0 
Saunders’ ‘ British Heterogyna and Fossorial 

EDA INONOPLEN Gri aacdacecadecesevossetvecesdetteestes 0 4 6 03 4 
Saunders’ ‘ Synopsis of British Hymenoptera,’ 

att Lscccsaveus svoteedscmcs she teoeevaersonreeeamtee 060 0 4 6 
Newport's ‘ Athalia centifolie ’ (Prize Essay) O10 OF 140 


The JoURNAL OF PROCEEDINGS is bound up with the TRANSACTIONS. 


Fellows who have paid their Subscription for the current year, are 
entitled to receive the Transactions for the year without further payment, 
and they will be forwarded free, by post, to any address. 


(ule 


CONTENTS. 


Explanation of the Plates... ary re Ds ae nie eee 
Errata... Lf me a aa owe in ac Ses oo 
List of Fellows ... ee aes aR aH Vie eas ae aes 


Additions to the Tabrany ice ae ahs do ee 50 oe 


MEMOIRS. 


I. On a collection of Lepidoptera from ea Burma. Ey, 
Epwarp Meyrick, B.A., F.Z.S. 


II. Further observations on the Tea-bugs (Helopeltis) ae resin 
By CHarLEes OWEN WATERHOUSE, F.E.S. 


III. Notes on some Lepidoptera received from the peiskbouhaed 
of Alexandria. By Grorcr T. BretHunE-Baker, F.L.S. ... 


IV. The Rhynchophorous Coleoptera of Japan. Part LIT. 
Scolytidae. By Wattrer F. H. Buanprorn, M.A., F.Z.8. 


YV. Description of the female of Eran serony 
Butl. By Haminton H. Drucz, F.Z.S. a0 


VI. A list of the Lepidoptera of the Khasia Hills. Bart IOI By 
Colonel CHARLES SwinHogk, M.A., F.L.S., ete. ws: 


VII. An Entomological Temcaina to Corsica. By GroRGE C. 
CuaAmpPion, F.Z.S.... oft 


VUI. A list of the Hemiptera-Heteroptera agllapted “e Mr. 
Champion in Corsica in June, 1893, with a description of 
one new species. By Epwarp SAUNDERS, F.L.S. 


IX. On the Phylogeny of the Pierinw, as illustrated by fhe 
Wing-markings and Geographical Distribution. By Dr. 
Freperick A. Dixty, M.A., SLIBE F.E.S., Fellow of 
Wadham College, Oxford .. = 


X. Some notes on the Micro- hoes a eae tas are aa 
feeders, and chiefly on the early stages of Hriocephala cal- 
thella '(Lygenide, pineeedioe eae ee By Drs 
Tnomas A. CHapMan, M.D., 

XI. On the Tenebrionide collected in es aval Tasaunh oe 

Mr. James J. Watker, R.N., F.L.S., during the voyage of 
H.M.S. ‘ Penguin,” with descriptions of new ace and 
species. By Greorcre C. CuamPION, F.Z.8. ... = 

XII. Description of the Larva and Pupa of ps homem be. Fab. 
By Cuar.uzs B. Taytor, F.E.S. 

XIII. Notes on Dorydium (?) Westwoodi, Buchanan, White, with 
observations on the use of the name agen By WIL- 
LIAM F. Kirpy, F.L.8. ... 

XIV. Some new species of todos By fe Bae Caxos 
Fowuer, M.A., F.L.8., Sec. Ent. Soc. Lond. 20 aes 


( viii ) 


PAGE 
XV. Temperature Experiments in 1893 on several species of 
Vanessa and other Peper x FREDERIC MERRI- 

FIELD, F.E.S. 200 ocr renee 
XVI. Mr. Merrifield’s ixparinante in aniperatane: Wraristion as 


bearing on Theories of Heredity. By Dr. FrepErRIcK A. 
Drxey, M.A., M.D., F.E.S., Fellow of Wadham Ogleze, 


Oxford 508 365 439 
XVII. Descriptions of new Uae pene Mashunaland. By 
Louis Périnevry, F.E.S. Aa we 447 


XVIII. On Pyralidina from the ey Archipelago. By ‘Epwarp 
Meyrick, B.A., ¥.Z.8. .. 455 


XIX. Supplemental Tet of the Lienheohn Colsonters chtannee ig 
iMineseedeutells WALKER, R.N., F.L.8., during the voyage of 
H.M.S. “ Penguin,’’ under the ponynand of Captain Moorg, 

R.N. By Craruss J. Gawan, M.A., F.E.S. tise 481 


XX. Descriptions of a new species of Raph, L, and of fines 
new species of T'richoptera from the Balkan Peninsula, with 
critical remarks on , Panorpa gibberosa, McLach. ~ Pro- 
fessor FRANZ KLAPALEK, E.E.S. ave 489 
XXI. A Monograph of British Braconide. Part v2 By ite Bay. 
THomAS A. Marsuatt, M.A., F.H.S., Memberof the Société 
Entomologique de France .. : 497 
XXII. Catalogue of the Pter canoe: Tor feiediee ariel Tineide of 
the Madeira Islands, with notes and descriptions of new 
species. By the Right Honourable Lord WatsinecuaM, M.A., 
LL.D., F.B.S8. 535 


XXIII. Palearctic Nemoure. ‘By Tene Alp Wont E. E. S. pe aay 


XXIV. Supplementary Notes on the Scolytide of Japan, with a list of 
species. By Watrrr F. H. Buanprorp, M.A., F.Z.8.  ... 575 

XXV. Descriptions of the Pyralide, Crambide, and Phycide col- 
lected by the late T. Vernon Wollaston in Madeira. By 


Gercorcs T. BurauNne-BAkER, F.L.S. rece “tell, 
Proceedings for 1894... ee She 0 se oes a Pr 1 
President’s Address AAs Ss ies ios re és es $33 1 
Index oo & esi se re as ae iss ase lxxxv 


EXPLANATION OF THE PLATES. 


Plate I. See pages 383—51 Plate VIII. See pages 351—408 

elds See ,, 145—223 np IBS See ,, 425—438 

By ODES Ian cen WY 5p Ok See ,, 489—495 
See ,, 249—334 ty OLR gale 

VI. & VII. See ,, 497—531 


”» 


See ,, 335—350 », AIT. & XIV. 
See ,, 557—874 


Page 145 
» 146 
an 
” 154 
” 162 
(aalee 
164 
*, 166 
”” 168 
7 LOS 
* 169 
», 172, 220, etc. 
Lf 
a lAd, 
aye eS 
y 179 
Seren 
” 182 
cep 184 
” ” 
re deta) 
3 86 
” ” 
go 
so le 
rit bbs 
;, 198, 199 
loo 
” 9? 
,, 200 
204 
” oD 
”» 205 
” ” 
” ” 
», 208 
pare olie 
” 99 
ar ails 
ty 84 
ay 2b 
5, 9878 lines 4 and 5 


(ix ) 


ERRATA. 


Se 


Geometers 
Langlia 

pl. 3663 
hearsayt 
Epicoia 
Sunbury’s 
mimicusaria 
Menocteniide 
corta 

this wrong 
Herr Schiff 
veination 
seintiligera 
aventiara 
pulverentula 
Dithalma 
Brithrolophus 
Acidulidee 
Orgalivia 
antenne is 
hypospitata, 
aylinearia 
felicata 
exquisata 
calaminia 
Metoxidia 
Ismisea 
Maylayan 
Erinnis 
chromotaria 
forewing 
hemearia 
pairs spurs 
then 
antennz has 
Ath 

Calcula 
antenne is 
Amphidaris 
bengalaria 
Opthalmodes 
limacoides 


read 


Geometrina 
Langia 
pl. 36, fig. 3 
hearseyt 
Epicopeia 
Swinhoe’s 
numicusaria 
Monoctendiiz 
costa 
this is wrong 
Herr Schatt 
venation 
scutiligera 
aventiaria 
pulverulenta 
Dithalama 
Erythrolophus 
Acidalidz 
Orgalima 
antenne are 
hypospilata 
axylinaria 
filicata 
exquisita 
calamina 
Metowydia 
Isniscw 
Malayan 
Brinvys 
chromataria 
hindwing. 
hermearia 
pairs of spurs 
than. 
have. 
Atk. 
Culeula 
are 
Anvphidasys 
bengaliaria 
Ophthalmodes 
limacella 


——* 
‘a 
~_ 


Hist of Fellotus 


OF TILE 


ENTOMOLOGICAL SOCIETY OF LONDON. 


Date of HONORARY FELLOWS. 


Election. 

1894 Foret, Professor August, M.D., The University, Ziirich. 
1884 Mixer, Dr. Fritz, Blumenau, Santa Catarina, Brazil. 

1884 OsTEN SACKEN, Baron C. R. von, Wideplatz, Heidelberg. 
1884 Packarp, Dr, Alpheus S., Providence, Rhode Island, U.S.A. 


1889 Ru.e¥y, Prof. Charles V., U.S. National Museum, Washington, D.C., 
U.S.A. 


1872 Saussure, Henri F. de, Tertasse, 2, Geneva. 

1871 Srtys-Lonecuamps, Baron M. E. de, Liége. 

1885 SNELLEN, Pieter Carl T., Rotterdam. 

1893. WarrenwyL, Hofrath Dr. Carl Brunner Von, Thereseinamagasse, 
25, Vienna. 


FELLOWS. 
Marked * are Original Members. 
Marked + have compounded for their Annual Subscriptions. 


Date of 
Election. 


1877. Apams, Frederick Charlstrom, 68, St. Ermin’s Mansions, Caxton- 
street, Westminster, S.W. 

1877. Apbams, Herbert J., Roseneath, London-road, Enfield, N. 

1885 ApkIN, Robert, Wellfield, Lingard-road, Lewisham, S.E. 

1891 Apyer, J. M., Brockenhurst, Lymington, Hants. 

1856 ARMITAGE, Edward, R.A., 3, Hall-road, St. John’s Wood, N.W. 

1886 ArMmorE, E. A., 3, Haylett-terrace, Exton’s-road, King’s Lynn 
Norfolk. 

* + Bapincron, Professor Charles Cardale, M.A., F.R.S., F.L.S., &c. 

Professor of Botany in the University of Cambridge, 5, Brook- 
side, Cambridge. 


LIS! OF FELLOWS. X1 


Batty, William Edward, Lyuwood Mouse, Paul Churchtown, near 
Penzance, Cornwall. 

Baker, Walter F., Trent House, Gainsborough. 

Bankes, Eustace R., M.A., The Rectory, Corfe Custle, Wareham, 
Dorset. 

Barciay, Francis H., F.G.S., Knott’s Green, Leyton, Hssex. 

BarGaGuLt, Nobile Cavaliere Piero, Piazza S. Maria, Palazzo 
Tempi No. 1, Florence, Italy. 

BarkeEr, H. W., 147, Gordon-road, Peckham, S.E. 

Barrett, Charles Golding, Inland Revenue Department, Somerset 
House, W.C. ; and 39, Linden-grove, Nunhead, 8.E. 

Barton, Stephen, 114, St. Michael’s Hill, Bristol. 

Bateson, William, M.A., F.R.S., Fellow of St. John’s College, 
St. John's College, Cambridge. 


1851 +t Beaumont, Alfred, The Red Cottage, Pond Road, Blackheath, S.E. 


1893 
1891 
1882 
1885 
1892 
1886 
1880 


1879 
1891 


BepparpD, Frank E., M.A., F.R.S., Zoological Gardens, Regent's 
Park, N.W. 

BEECHING, Robert A. Dallas, 24, St. James-road, Tunbridge Wells, 
Kent. 

Bere, Prof. Dr. Carlos, Director del Museo Nacional, Buenos 
Aires (Rep. Argent.), South America. 

BeruHune-BaAkER, George T., F.L.S., 19, Clarendon-road, Edgbaston, 
Birmingham. 

BIDvELL, Walter Cuthbert, 32, The Grove, Bolton Gardens, S.W. 

BippLE, F. W., M.A., Lanherne, Albemarle-road, Beckenham, Kent. 

BIGNELL, George Carter, 7, Clarence-place, Stonehouse, Plymouth. 

Biuuvrs, T. R., 20, Swiss Villas, Coplestone-road, Peckham, S.E. 

Biaser, W. H., F.L.S., 34, Cromwell-road, West Brighton. 


1894 + BLACKBURNE-Maze, W. P., Shaw House, Newbury, Berkshire. 


1889 
1890 
1885 
1886 
1876 
1875 


1876 
1891 


1892 
1888 


BLanprorD, Walter F. H., M.A., F.Z.S., 48, Wimpole-street, W. 

Buatcu, W. G., Knowle, near Birmingham. 

Buiatuway?, Lieut.-Col. Linley, F.L.S , Hagle House, Batheaston, 
Bath. 

BLoomriELtp, The Rev. Edwin Newson, M.A., G'uestling Rectory, 
fTastings. 

Borre, Alfred Preudhomme de, Villa la Faucette, Petit Saconnen, 
Geneva. 

Borrer, Wm., junr., F.G.S., Pakyns Manor House, Hurstpierpoint, 
Sussex, 

BoscuHer, Edward, Bellevue House, Twickenham. 

Booru, George A., Fern Hill, Grange-over-Sands, Carnforth, 
Lancashire. 

BovuskELL, Frank, 11, Lansdowne-road, Stoney Gate, Leicester. 

Bowen, B.A., Langley, Elitham-road, Lee, S.E. 


1894+ Bowes, E. Augustus, M.A., Myddelton House, Wultham Cross, 


Hertfordshire, 


xii 


LIST OF FELLOWS. 


1852 + Boyp, Thos., Woodvale Lodge, South Norwood Hill, 8.E. 


1893 


1894 


1877 


1870 
1894 
1890 
1893 
1879 


1878 
1887 
1886 
1892 


1890 
18835 


1889 


Brawsant, Edouard, Chateau de Morenchies, par Cambrai (Nord), 
France. 

Breyer, Professor H. G., M.D., Gymnasium, Pretoria, Transvaal, 
Africa, 

Briaas, Charles Adolphus, 55, Lincoln’s Inn Fields, W.C. ; and 
Surrey House, Leatherhead, Surrey. 

Briaes, Thomas Henry, M.A., Surrey House, Leatherhead, Surrey. 

Bricut, Percy M., Roccabruna, Bournemouth. 

BristoweE, B. A., Durlstone, Champion Hill, 8.B. 

BromiLow, Frank, Selborne, Poole road, Bournemouth. 

Broneniart, Le Chevalier Charles, Assistant d’Entomologie au 
Muséum WVhistoire naturelle de Paris, Memb. Ent. Soc. France, 
and Memb. Geol. Soc. France, Foreign Corr. Geol. Soc. Lond., 
&e., 9, Rue Linné, Paris. 

Brown, Capt. Thomas Drury, Auckland, New Zealand. 

Brown, Henry Rowland, M.A., 3, Pump-court, Temple, B.C. 

Brown, John, 5, King’s Parade, Cambridge. 

Browne, Capt. Clement Alfred Righy, R.E., cjo Grindlay & Co., 
55, Parliament-street, Westminster, S.W. 

Bryant, George, Somerset Lodge, Old Shirley, near Southampton. 

Buck ton, George Bowdler, F.R.S., F.L.8., Weycombe, Haslemere, 
Surrey. 


Burns, Henry, The Free Public Library, Fulham, 8.W. 


1868 + BuTLeR, Arthur Gardiner, Ph.D., F.L.S., F.Z.8., British Museum, 


1883 
1886 
1886 
1885 
1860 
1880 
1889 
1890 


1886 
1894 


1886 


South Kensington, 8.W.; and The Lilies, Penge-road, Beckenham, 
Kent. 
But er, Edward Albert, B.A., B.Sc.,39, A shley-road, Crouch Hill, N. 


CALVERT, Wm. Barilett, [iceo de Quillota, Quillota, Chili, South 
America. 

CAMERON, Peter, The Rookery, Bridgemont, Whaley Bridge, 
Cheshire. 

CAMPBELL, Francis Maule, F.L.S., F.Z.8., &c., Rose Hill, Hoddes- 
don, Herts. 

CaNnbDbzE, Dr. E., Glain, Liége. 

CANSDALE, W. D., Sunny Bank, South Norwood, S.E. 

Cant, A., c/o Fredk. Du Cane Godman, Esq., F.R.S., 10 Chandos- 
street, Cavendish-square, W. 

CAPPER, Samuel James, F.L.S. (President of the Lancashire and 
Cheshire Entomological Society), Huyton Park, near Liverpool. 

Capron, Edward, M.D., Shere, Guildford, Surrey. 

CaraccioLo, H., H.M. Customs, Port of Spain, Trinidad, British 
West Indies. 

CARMICHAEL, Sir Thomas David Gibson, Bart., M.A., F.LS., 
Castlecraig, Dolphinton, N.B. 


LIST OF FELLOWS. Xill 


1892 CarrEenter, The Honble. Mrs. Beatrice, Kiplin, Northallerten, 
Yorkshire. 

1868 CarRrINnGTon, Charles, Carylls, Fay Gate, Sussex. 

1890 Carrer, George Wm., M.A., F.L.S.. Clif End House, Scarboro’. 

1889 + Cave, Charles, 13, Lowndes-square, S.W. 

1871 Campion, George C., F.Z.S., Liprartan, Heatherside, Horsell, 
Woking, Surrey ; and 10, Chandos-street, Cavendish-square, W. 

1891 COHapman, Thomas Algernon, M.D., Fir Bank, Hereford. 

1890 CuarrERTON, Frederick J.S., 78, Clissold Road, Stoke Newington, 
Ne 

1891+ Currry, Arthur J., M.A., 27, Hereford-square, S.W. 

1889 Curisty, W. M., M.A., Watergate, Emsworth, Sussex. 

1886 + Ciark, John Adolphus, The Broadway, London Fields, N.E. 

1867 CrarKE, Alex. Henry, 109, Warwich.road, Earl’s Court, 8.W. 

1886 CLARKE, Charles Baron, M.A., F.R.S., President L.S., F.G.S., 
13, Kew Gardens-road, Kew, S.W. 

1891 CrLarkE, Henry Shortridge, 2, Osborne-terrace, Douglas, Isle of Man. 

1891 CockERELL, Theodore D. A., F.Z.S., Las Cruces, New Mexico, 
CS A 

1874 CockiE, Major George, M.A., B.Mus., Oxon., 9, Bolton-gardens, 
S.W. 

1873 Cour, William, 7, Knighton Villas, Buckhurst Hill, Essex. 

1880 CopLaNnD, Patrick F., 2, Hope Villas, Buckhurst Hill, Essex. 

1894 Corrs, E. C., The Indian Museum, Calcutta. 

1892 Cowan, Thomas William, F.L.S., F.G.S., F.R.M.S., 31, Belsize 
Park Gardens, Hampstead, N.W., and Penleaze, Fowey, Cornwall. 

1886 CoweELL, Peter (Librarian of the Liverpool Free Public Library), 
William Brown-street, Liverpool. 

1867 Cox, Herbert Ed., c/o F. 8. Eve, Esq., 125, Harley-street, W. 

1888 Crecosn, J. P., P.O. Bow 1420, Johannesburg, South Africa. 

1890 Crewe, Sir Vauncey Harpur, Bart., Caike Abbey, Derbyshire. 

1880 +Crisp, Frank, LL.B., B.A., Treasurer L.S., F.G.S., Treasurer 
R.M.S., 5, Lansdowne-road, Notting Hill, W. 

1888 Croker, A. J., 90, Albert-road, Walthamstow. 

1883 CrowLey, Philip, F.L.S., F.Z.8., Waddon House, Croydon. 


1873 Date, C. W., Glanville’s Wootton, Sherborne, Dorset. 

1887 Dattry, The Rev. Thomas W., M.A., F.L.8., Madeley Vicarage, 
Newcastle, Staffordshire. 

1886 Dannatt, Walter, F.Z.S., Ivy Dene, Westcombe Park, Blackheath, 
S.E. 

1892 +DENNIS, George Christopher, 39, Blossom-street, York. 

1885 Denv, Hastings Charles, C.E., F.L.S., 20, Thurloe-square, S.W. 

1886 Dickson, The Rev. Prof. William Purdie, D.D., LL.D., Professor 
of Divinity in the University of Glasgow, Glasgow. 


X1V 


1875 
1887 


1891 
1885 


1873 
1886 


1845 
1889 
1874 
1884 
1867 


1894 


LIST OF FELLOWS. 


Distant, Wm. Lucas, Bow 352, Pretoria, Transvaal, South Africa. 

Drixey, Frederick Augustus, M.A., M.D., Fellow and Bursar of 
Wadham College, Wadham College, Oxford. 

DoNIsTHORPE, Horace St. John K., 73, West Cromwell-road, 8.W. 

Donovay, Surg.-Captain Charles, M.D., 1st Burma Rifles, Fort 
Dufferin, Mandalay, Burma. 

Doria, Marquis Giacomo, Strada Nuova, Genova. 

Dormer, The Right Honourable Lord, Cox’s Hotel, Jermyn-street, 
S.W. 

Dovueias, John Wm., Dartmouth Lodge, 153, Lewisham - road, 
Lewisham, 8.E. 

DowninG, John W., 59, Lupus-street, St. George’s-square, S.W. 

DowsetT, Arthur, Castle Hill House, Reading. 

Druce, Hamilton H. C. J., F.Z.8., 43, Circus-road, St. John’s 
Wood, N.W. 

Druce, Herbert, F.L.S., F.Z.S., 43, Circus-road, St. John’s Wood, 
NAW f 

DunpGEon, G. C., 58, Wontagu-square, W. 


1849 }DUNNING, Joseph Wm., M.A., F.L.8., F.Z.8., 4, Talbot-square, 


1883 


1890 
1865 


1886 
1884 


1886 
1886 
1878 


1886 


1890 


1892 
1861 
1886 


1881 
1889 


Paddington, W. 
Durrant, John Hartley, The Cottage, Merton Hall, Thetford, 
Norfolk. 


Eastwoop, John Edmund, Exton Lodge, Witley, Surrey. 

Eaton, The Rev. Alfred Edwin, M.A., care of R.S. Eaton, Esq., 
4, Belfield-terrace, Weymouth, Dorset. 

Epwarps, James, Colesborne, Andoversford, R.S.O., Gloucestershire. 

Epwarps, Stanley, F.L.8., F.Z.8., Kidbrook-lodge, Blackheath, 
S.E. 

Exisua, George, 122, Shepherdess-walk, City-road, N. 

Euis, Joho W., M.B., L.R.C.P., 18, Rodney-street, Liverpool. 

Ewes, Henry John, J.P., F.L.S., F.Z.S., Vice-PResIpENT, 
Colesborne, Andoversford, R.S.O., Gloucestershire. 

Enock, Frederick, F.L.S., 21, Manor-gardens, Holloway, N. 


Farn, Albert Brydges, fount Nod, Greenhithe, Kent; and 
Medical Department, Local Government Board, Whitehall, 8.W. 

Farren, William, Fern House, Union-road, Cambridge. 

FENN, Charles, Eversden House, Burnt Ash Hill, Lee, S.E. 

Fenwick, Nicholas Percival, Holmwood, South Bank, Surbiton 
Hill, Surrey. 

FEREDAY, R. W., Christchurch, Canterbury, New Zealand. 

FERNALD, Prof, C. H., Amherst, Mass., U.S.A. 


1878 


1874 
1886 
1865 


LIST OF FELLOWS. XV 


Finzt, John A., Hanover Lodg2, 77, St. Helen’s-gardens, N., 
Kensington, W. 

Fircu, Edward A., F.L.8., Brick House, Maldon, Essex. 

Fircu, Frederick, Hudleigh House, Highbury New Park, N. 

FLercuer, J. E., 2, Bedwardine-road, St. Johns, Worcester. 


1883+ FLETCHER, William Holland B., M.A., Fairlawn, Worthing, 


1892 
1885 
1880 


1883 
1888 


1891 
1855 


1889 
1884 


1887 


1887 
1892 


1890 
1893 


1865 


1890 
1886 
1855 
1874 
1886 
1891 


1894 
1865 


Sussex. 
FLevuTIAux, Edmond, 1, Rue Malus, Paris. 
Foxkerr, A. J, F., Zierikzee, Zeeland, Netherlands. 
Fow1er, The Rev. Canon, M.A., F.L.5., Secrerary, The School 
House, Lincoln. ; 
FREEMAN, Francis Ford. A bbotsfield, Tavistock, South Devon. 
FreMLIN, H. Stuart, M.R.C.S., L.R.C.P., Mereworth, Maidstone, 
Kent. 
Frouawk, F. W., 39, Dornton-road, Balham, 8.W. 


Fry, Alexander, F.U.5., Zhornhill House, Dulwich Wood Park, 
Norwood, S.E. 


Fryer, Charles John, 410, Wandsworth-road, 8.W. 
Fuuuer, The Rev. Alfred, M.A., The Lodge, 7, Sydenham-hill, 
Sydenham, 8.E. 


GAHAN, Charles Joseph, M.A., British Museum (Natural History), 
South Kensington, S.W.; and 16, Ashchurch-grove, Shepherd's 
Bush, W. 

Seen Francis, M.A., D.C.L., F.R.S., F.G.S., 42, Rutland Gate, 

GaRDE, Philip de la, R.N., H.M.S. “Raleigh,” c/o Miss Purkis, 
12, Eversfield-road, St. Leonard’ s-on-Sea. 

GARDNER, John, 6, Friars-gate, Hartlepool. 

Gipps, Arthur Ernest, F.L.8., Avenue House, St. Albans, Hert- 
fordshire. 

+Gopman, Frederick Du Cane, F.R.S., F.L.S., F.Z.S., South Lodge, 
Lower Beeding, Horsham, Sussex ; 7, Carlos-place, Grosvenor- 
square ; and 10, Chandos-street, Cavendish-square, W. 

GOLDTHWAITE, Oliver, 3, Duke of Edinburgh-road, Carshalton, 

Surrey. 
+Goopricu, Captain Arthur Mainwaring, Aubrey, Lymington, 
Hants. 

GorHam, Rev. Henry Stephen, F.Z.S., The Chestnuts, Shirley 
Warren, Southampton. 

Goss, Herbert, F.L.S., F.G.S., Secretary, The Avenue, 
Surbiton-hill, Surrey; and 11, Chandos-street, Cavendish- 
square, W. 

GREEN, A. P., Colombo, Ceylon. 

GREEN, E. Ernest, Eton Estate, Punduloya, Ceylon. 

GREEN, Joseph F., West Lodge, Blackheath, S.K. 

GREENE, The Rev. Joseph, M.A., Rostrevor, Clifton, Bristol. 


Xvi LIST OF FELLOWS. 


1893+ GREENWOOD, Henry Powis, F.L.S., Hornham Cliff, near 
Salisbury. 

1888 Gruirritus, G. C., 43, Caledonian-place, Clifton, Bristol. 

1894 GrimsHaw, Percy H., Natural History Department, Edinburgh 
Museum of Science and Art, Edinburgh. 


1893 Hatrorp, Frederick Michael, 6, Pembridge-place, W. 

1890 + Hauu, A. E., Norbury, Pitsmoor, Sheffield. 

_ 1885 Hau, Thomas William, “ Stanhope,” The Crescent, Croydon. 

1891 Hampson, G. F., B.A., 36, Tedworth-square, Chelsea, S.W. 

1891 Hanpury, Frederick J., F.L.S., 69, Clapton Common, Clapton, N.E. 

1891 Hanson, R. E. Vernon, B.A., c/o Glen Coats, Esq., Ferguslie Park, 
Paisley, N.B. 

1877 HarpineG, George, The Grove, Fishponds, Bristol. 

1889 Harrison, John, 7, Gawber-road, Barnsley, Yorkshire. 

1892 Heapty, Charles Burnard, Stoneygate-road, Leicester. 

1892 Heatu, Edward Alfred, M.D., F.L.S., 114, Ebury-street, Pimlico, 
S.W. 

1889 HeENN, Arnold Umfreville, Box 1282, Post Office, Sydney, N.S. W. 

1881 Henry, George, 38, Wellington-square, Hastings. 

1895 Hippert, Charles R. C., Sefton Purk, Slough, Bucks. 

1888 Huices, Martin Stanger, F.C.S., Clarence House, Russell-street, 
Gloucester. 

1891 Huiuu, Henry A., 4, Rosslyn-gardens, Hampstead, N.W. 

1876 + HiLLMAN, Thomas Stanton, Hastgate-street, Lewes. 

1890 HopeKinson, J. B., Roseberry House, Powys-road, Ashton-on- 
Ribble. 

1888 Hopson, The Rev. J. H., B.A., 70, Raincliffe-road, Walton, 
Liverpool. 

1887 Ho.uanp, The Rev. W. J., D.D., Ph.D., 5th Avenue, Pittsburg, 
Penn., U.S.A. 

1876 t Horniman, Fredk. John, F.L.S., F.Z.8., F.R.G.S., &e., Surrey 
Mount, Forest Hill, S.E. 

1892 Hoyue, Samuel, Audley House, Sale, Cheshire. 

1865 + Hupp, A. E., “ Clinton,” Pembroke-road, Clifton, Bristol. 

1888 Hupson, George Vernon, The Post Office, Wellington, New 
Zealand. 


1880+ INcuBALD, Peter, F.L.S., F.Z.8., Grosvenor-terrace, Hornsea, 
Holderness. 

1893 Irby, Lieutenant-Colonel Leonard Howard Loyd, F.L.S., F.Z.S., 
14, Cornwall-terrace, Regent’s Park, N.W. 

1891 IsapeuL, The Rev. John, 65, Waddon Old-road, Croydon. 


1886 Jacosy, Martin, 7, Hemstall-road, West Hampstead, N.W. 


LIST OF FELLOWS. Xvi 


1892 Jarrrey, Francis, M.R.C.S., L.R.C.P., 8, Queen's Ride, Burnes, 
S.W. 

1869 Janson, Oliver E., Perth-road, Stroud Green, N.; and 44, Great 
Russell-street, Bloomsbury, W.C. 

1886 JENNER, James Herbert Augustus, 4, Husi-strect, Lewes. 

1886 Joun, Evan, Llantrissant, Pontypridd, Glamorganshire. 

1889 Jounson, The Rev. W.F., M.A., Winder-terrace, Armagh, Ireland. 

1888 Jones, Albert H., Shrublands, Eltham, Kent. 

1894 Jones, Frederic Whitworth, “ Sherwood,” Setlagoli, British 
Bechuanaland, Africa. 

1894 Jorpay, Dr. K., The Museum, Tring, Hertfordshire. 


1884 Kane, W. F. de Vismes, M.A., M.R.LA., Sloperton Lodge, Kings- 
town, Ireland. 

1884 Kappe, A. W., F.L.S., 5, Burlington-gardens, Chiswick, W. 

1876 + Kay, John Dunning, Leeds. 

1884 Keays, F. Lovell, F.L.S., 26, Charles-street, St. James's, S.W. 

1894 Kerrsie, Henry, 10, Coleman-street, E.C. 

1890 Kenrick, G. H., Whetstone, Somerset-road, Edgbaston, Bir- 
mingham. 

1890 Kimper, Miss M., Cope Hall, Enborne, Newbury, Berks. 

1899 Kine, J. J. F. X., 207, Sauchiehall-street, Glasgow. 

1861 Krirpey, William F., F.L.S., 5, Burlington-gardens, Chiswick, W. 

1893 Krrkaupy, George Willis, St. Abbs, Worple-road, Wimbledon, 
S.W. 

1889 KULAPALEK, Professor Franz, Trebon, Wittingau, Bohemia. 

1887 + KEIN, Sydney T., F.L.S., F.R.A.S. (Hon. Treasurer, Middlesex 
Natural History and Science Society), The Red House, Stanmore, 
Middlesex. 

1876 Kraatz, Dr. G., 28, Link-strasse, Berlin. 


1868 Lana, Colonel A.M.,R.E., 31, Shooter's Hill-road, Blackheath, 8.E. 

1887 + Leecu, John Henry, B.A., F.L.S., F.Z.5., F.R.G.S., &c., Vachery, 
Cranleigh, Surrey. 

1883 Lemann, Fredk. Charles, Blackfriars House, Plymouth. 

1892 Lesuin, J. H., 58, Forbourne-road, Upper Tooting, 8.W. 

1876 Lewis, George, F.L.S., St. Regulus, Archer’s-road, Southampton. 

1892 Licurroot, R. M., Bree-st., Cape Town, Cape of Good Hope. 

1886 Livett, H. W., M.D., Wells, Somerset. 

1865 t LLEWELYN, Sir J. Talbot Dillwyn, Bart., M.A., F.L.S., Penller- 
gare, Swansea. 

1881 + Luoyp, Alfred, F.C.S., The Dome, Bognor, Sussex. 

1885 + Luoyp, Robert Wylie, St. Cuthberts, Thurleigh-road, Nightingale- 
lane, Clapham Common, 8.W. 

b 


XVill LIST OF FELLOWS. 


1894 Lowkr, The Rev. Frank E., M.A., St. Stephen’s Vicarage, 
Guernsey. 

1850 Lown, W. H., M.D., Woodcote Lodge, Inner Park-road, Wimbledon 
Park, 8.W. 

1893 Lower, Oswald B., Bleak House, Park Side, Adelaide, South 
Australia. 

1850+ Lupnock, The Right Honble. Sir John, Bart., M.P., D.C.L., 
F.R.S., F.LS., F.G.S., etc., High Elms, Farnborough, 
Kent. 

1880 Lupron, Henry, Lyndhurst, North Grange-road, Headingley, 
Leeds, 


1887 M‘DovcatL, James Thomas, Dunolly, Morden-road, Blackheath, 
S.E. 

1851 + M‘Inrosu, J. 

1888 Mackinnon, P. W., Lynndale, Mussoorie, N.W.P., India. 

1892 Mackonocuir, The Rev. J. A., B.A. (Chaplain to the Earl 
of Home), Douglas Castle, Lunarkshire; and The Hirsel, 
Coldstream. 

1858 McLacnuan, Robert, F.R.S., F.L.S., F.Z.S., Treasurer, West- 
view, 23, Clarendon-road, Lewisham, 8.E. 

1887 Manpers, Surgeon-Captain Neville, L.R.C.P., M.R.C.S., Medical 
Staff Mess, Aldershot. 

1891 Mancer, William T., 100, Manor-road, Brockley, S.E. 

1892 ManspripGs, William, 21, Rosenau-crescent, Battersea Park, 8.W. 

1894 + Marsuatt, Alick, Auchinraith, Bexley, Kent. 

1865 Marsuaut, The Rev. Thos. Ansell, M.A., Botusfleming Rectory, 
Hatt, Cornwall. 

1856+ Marsuaur, William, Auchinraith, Bexley, Kent. 

1874+ Mason, Philip Brookes, M.R.C.S., P.LS., Trent House, Burton- 
on- Trent. 

1865 Maruew, Gervase F., R.N., F.L.S., F.Z.S., F.R.G.S., Lee House, 
Dovercourt, Essex. 

1887 Marruews, Coryndon, Plympton St. Mary, South Devon, 

1860 May, John William, K.N.L., Blenheim House, Parson’s Green- 
lane, Fulham, S.W. 

1872 +t MeLDoLa, Professor Raphael, F.R.S., F.C.S., PRESIDENT, 6, Bruns- 
wick-square, W.C. 

1885 MELvILL, James Cosmo, M.A., F.L.S., Brook House, Prestwich, 
Lancashire. 

1887 MERRIFIELD, Frederic, 24, Vernon-terrace, Brighton. 

1888 Mryer-Danrcis, G., c/o Sogin & Meyer, Wohlea, Switzerland. 

1880 Meyrick, Edward, B.A., F.Z.S., Ramsbury, Hungerford, Berk- 


shire. 


LIST OF FELLOWS. X1X 


1894 Mratu, Professor Lewis Compton, F.R.S., Crag Foot, Ben Rhydding, 
Leeds. 

1883 Mies, W. H., The New Club, Calcutta. 

1879 Monrerro, Senhor Antonio Augusto de Carvalho, 72, lua de 
Alecreon, Lisbon. 

1853 Moors, Frederic, D.Sc., A.L.S., F.Z.S., Claremont House, Avenue- 
road, Croydon-road, Penge, 8.E. 

1886 Morean, A.C. F., F.L.S., 24, Leinster-square, W. 

1889 +t Morice, The Rev. F. D., M.A., Fellow of Queen’s College, Oxford, 
27, Hillmorton-road, Rugby. 

1893 Morton, Kenneth J., Glenview Cottage, Carluke, N.B. 

1889 Mostey, 8. L., Beaumont Park, Huddersfield. 

1869 + Miuuer, Albert, F.R.G.S. 

1872 t Murray, Lieut.-Colonel H., 43, Cromwell Houses, Cromweill- 
road, S.W. 

1886 Muorcn, J. P., 359, Hornsey-road, N. 


1889 Nevrnson, Basil George, M.A., F.Z.8., 3, Yedworth-square, 
Chelsea, S.W. 

1887 Newman, The Rev. W. J. H., M.A., Uhe Vicarage, Steeple 

Barton, Oxon. 

1878 Newman, Thomas P., 54, Hatton-garden, H.C.; and Hazelhurst, 
Haslemere, Surrey. 

1890 NeEwstTEAD, R., The Museum, Chester. 

1882 NICEVILLE, Lionel de, F.L.S., C.M.Z.S., Indian Museum ; and 13, 
Kyd-street, Calcutta. 

1886 NicHoxson, William E., School Hill, Lewes, Sussex. 

1893 NOoNFRIED, A. F., Rakonitz, Bohemia. 

1886 Norris, Robert E., 15, Market-place, Cirencester. 

1878 NorripGe, Thomas, Ashford, Kent. 


1869 OBEeRtTHUR, Charles, Rennes, France. 

1877 OxBerTHUR, René, Rennes, Vrance. 

1893 + OcLE, Bertram 8., Steeple Aston, Oxfordshire. 

1883 OLDFIELD, George W., M.A., F.L.S., F.Z.S., 21, Longridye-road, 
Earl's Court, S.W. 

1893 OLtverR, John Baxter, 12, dAvenue-road, St. John’s Wood, 
NEW: 

1873 OLtviER, Ernest, Iamillons, prés Moulins (Allier), France. 

1886 Oxtirr, Arthur Sidney, Government Entomologist, Department of 
Agriculture, Macquarie-street, Sydney, N. S. Wales. 


xx LIST OF FELLOWS. 


1878 OnrmeERop, Miss Eleanor A., F.R.Met.S., Torrington House, Holy- 
well Hill, St. Albans, Herts. 

1880 ORrmEROD, Miss Georgiana, Torrington House, Holywell Hill, 
St. Albans, Herts. 


1893 PauLcKE, Wilhelm, 33, Langstrasse, Baden-Baden, Germany. 

1888 PENNINGTON, F., jun., 7, Park-place, St. James’s, 8.W. 

1883 PrRinGuEY, Louis, South African Museum, Cape Town, South 
Africa. 

1879 PERKINS, Vincent Robt., Wotton-under-Hdge, Gloucestershire. 

1887 Puriiips, Charles Edmund Stanley, Castle House, Shooter's 
Hili, Kent. : 

1891 Purerce, Frank Nelson, 7, The Elms, Dingle, Liverpool. 

1885 Pout, J. R. H. Neerwort van de, Heerengracht 476, Amsterdam. 

1870 + Porritt, Geo. T., F.L.8., Crosland Hall, Huddersfield. 

1884+ PouLtTon, Professor Edward B., M.A., F.R.S., F.L.S., F.G.S., 
F.Z.S., VicE-PRESIDENT, Hope Professor of Zoology in 
the University of Oxford, Wykeham House, Banbury-road 
Oxford. 

1894 Pratt, John, The Cedars, New Barnet, Hertfordshire. 

1851 Preston, The Rev. Thomas Arthur, M.A., F.L.8., Thurcaston 
Rectory, Leicester. 

1878 Prick, David, 48, West-street, Horsham, Sussex. 

1893 Prout, Louis Beethoven, 12, Greenwood-road, Dalston, N.E. 


1887 Raconot, E. L. (Ex-President Entom. Soc. France), 12, Quai de la 
Rapée, Paris. 

1882 + RamspEN, Hildebrand, M.A., F.L.S., 26, Upper Bedford-place, 
Russell-square, W.C. 

1874 ReeEp, Edwyn C., C.M.Z.S., Banos de Cauquenes, Chili. 

1893 Reip, Captain Savile G., late R.E., Thornhaugh, Swanage, 
Dorset. . 

1891 ReEr, William, Pitcaple, Aberdeenshire. 

1890 RenpLesuAm, The Right Honble. Lord, Rendlesham Hall, Wood- 
bridge, Suffolk. 

1886 Ruopes, John, 360, Blackburn-road, Accrington, Lancashere. 

1891 Ricnarpson, Nelson M., B.A., Monte Video, near Weymouth, 
Dorset. 

1894 Ripine, William Steer, B.A., M.D., Buckerell Lodge, Buckerell, 
near Honiton, Devon. 

1853 Ripon, The Most Honourable the Marquis of, K.G., D.C.L., 
F.R.S., F.L.S., ete., 9, Chelsea Embankment, 8.W. 

1889 Rosinson, Arthur, B.A., 1, Mitre Court Buildings, Temple, B.C. 

1892 Rosinson, Sydney C., Goldsmith’s Hall, E.C. 

1869 + Roprnson-DovuGcias, William Douglas, M.A., F.L.S., F.R.G.S., 
Orchardton, Castle Douglas, N.B. 


LIS? OF FELLOWS. X81 


1890 Rosson, John Emmerson, Hartlepool. 

1886 Rose, Arthur J., Brunnen Lodge, Upper Walthamstow-road; 
Walthamstow. 

1868 Rorunry, George Alexander James, 15, Versailles-road, Nor- 
wood, S.E. 

1894 RoruscuiLp, The Honble. Nathaniel Charles, F.Z.S., 148, Picca- 
dilly ; and Tring Park, Tring, Herts. 

1888 RoruscuiLtp, The Honble. Walter, F.Z.S., 148, Piccadilly, W ; 
and Tring Park, Tring, Herts. 

1890 RovrLenGs, G. B., 50, Russell-square, W.C. 

1892 RwvssE 1, S. G. C., 19, Lombard-street, E.C. 

1894 Rye, Bertram George, 212, Upper Parliament-road, Putney, 
S.W. 

1894 Ry.anps, Thos. Glazebrook, F.L.S., F.G.S., Highfields, Thelwall, 
Warrington. 


1885 Saper, Ernest, F.Z.S., F.R.G.S., Lynton House, South Side, Clap- 
ham Common, S.W. 

1891 Sr. Jonny, The Rev. John Seymour, B.A., 42, Castlewood-road, 
Stumford Hill, N. 

1875 Sane, Auguste, 13, Rue Guy de lu Brosse, Paris. 

1866 + SALVIN, Osbert, M.A., F.R.S., F.L.S., V.P.Z.S., 10, Chandos- 
street, Cavendish-square, W.; and Hawksfold, Fernhurst, 
Haslemere. 

1886 Sautwey, Reginald'E., Sungate, Hook-road, Kingston-on-Thames. 

1865 + Saunpers, Edward, F.L.S., St. Ann’s, Mount Hermon, Woking, 
Surrey. 

1861 t SaunpERs, G. 8., 20, Dents-road, Wandsworth Common, S.W. 

1886 Saunpers, Prof. Wm., Central Experimental Farm, Ottawa, 
Canada (President of the Eutomological Society of Ontario). 

1881 Sconuick, A. J., Allandene, Dorset-road, Merton Park, Wimble- 
don, S.W. 

1886 ScuppeER, Samuel H., Cambridge, Mass., U.S.A. 

1864 SEMPER, George, care of Bernhard Beer, Esq., 10, Newgate- 
street, E.C, 

1862 Suarp, David, M.A., M.B., C.M., F.R.S., F.L.S., F.Z.S:, Haw- 
thorndene, Hills-road, Cambridge; and University Museum of 
Zoology and Comparative Anatomy, Cambridge. 

1883 SHaw, A. Eland, M.R.C.S., Fakenham, Norfolk. 

1883 + SHELLEY, Capt. George Ernest, F.G.S., F.Z.S., 10, Thurlow- 
square, S.W. 

1887 Sticu, Alfred, Burlington-lune, Chiswick, W. 

1887 Sipawicx, Arthur, M.A., Fellow of Corpus Christi College, Oxford, 
64, Woodstock-road, Oxford, 


XXil 


1869 
1885 


LIST OF FELLOWS. 


Smitu, Henley Grose, F.Z.8., 5, Bryanston-square, Hyde Park, W. 
SoutH, Richard, Oxford-road, Macclesfield, Cheshire. 


* + Spence, William Blundell, Florence, Italy. 


1889 
1890 
1862 
1837 
1891 


1889 
1886 


1882 
1884 


1894 
1876 
1893 
1892 


1886 


1889 


1892 
1893 


SranpDEN, Richard §., F.L.8., Thorpe Hall, near Colchester. 

SrEarns, A. E., 99, Gloucester-terrace, Hyde Park, W. 

SreveEns, John §., 4, Pope’s-grove, Twickenham. 

SrEvENS, Samuel, F.L.S., Loanda, Beulah Hill, Upper Norwood, 
S.E. 

Stitt, Major John Nathaniel, 12, Seafield-terrace, Seaton, Devon ; 
and Junior United Service Club, Charles-street, St. James’s, S.W. 

Srraton, C. R., F.R.C.S., West Lodge, Wilton, Wilts, 

SurraGe, J. Lyddon, B.A., 82, Mornington-road, Regent's Park, 
N.W. 

Swanzy, Francis, Stanley House, Granville-road, Sevenoaks. 

SwINHOEk, Colonel Charles, M.A., F.L.8., F.Z.S., Avenue House, 
Cowley-road, Oxford. 

SwINnuHoe, Ernest, Avenue House, Cowley-road, Oxford. 

Swinton, A. H., Promenuden Gasse, 12, Ziirich, Switzerland. 


Taytor, Charles B., Rae-street, Rae Town, Kingston, Jamaica, 

TayLor, The Rev. George W., F.R.S. (Canada), St. Alban’s Rectory, 
Nanaimo, British Columbia. 

THEOBALD, F. V., M.A., Lecturer in Economic Entomology and 
Zoology to the South Eastern Agricultural College, Wye Court, 
near Ashford, Kent.. 

THORNEWILL, The Rev. C. F., M.A., Calderhall Vicarage, 
Whitchurch, Salop. 

THORNLEY, The Rev. A., M.A., South Leverton Vicarage, Lincoln. 

TownsEnD, Professor C. H. Tyler, Las Cruces, New Mexico, 
U.S.A. 


1859 + TRIMEN, Roland, F.R.S., F.L.S. (Curator of the South African 


1891 
1893 


1894 
1886 


1893 


1866 
1889 


Museum), Cape Town, Cape Colony. 

TUFFNELL, Carleton, Langley, Kenley, Surrey. 

TurNER, Henry Jerome, 13, Drakefell-road, St. Catherine's Park, 
Hutcham, 8.E. 

TuRNER, Thomas, Cullompton, Devon. 

Tutt, J. W., Rayleigh Villa, Westcombe Park, Blackheath, S.E. 


Unicn, F.W., Trinidad, British West Indies. 


VERRALL, George Henry, Sussex Lodge, Newmarket. 
Vivian, H. W., Glenafon, Taibach, South Wales; and Trinity 
College, Cambridge. 


LIST OF FELLOWS. XXili 


1876 WAKEFIELD, Charles Marcus, F.L.S., Belmont, Uxbridge. 

1886 Waker, Alfred O., F.L.S., Nant Glyn, Colwyn Bay, Denbigh- 
shire. 

1870 Waker, The Rev. Francis Augustus, D.D., F.L.S., Dun Mallard, 
Cricklewood, N.W. 

1878 Waker, James J., R.N., F.LS., 23, Ranelagh-road, Marine 
Town, Sheerness. 

1863 +Wa.tacr, Alfred Russel, D.C.L., Oxon., F.R.S., F.LS., F.ZS., 

Corfe View, Parkstone, Dorset. 

1866 + WatstnauaM, The Right Hon. Lord, M.A., LL.D., F.R.S., F.LS., 
F.Z.S., Vice-PresipENT, High Steward of the University of 
Cambridge, Merton Hall, Thetford, Norfolk ; and 66a, Eaton- 
square, S.W. 

1886 Warren, Wm., M.A., c/o The Honble. Walter Rothschild, Tring 
Park, Tring, Bucks. 

1869 Wareruouss, Charles O., Ingleside, Avenue Gardens, Acton, W. ; 
and British Museum, Cromwell-road, 8.W. 

1891 + Watson, Capt. E. Y., F.Z.S., Indian Staff Corps, care of Messrs, 
King & Co., 45, Pall Mall, S.W. 

1893 Wess, John Cooper, 32, Henslowe-road, Dulwich, S.E. 

1876+ WestEeRN, E. Young, 36, Lancaster Gate, Hyde Park, W. 

1882 Werymer, Gustav, Sadowa-strasse 21a, LElberfeld, SIthenssh 
Prussia. 

1886 WHEELER, Francis D., M.A., LL.D., Paragon House School, 
Norwich, 

1865 Wuitrr, The Rev. W. Farren, M.A., Stonehouse Vicarage, 
Gloucestershire. 

1884 Wuuitr, William, The Ruskin Museum, Meersbrook Park, 
Sheffield. 

1882 WiuiaMs, W. J., Zoological Society, 3, Hanover-square, W. 

1894. Witson, Edwin, Cherry Hinton-road, Cambridge. 

1894 Wo.tey-Dop, F. H., Box 225, Calgary, Alberta, N.W.T. Canada. 

1881 Woop, The Rev. Theodore, 23, Broderichk-road, Upper Tooting 
S.W. 

1894 Wootr, Michael Yeatman, 1, Marlborough-place, St. Johws Wood, 

NW, 

1891 WrovuacutTon, R. C., Conservator of Forests, Indian Forest Service, 

Bombay, India. 


1888 YeErsury, Colonel J. W., R.A., Army and Navy Club, Pall Mall, 
S.W. 

1892 Youpa.e, William Henry, F.R.M.S., 52, Main-street, Cockermouth, 
Cumberland. . 

1886 Youna, Morris, Free Museum, Paisley, N.B. 


C exiv 


ADDITIONS TO THE LIBRARY 


Durinc THE YEAR 1894. 


Apter (H.). Alternating Generations; a study of Oak Galls and Gall 
Flies; translated by G. R. Straton. 8vo, Oxford, 1894. 
The Translator. 


Aucock (A.). Natural History Notes from H.M.S. ‘‘ Investigator,”’ 
Deep-sea Dredging. Ser. II., No. 1. 


[Ann. & Mag. N. H. (6) xii., 1894. | The Author. 
AsumeEapD (W. H.). Monograph of the North American Proctotrypide. 
(Bull. U. S. N. Mus., No. 45, 1893. | The Author. 


BarTLert-Canvert (W.). Catalogo de los Lepiddépteros, Rhopaloceros i 
Heteroceros de Chile. 
[An. Univ. Chile, 1886. ] The Author. 


Bere (C.). Molus pyroblaptus, Berg. 
Las Cuestiones de Limites. 
{An. Soe. Cien. Arg., 1892. | 
Pseudoscorpionidenkniffe. 
{Zool. Anzeiger, 1893. | 


Descripciones de Algunos Heterdépteros nuevos 6 poco conocidos. 
[An. Mus. Nac. Montevideo, 1894. | The Author. 


BERTOLINI (Guiseppe). Illustrazione dei prodotti naturali del Mozambico. 
Dissertazione intorno ad insetti Ditteri. 4to, Bologna, 1862. 
[Mem. Acad. Sci. Vol. XII. | The Author. 


Bicor (J. M. F.). Insectes diptéres pour servir A la faune du Gabon. 

[Thoms. Arch. Ent. Il., 1858. ] 

Enumeration des Diptéres recueillis en Tunisie par M. Val 
Mayet. 8vo, Paris, 18838. Y ppd 

Descriptions de Diptéres Nouveaux. 

[Mem. Soc. Zool. France, 1892. | ; 

Catalogue of the Diptera of the Oriental region. Parts IT. and III. 

{Journ. As. Soe. Bengal, 1891. } ‘ Mr. Verrall. 


BLANDFORD (W. F. H.). Report on the destruction of beer-casks in India 
by a boring-beetle (Xyleborus perforans, Woll.). London, 1893. 
The Scolyto-platypini, a new Subfamily of Scolytide. 
[Trans. Ent. Soc. Lond., 1893.] 
Description d’un nouveau genre de Scolytides, Aricerus. 
{Ann. Soc. Ent. de Belgique, 1894. } ; 


The Rhynchophorous Coleoptera of Japan. Part III.. Scolyti 
|Trans. Ent. Soc. Lond., 1894. } , The Dyes 2 


(yaw) 


Boutuman (C. H.). The Myriapoda of North America. 


[ Bull. U. S. N. Mus., No. 46, 1898. | The Author. 
Boucusé (P. Fr.). Beitriige zur Insectenkunde. 
[Nov. Act. Acad. Leopold, 1833. ] Mr. Verrall. 
BRAUvER (Friedrich). Uber die Stellung der Gattung Lobogaster, Phil., im 
Systeme. 
[Sitzb. Akad. Wissensch., 1883. | Mr. Verrall. 


Broun (Thomas). Manual of the New Zealand Coleoptera. Parts V.—VLI. 
8vo, Wellington, 1893. 
(Col. Mus. and Geol. Surv. N.Z. | N.Z. Institute. 


CAMBRIDGE {O. P.). See Gopman (I. D.) and Satvin (0.). Biologia 
Centrali-Americana. 
CAMERON (P.). See Gonman (F. D.) and Satvry (0.). Biologia Centrali- 
Americana. 


Canptze (E.). Elatérides recueillis par M. EK. Modigliani aux bords du lac 
de Toba, & Sumatra. 


[Ann. Mus. Civ. St. Nat. Genova, 1894. The Author. 
Caszy (T. L.). Coleopterological Notices, V. New York, 1893. 
[Ann. N.Y. Acad. Sci. VII. ] The Author. 


CuHampion (G. C.). Onthe Tenebrionide collected in Australia and Tas- 
mania by Mr. J. J. Walker, R.N., during the voyage of H.M.S. 
‘Penguin,’ with descriptions of new genera and species (1 
plate). 

[Trans. Ent. Soe. Lond., 1894. } 


An Entomological Excursion to Corsica. 


(‘l'rans. Hnt. Soc. Lond. 1894. | The Author. 
See also GopmMAN (F. D.) and Satvin (O.). Biologia Centrali- 
Americana, 


Cowan (T. W.). The Honey-Bee. 12mo, London, 1890. The Author. 


Dauua Torre (C. G. de). Catalogus Hymenopterorum hucusque descrip- 
torum systematicus et synonymicus. Vols. I., II., VI., VI1., 
IX. 8vo, Lipsiae, 1892—4, Mr. Dunning. 


Eicunorr (W.). Huropiischen Borkenkifer. S8vo, Berlin, 1881. 
Mr. Dunning. 


Ericuson (W. EF). Naturgeschichte der Insecten Deutschlands. Abthe’- 
lung [. Coleoptera. 8vo, Berlin, 1893 —4. 
Band V., Lief. 2, 3, G. Seidlitz, 1893—4. 
Band VI., Lief. 6, J. Weise, 1893. Purchased. 


Fasre (J. H.). Souvenirs Entomologiques (1st Ser.), 2nded. S8vo, Paris, 
1894, Purchased. 


Fea (Leonardo). Viaggiodi . . . im Birmania e regioni vicine. 

Longicorn Coleoptera. By C. J. Gahan. S8vo, Genova, 1894. 

The Author. 

Foret (A.). Les Formicides des Indes et de Ceylan, I —III. 

[Journal Bombay N. H. Soe., 1892. ] 

Nouvelles espéces de Formicides de Madagascar (recoltées par M. 

Sikora), lere serie. — 
fAnn. Soc. Ent. Belg., 1892. | 


(| xxvi_) 


Foret (A.), continued— 
Quelques Fourmis de la Faune Mediterranéene. 
[Ann. Soe. Ent. Belg., 1892. | 
Note sur les Attini. 
[Ann. Soc. Ent. Belg., 1893. ] 
Abessinische und andere Afrikanische Ameisen. 
[Mittheil. Schweiz. Ent. Gesell., 1893. | 
Nouvelles Fourmis de |’ Australie et des Canaries. 
Note préventive sur un nouveau genre et une nouvelle espéze de 


Formicide (Camponotide). 
[Ann. Soc. Ent. Belg., 1893. | 


Die Ameisen Neu Seelands. Attini und Cryptocerini. 
[Mittheil. Schweiz. Ent. Gesell. Bd. 8. 1893.| 

Etudes Myrmecologiques en 1879, 2e partie. 

Hermaphrodite de |’ Azteca instabilis, Smith. 

Observations nouvelles sur la biologie de quelques Fourmis. 
[Bull. Soc. Vaud. Sc. Nat., XVI., XXVIIL, XXIX.] 

Die Ameisenfauna Bulgariens. 

[ Verhand. zool.-bot. Gesell. Wien, 1892. | 

Die Nester der Ameisen. Zurich, 1892. 

Les Formicides d’ Oran (Algerie). 

[Bull. Soc. Vaud. Sc. Nat., XXX. | 

Quelques Fourmis de Madagascar, de Nouvelle Zélande, ete. 
[Ann. Soc. Ent. Belg., 1894. | The Author. 
See also GRANDIDIER (A.). 


Fow.er (W. W.). See Gopman (F. D.) and Satvin (0.). Biologia Centrali- 
Americana. 


Gauan (C. J.). Description of a new Longicorn Beetle of the genus 
Diaxenes. 
| Ann. and Mag. N. H. (6), xiii., 1894. | 


On the characters of a new genus and six new species of Longicorn 
Coleoptera from New Guinea. 
[Ann. and Mag. N. H. (6), XIII., 1894. ] 


See Fra (L.). Viaggio di. Longicorn Coleoptera. The Author. 


Gopman (F. D.) and Satvin (O.). Biologia Centrali-Americana. 
Araneidea by O. P. Cambridge. Insecta by P. Cameron, G. C. 
Champion, W. W. Fowler, F. D. Godman, H. S. Gorham, O. 
Salvin, and D. Sharp. Parts CXIII.—CXI1X. 1894. 
The Editors. 


Goruam (H.8.). See Gopman (F. D.) and Satvin (0.). Biologia Centrali- 
Americana. 


GRanpIpIER (A.). Histoire Physique, Naturelle et Politique de Madagas- 
car. 28e fascicule et supplément, 
Les Formicides, par A. Foren. 4to, Paris, 1891. The Author. 


GuERIN-MENEVILLE (F. E.). Situation, Maladies et Amélioration des 
yaces du Vera Soie. S8vo, Paris, 1857. Mr. Bromilow. 


Hampson (G. F.).. The Fauna of British India, including Ceylon and 


Burmah. 
Moths, Vol. II. S8vo, Lond., 1894, The Author. 


a 


( xsi) 


HormaNnN (Ernst). Die Raupen der Gross-Schmetterlinge Europas. 4to, 
Stuttgart, 1893. Mr. Dunning. 


Houianp (W. J.). New and undescribed Genera and Species of West 
African Noctuide. 
[Psyche, 1894. | The Author. 


ImnorF (Ludwig). Studium der Koleoptern, S8vo, Basel, 1856. 
Mr, Dunning. 


JANET (C.). Sur les nématodes des glandes pharyngiennes des Fourmis. 
[C.R. Acad. des Sci. Paris, 1898. | 


Etudes sur les Fourmis, Notes 1-6. 
[Ann. Soc. Hut. France, &e., &c. 1893—94. ] 


Sur les Nerfs de Vantenne et les Organes chordotonaux chez les 
Fourmis. 


Sur le Systéme glandulaire des Fourmis. 
[C.R. Acad. Sci. Paris, 1894. | The Author. 


Karscu (F.). Westafrikanische Dipteren. 
[Zeitschr. f.-d.-ges. Naturwiss. Berlin, 1879. ] Mr. Verrall. 


Kine (Capt. P. P., R.N.). Descriptions, &c., of the Insects collected by 
° in the Survey of the Straits of Magellan. 
Hymenoptera (A. H. Haliday). 
Diptera (I. Walker). 


[Trans. Linn. Soc. Lond., 1837. ] Mr. Verrall. 
Kinestey (J. 8.). Classification of the Arthropoda. 
[Tufts’ College Studies, No. I., 1894. | The College. 


Kirsy (W. F.). Synonymie Catalogue of Lepidoptera-Heterocera (Moths). 
Vol. I. Sphinges and Bombyces. 8vo, London, 1892. 
Mr. Dunning. 


Kouse (H. J.). Hinfithrung in die Kenntniss der Insekten. 8vo, Berlin, 
1893. Mr. Dunning. 


KRASILSTSHIK (J.). La Graphitose et la Septicémie chez les insectes. 
[Mem. Soe. Zool. France, 1893. | The Author. 


LetTHiERRyY (L.), et SEVERIN (G.). Catalogue Général des Hemiptéres. 
Tome 1. Hetéroptéres-Pentatomide. Bruxelles, 1893. 
Purchased. 


Lintner (J. A.). Highth and Ninth Annual Reports on the Injurious and 
other Insects of the State of New York. 8vo, Albany, 
1892—93, The Author. 


Lorw (Hermann). Bemerkungen tiber die Familie der Asiliden.  4to, 
Berlin, 1851. 


Neue Beitrage zur Kenntniss der Dipteren. 4to, Berlin, 1856. 


Bidrag till kannedomen om Afrikas Diptera. 
[Ofvers. K. Vet.-Akad. Forh. 1862. | 


(| xxvii) 


Loew (Hermann), continued— 


Uber die bisher auf der Galizischen Seite des Tatragebirges beobach- 
teten Dipteren. 
[Fahrbuche Gel. Gesell. Krakau, 1870. ] 


Bemerkungen tiber die von Herrn F. Walker im 5 B. des Ento- 
mologist’’ beschriebenen igyptischen und arabischen Dipteren. 
[Zeits. Ges. Nat., 1873. ] Mr. Verrall. 


Lowne (EH. T.). The Anatomy, Physiology, Morphology, and Development. 


of the Blow Fly. Part V. 8vo, London, 1894. Purchased. 


McLacutan (R.). On two small collections of Neuroptera from Ta-chien-lu, 
Western China. 


[Ann. and Mag. N. H. (6) xiii, 1894. ] The Author. 
Two new species of Myrmeleonide from Madagascar. 
[Ann. and Mag. N. H. (6) xiii., 1894. } The Author. 


Martin (Matthew). The Aurelian’s Vade Mecum, 8vo, Exeter, 1785. 
Mr. Mosley. 


Mostey (S. L.). Insects on Forest Trees. 12mo, Huddersfield, 1890. 
The Author. 


OLIviER (Ernest). Descriptions d’espéces nouvelles de Lampyrides. 
[Ann. Soe. Ent. I'rance, 1894. | The Author. 


Ormerod (Miss Eleanor A.). Report of observations of Injurious Insects 
and Common arm Pests for 1893. The Authoress. 


OsTEN-SACKEN (Baron C.R.). Explanatory note of my views on the Sub- 
orders of Diptera. 
| Ent. Monthly Mag., 1893. | 


Zur Geschichte der sogenannten Brustgriite (breast-bone) der 
Cecidomyien. Rejoinder to Professor Brauer’s 'Thatsichliche 
Berichtigung, &c., in the Berliner Ent. Zeitschr., 1892. 


Two critical remarks about the recently published third part of the 
**Muscaria schizometopa’’ of Messrs. Brauer and Beryenstamm. 
[Berl. Ent. Zeitschr., 1893. | } 


On the characters of the three divisions of Diptera: Nemocera vera, 
Nemocera anomala, and Hrenochaeta. 
[Berlin. Ent. Zeitschr., 1892. | 


On the Atavic Index-characters, with some remarks about the classi- 
fication of the Diptera. 
{Berl. Ent. Zeitschr., 1894. ] 


Synonymica about Tipulide. 

Three Trochobolx from New Zealand and Tasmania. 

A remarkable case of malformation of the discal cell in a specimen 
ot Liogma glabrata. 

{Berl. Ent. Zeitschr., 1894. ] 


On the oxen-born Bees of the ancients (Bugonia), and their relation 


to Hristalis tenaz, a two-winged insect. S8vo, Heidelberg, 1894.- 


The Author. 


PacKarp (A. Bo; Notes on Gliuphisia and other Notodontidxe. Parts 1 
and 2. 


{| Psyche, 1893. | 


———————eE——eeerorrr err eee 


€ ets) 


Packarp (A. 8.), continued— 
On the Systematic Position of the Diptera. 
[‘* Science,’? N. York, 1893. | 
A Study of the Transformations and Anatomy of Lagon crispata, a 
Bombycine Moth. 
[Proc. Am. Phil. Soc., 1894. ] 


On the origin of the Subterranean Fauna of North America. 
[Am. Naturalist, 1894. | 


On the Inheritance of Acquired Characters in Animals, with a com- 


plete Metamorphosis. 
[Proc. Am. Acad. Sci., 1894.] 


Note on Thermobia domestica, and its occurrence in the United 
States. 
[Ent. Monthly Mag., 1894.] 


On the Systematic Position of the Siphonaptera, with notes on their 


structure. 
[Proc. Boston Soc. N. H., 1894.] The Author. 
Prout (Louis B.). Coremia ferrugaria, Haw., and unidentaria, Haw. 
8vo, London, 1894. The Author. 


Reep (E. C.). Introduccion al Estudio de los Insectés Himendpteros de 
Chile. Las Fosores 6 avispas cavadoras. 
[An. Univ. Chile, 1894. ] The Author. 


ReErp (Clement). Norfolk Amber. 
[Trans. Norf. and Norwich N. H. Soc. 1884, 1886.] Mr. Verrall. 


Riey (C. V.). Report on the Insecta, Arachnida, and Myriapoda collected 
by US. ‘ Eclipse’ expedition to West Africa, 1889—90. 
[Proc. U.S. Nat. Mus., Vol. XVI., 1893. | The Author. 


Ronpani (Camillo). Diptera Italica. Fase. I.—IIL. 
[Atti Soc. Ital. Milano, 1865—68. | 


Nota sugli Insetti Parassiti della Galleruca dell’ olmo. 
[Bull. Com. Agrar., Parma, 18/0. | Mr. Verrall. 


Satvin (O.). [See GopMan (F. D.). Biologia Centrali-Americana. } 


Saussure (H. de), et ZeHNTNER (Léo). Notice morphologique sur les 
Gryllotalpiens. 
[Revue suisse de Zoologie, 1894. | The Author. 


[See also Gopman (F. D.) and Sauvin (O.). Biologia Centrali- 
Americana. | 


Scupper (S. H.). The Butterflies of the Eastern United States and Canada, 
with special reference to New England. 3 Vols., Cambridge, 
Mass., 1889. 


The American Tertiary Aphid. 4to, Washington, 1594. 
Insect Fauna of the Rhode Island Coal Field. Syo, Washington, 


1893. : 
[Bull. U.S. Geol. Surv., No. 101, 1898. | 


Tertiary Tipulide, with special reference to those of Florissant, 
Colorado. 
[Proc. Am. Phil. Soe. X XXII., 1894. ] 


Scupper (8S. H.), continued— 


The North American Centhophili. 
[Proc. Am. Acad., Vol. XXX., 1894. | 


The effect of Glaciation and of the Glacial Period on the present 
Fauna of North America. 
[Am. Jour. Sci., 1894. | 


Tertiary Rhynchophorous Coleoptera of the United States. 4to, 
Washington, 1893. The Author. 


Sewuirz (Georg). Fauna Transsylvanica. Die Kifer (Coleoptera) Sieben- 
biirgens. 8vo, Konigsberg, 1891. 


Fauna Baltica. Die Kafer (Coleoptera) der Deutschen Ostee- 
provinzen Russlands. 8vo, Konigsberg, 1891. 


[See also Ericuson (W. F.). | Mr. Dunning. 


Semper (Georg). Die Schmetterlinge der Philippinischen Inseln. Beitrag 
zur Indo-Malayischen Lepidopteren-Fauna. Band I.: Die 
Tagfalter-Rhopalocera. 4to, Wiesbaden, 1886—92. 

The Author. 


Suarp(D.). [See Gopman (F. D.) and Sanvin (O.). Biologia Centrali- 
Americana. | 


Simmons (Orville L.). Development of the Lungs of Spiders. 
| Tufts’ College Studies, No. II., 1894. ] 


SinTenis (O.). Neu gefundene Tipuliden. 
[Sitzungsb. Dorpat Naturf. Ges. 1884. ] 


Srx (G. A.). Eine Entomologische Wandeling in Augustus. 
[' Tijdschr. voor Ent., 1858. | 


SmitH (John B.). Catalogue of the Lepidopterous Super-family Noctuidee 
found in Boreal America. 
[Bull. U.S. N. Mus., No. 44, 1893. ] The Author. 


SNELLEN VAN VOLLENHOVEN (S. C.). Beschrijving van eenige Nieuve 
Soorten van Diptera. 8vo, Amsterdam, 1862. Mr. Verrall. 


STENHAMMAR (C.). Forsdk till gruppering och revision af de Svenska 
.. Ephydrine. ; 
[Ofy. K. Vet.-Akad. Férh. 1844. | Mr. Verrall. 


Tepper (J. G. O.). The Blattarize of Australia and Polynesia. 
Descriptions of South Australia Brachysceloid Galls. 
Notes and Remarks on South Australian Rhopalocera. 
[Trans. Roy. Soc. South Austral., 1893. ] The Author. 


Tomson (C. G.). Opuscula Entomologica, Fase. 17,18. Purchased. 


TownsEnp (C. H.). A new Simulium from Southern New Mexico. 
[ Psyche, 1891. | 


Description of a Sarcophaga bred from Helix. 
[Psyche, 1892. ] The Author. 


Turt (J. W.). British Noctue and then Varieties. Vols. II., III. 8vo, 
London, 1892. Purchased. 


Geer, 


Voet (Jean |E.). Beschreibungen und Abbildungen hartschaaligter In- 
sekten. Coleoptera, Linn. Commentar von Dr. G. W. F. 
Panzer. 4to, Erlangen, 1798. Mr. Dunning. 


WALKER (J. J.). A visit to Damma Island, East Indian Archipelago. With 
notes on the Fauna by R. B. Sharpe and others. Parts I., II. 
[Ann. and Mag. N. H. (6) xiv., 1894. | The Author. 


Watson (HE. Y.). A proposed Classification of the Hesperiide, with a 
Revision of the Genera. 
[P. Z. S. Lond., 1893. ] The Author. 


WEED (Clarence M.). A Descriptive Catalogue of the Harvest Spiders 
(Phalangiidz) of Ohio. 3. 


[Proc. U. S. Nat. Mus., Vol. XVI. ] The Author. 
Waite (A. E. Holt). Butterflies and Moths of Teneriffe. (Kdited by Rash- 
leigh Holt White.) 4to, London, 1894. Mr. Dunning. 
Wittiston (S. W.). Contribution to a Monograph of the North American 
Syrphide. 


[Proc. Am. Phil. Soc., 1882. ] 


Dipterous Larve fromthe Western Alkaline Lakes, and their use as 


Human Food. 
[Trans. Conn. Acad., 1883. ] 


On the Classification of North American Diptera. 
[Ent. Am,, 1885.] Mr. Verrallt. 


ZEHNINER (Léo). [See Saussure (H. de), and Gopman.(F. D.), and 
SALVIN (O.). Biologia Centrali-Americana. | 


Periodicals and Publications of Societies. 


AFRICA. 
Care Town. South African Philosophical Society. Transactions. Vol. VII.; 
Vol. VIII., Part 1. The Society. 


AMERICA (NORTBRB). 


CANADA. 
Hatirax. Nova Scotian Institute of Science. Proceedings and Transac- 
tions. 2Ser., Vol. I., Part 3. The Institute. 


Lonpon, Ontario. The Canadian Entomologist. Vol. XXVI., 1894. 
By Exchange. 


Monrreat. Royal Society of Canada. Proceedings and Transactions. 
Vol. XI., 1894. The Society. 


() SExie of) 
Toronto. Entomological Society of Ontario. Report XXIV., 1894. 
The Society. 
Fruit Growers’ Association of Ontario. Report for 1893. 
The Association, 


UNITED STATES. 
Boston. Boston Society of Natural History. Proceedings. Vol. XXVI., 
1893. By Exchange. 


CAMBRIDGE, Mass. Museum of Comparative Zoology at Harvard College. 
Annual Report, 1892—93. The Curator. 


Davenport, Iowa. Academy of Natural Sciences. Proceedings. Vol. II., 
Part 2; Vol. V., Part 2; Vol. VI., PartI. The Academy. 


PHILADELPHIA. Academy of Natural Sciences of Philadelphia. Proceed- 
ings 1885; 1893, Parts 2, 3; 1894, Part 1. By Exchange. 


Entomological News and Proceedings of the Entomological Section. 
Vol. V. By Exchange. 


American Philosophical Society. Proceedings, 1893. 
The Society. 


American Entomological Society. Transactions, 1894. 
By Exchange. 


WaAsHINGTON. Insect Life, 1894. U.S. Dept. of Agric. 


* 


AMERICA (SOUTH). 
ARGENTINE REPUBLIC. 


Buenos Airrs. Academia Nacional de Ciencias, Cordova.  Boletin, 


Tomo XII. By Exchange. 
BRAZIL. 
Rio pe JANEIRO. Museo Nacional. Archivos, 1892. The Museum. 
CHILI. 


SantiaGo. Société Scientifique du Chili. Actes, 1893. The Society. 


WEST INDIES. 
JAMAICA. Institute of Jamaica. Journal. Vol. II. The Institute. 


TRINIDAD. Trinidad Field Naturalist’s Club. Journal. Vol. II. 
The Club. 


ASIA. 


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Bombay. Natural History Society. Journal. Vol. VIII., Parts 1—5; 
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(. xx) 


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Sypney. The Agricultural Gazette of New South Wales, 1894. 
Agric. Dept. 
Linnean oes of New South Wales. Proceedings, 2nd Series, 


Vol. VII By Exchange. 
WELLINGTON. New sitet Institute. Transactions and Proceedings 
Vol. XXVI. The Institut 
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Brunn. Naturforschender Verein in Briinn. Verhandlungen. Band 


XXXI. By Exchange. 
Vienna. K. k. zoologisch-botanische Verein (Gesellschaft) in Wien. 
Verhandlungen. Band XLIV. By Exchange. 


Wiener entomologische Verein. Jahresbericht, 1892—93. 
By Purchase. 


Wiener entomologische Zeitung, 1894. By Purchase. 


BELGIUM. 
BRuSsELs. Société Entomologique de Belgique. Annales. Tome XXXVIII. 
By Exchange. 
DENMARK. 
CoPENHAGEN. Entomologiske Meddelelser udgivne af Entomologisk 
Forening ved F’. Meinert. Band I—VII. 1888—94. 
Mr. Dunning. 
FRANCE. 
CaEn. Société Linnéenne de Normandie. Bulletin, 1893. By Exchange. 


Société Frangaise d’Entomologie. Revue. Tomes I.—XIII. 
1882—94, Mr. Dunming. 


Lyons. Société Linnéenne de Lyon. Annales, 1891—93. By Exchange. 
Paris. Société Entomologique de France. Annales, 1892. By Exchange. 
L’Abeille, 1894. By Purchase. 


ToutousE. Société d’ Histoire Naturelle. Bulletins, 1892—93. 
By Exchange. 


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Berutn. Entomologische Verein in Berlin. Berliner entomologische 
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Zeitschrift. 1894. By Eachange. 
Entomologische Nachrichten. 1893. Dr. Fremlin. 


Bericht iiber die wissenschaftlichen Leistungen des Jahres 1892. 
Purchased. 


c 


( xxx) 
Drespen. ‘‘Tris.’”’ Deutsche entomologische Ceieehe ey, Band VI.» 
Hit. 2. By Exchange. 


FrankFrort. Senckenbergischen naturforschenden Gesellschaft. Bericht. 
1894. By Exchange. 


Srertin. Entomologische Verein. Entomologische Zeitung. Jahrg. LY. 
By Exchange. 


WERNIGERODE. Naturwissenschaftliche Verein des Harzes. Band VIII. 
18 


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WIeEsBADEN. Nassauischer Verein fiir Naturkunde Jahrbucher. Jahrg. 
XLVI. By Exchange. 


GREAT BRITAIN AND IRELAND. 


BiRMINGHAM. Philosophical Society. ~ Proceedings, 1893. The Society. 


Dusiin. Royal Dublin Society. Transactions, Ser. 2, Vol. IV., pt. pe 
Vol. V., pts. 1—4. Proceedings, New Ser. SS avols VIL., pt. 
Vol. Vit, pts. 1, 2. The Society. 


Essex. Essex Naturalist, being the Journal of the Essex Field Club. 


Vol. VII., pts. 6—12. By Exchange. 


HertrorD. Hertfordshire Natural History Society and Field Club. 


Transactions. Vol. VII., pts. 8, 9. The Society. 
Lonpon. Annals and Magazine of Natural History. 1894. By Purchase. 
City of London Entomological and Natural History Society. 


Transactions for 1893. The Society. 
Entomologist (The). 1894. T. P. Newman. 
Entomologist’s Monthly Magazine, 1894. The Editors. 
Entomologist’s Record and Journal of Variation. 1894. 

The Editor. 
Linnean Society of London. Transactions, Vol. V., pt. 11; Vol.VI., 
pts. 1, 2. Journal, Nos. 157, 158. By Rechange. 
Nature. 1894. The Publishers. 
Nature Notes. 1894. The Publishers. 


- Quekett Microscopical Club. Journal. 1894. The Club. 


Royal Agricultural Society. Journal. 3rd Ser., Vol. V. 
The Society. 


Royal Microscopical Society. Journal. 1894. The Society. 
Royal Society. Proceedings. Nos. 328—338. The Society. 


Year Book of Scientific Societies. 1894. By Purchase. 
Zoological Record for 1893. By Purchase. 
Zoological Society. Proceedings, 1894. Transactions, Vol. XIII., 
pt. 9: By Exchange. 
Zoologist (The). 1894. T. P. Newman. 
HOLLAND. 


Tue Hagur. Tijdschrift voor Entomologie. Jahr. 1892 - 
By Exchange. 


A 


a 


(esr: |) 


ITALY. 


FLORENCE. Societa Entomologica Italiana. Bullettino. 1894. 
By Exchange. 
Genoa. Museo Civico di Storia Naturale di Genova. Annali. Ser. 2 


=! 


Vol. XIII. By Exchange. 
RUSSIA. 
Moscow. Société Impériale des Naturalistes de Moscou. Bulletin. 1893, 
Nos. 2—4; 1894, No. 1. By Exchange. 
SWEDEN. 
SrocKHOLM. Entomologiska Férenigen i Stockholm. Entomologisk 
Tidskrift. 1894. By Exchange. 


Kongliga Svenska Vetenskaps-Alkademie Bihang, XVIII. Afd. 4. 
The Academy. 
SWITZERLAND. 


ScHAFFHAUSEN. Schweizerische entomologische Gesellschaft. _ Mitthei- 
lungen. 1894. By Exchange. 


THE 
TRANSACTIONS 


OF THE 


ENTOMOLOGICAL SOCIETY 


OF 
LONDON 
For tHE Year 1894. 


I. On a collection of Lepidoptera from Upper Burma, 
by Epwarp Mrnyrick, B.A., F.Z.S. 


[Read Oct. 18th, 1893.] 


THE species enumerated in this paper were collected 
by Surgeon-Captain N. Manders, whilst taking part in 
those military movements, directed to the pacification 
of the Shan States and their neighbourhood, which 
followed the annexation of the territory. Most of the 
country visited was previously unknown to Europeans, 
and highly interesting. ‘The unhealthy climate, however, 
naturally kept Dr. Manders’ hands full of other work, 
and the continued presence of hostile natives made 
collecting always dangerous and often impossible ; whilst 
an unfortunate wound from an enemy in ambush 
eventually led to his being invalided home when just 
about to visit some of the most promising regions. 
Under these trying circumstances the collection is a 
remarkable record of persevering work. I had originally 
hoped to have worked out the whole, but in the face of the 
pressure of other work this has proved impossible, and 


TRANS, ENTs SOC. LOND. 1894.—PART I. (MARCH.) A 


9 Mr. Edward Meyrick on a collection 


part has been transferred to other hands. ‘The families 
or groups here presented, however, are worked out in 
full, viz., the slretiadw, Pyralidina, and Tineina. 


ARCTIADA. 


1. Pelosia tetrasema, sp. n. 


$9. 42—45 mm. Head and palpi ochreous-orange, apex of 
palpi blackish. Thorax whitish-ochreous, with an orange central 
spot, a round blackish posterior spot, and a black longitudinal mark 
on patagia; in ¢ also two blackish spots on collar, Abdomen 
ochreous-orange. Forewings whitish-ochreons ; a cloudy roundish 
blackish spot in dise posteriorly, anda second beneath it on fold ; in 
& both reduced to dots; cilia yellow. Hindwings in ¢ ochreous- 
whitish, hindmargin yellow, in @ wholly ochreous-yellow ; cilia 
yellow. 

Hab. oni; three specimens. 

Allied to P. puncticollis, Butl. 


2. Bizone javanica, Butl. 


Hab. Fort Stedman. 
3. Bizone harterti, Elwes. 
Hab. Jkoni. 


A. Ammatho defecta, Walk. 
Hab. Fort Stedman. ; 


5. Ammatho disticha, sp. n. 


¢. 27mm. Head, palpi, and thorax ochreous-rosy. Abdomen 
pale rosy. Forewings ochreous-rosy ; a black dot near base ; two 
nearly straight transverse series of small round blackish spots, first 
of four, at 1, second of five, median. Hindwings pale rosy. 


Hab. Fort Stedman; one specimen. 


6. Aimmatho epiaantha, sp. n. 


g. 28mm. Head deep yellow. Thorax deep yellow, with a 
transverse series of four black dots, and one near posterior ex- 
tremity. Abdomen ochreous-yellow. Forewings rather deep 
ochreous-yellow ; markings cloudy blackish-fuscous ; a black dot 
near base; a transverse row of dots about }, angulated above 
middle ; a nearly straight transverse row of dots before middle ; a 


of Lepidoptera from Upper Burma. 3) 


third series from 2 of costa to 2 of inner margin, rather strongly 

curved outwards on upper 2, five median dots produced into short 

streaks posteriorly, and two additional short streaks above upper- 

most of them ; cilia yellow. Hindwings and cilia pale yellowish. 
Hab. Koni; one specimen. 


7. Miltochrista rubricosa, Moore. 
ITab. Fort Stedman. 


8. Miltocnrista siinica, Moore. 
Hab. Koni. 


9. Miltochrista callinoma, sp. n. 


9. 27 mm. Head pale rosy, face ochreous-whitish. Thorax 
palo rosy, towards middle whitish, with two blackish dots. Abdo- 
men pale rosy. Forewings rosy ; all veins except costal branches 
marked with blackish-grey lines, edged on both sides with grey- 
whitish shades ; these do not reach hindmargin, which is wholly 
rosy ; cilia grey-whitish. Hindwings light rosy ; cilia whitish. 

Hab. Koni; one specimen. 


10. Miltochrista eccentropis, sp. n. 

g. 20 mm. Head white. Palpi dark fuscous, apex white. 
Thorax orange, posterior extremity white. Abdomen ochreous- 
yellowish. Forewings white; a dark grey roundish spot near 
base, followed by two curved transverse series of similar spots, five 
in each series ; a curved orange antemedian’ fascia; a small black 
discal spot beyond middle; a fine blackish transverse line from 
beyond middle of costa to 3 of inner margin, forming a rather 
angular bend outwards round discal spot; beyond this all veins 
marked by blackish well-defined lines; cilia white. Hindwings 
ochreous-whitish ; hairs towards base more yellowish ; veins 
towards hindmargin marked with short cloudy dark grey streaks, 
sometimes confluent, diminishing downwards and not reaching anal 
angle ; cilia white. 


Hab. ‘Koni; two specimens. 


11. Miltochrista celidopa, sp. n. 


2¢@. 20—25 mm. Head and thorax ochreous-yellow. Abdo- 
men paie ochreous-yellowish, anal tuft grey. Forewings whitish- 
ochreous, base, costa, and a hindmarginal band ochreous-yellow, 


4 Mr. Edward Meyrick on a collection 


sometimes wholly ochreous-yellow ; a black dot near base ; a round 
black discal dot beyond middle; cilia ochreous-yellow. Hind- 
wings pale ochreous-yellowish, sometimes suffused with grey except 
towards hindmargin. 


Hab. Koni; three specimens. 


12. Miltochrista geodetis, sp. n. 


$9. I18—22 mm. Head blackish, sides orange. Thorax 
orange, with black dots on shoulders and patagia, sometimes con- 
fluent, a black transverse mark in middle of back, and a spot near 
posterior extremity. Abdomen blackish, sides and apex orange. 
Forewings orange; markings black; a dot near base; a small 
round spot on or near costa at 1, another beneath it near inner 
margin, a third in disc at 3, and sometimes a fourth on anal angle ; 
a streak along posterior half of costa round apex and hindmargin 
to anal angle ; cilia blackish. Hindwings dull orange ; a blackish 
streak along hindmargin and round apex ; sometimes a more or 
less broad dark fuscous suffusion before this ; sometimes a small 
dark fuscous spot beneath costa at 2; cilia blackish. 


Hab. ‘Koni; three specimens. 


13. Owacme disstmilis, Hamps. 


Hab. Joni. 


14, Lebena fragilis, Swinh. 
Hab. Keni. 


15. Rajendra tripartita, Walk. 
Hab. Fort Stedman. 


16. Phissama transiens, Walk. 
Hab. Fort Stedman, 


17. Spilosoma indica, Guér. 
Hab. Fort Stedman. 


18. Detopeia vulchella, L. 
Hab. Koni. 


of Lepidoptera from Upper Burma. 5) 


PYRALIDINA. 
PYRAUSTID. 


1. Margaronia nitidicostalis, Gn. 
Hab, Koni. 


2. Margaronia amphitritalis, Gn. 


Hab. Fort Stedman. 


3. Margaronia unionalis, Hb. 
Hab. (Koni: large but otherwise typical. 


4, Margaronia celsalis, Walk. 
Hab. oni. 


5. Margaronia tyres, Cr. 


Hab. Koni. 


6. Margaronia callizona, sp. n. 

&. 25mm. Thorax dark fuscous, with two white stripes. Ab- 
domen dark fuscous, on sides and beneath white, above with a 
broad orange band occupying three segments before apex, edged 
with black. Forewings dark fuscous; markings violet-white, 
thinly scaled ; a small wedge-shaped mark in disc at 1 ; a narrow 
white streak along inner margin from base to 2, indented in middle; 
a large oblique elongate pear-shaped blotch in disc before middle, 
nearly reaching margins ; a pale grey transverse mark in middle 
of disc ; a large transverse oval blotch beyond middle, not reaching 
margins ; a transverse white spot beneath costa at 3, emitting a 
fuscous-whitish line to inner margin before anal angle ; cilia dark 
fuscous, with a white patch above anal angle. Hindwings violet- 
white, thinly scaled; a moderately broad dark fuscous hind mar- 
ginal band, including a cloudy fuscous-whitish line ; cilia fuscous 
with a cloudy dark fuscous line, on lower half of hindmargin 
white except towards anal angle. 


Hab. Fort Stedman; one specimen. 


Distinct from all near allies by the orange band of 
abdomen. 


7. Margaronia principalis, Walk. 
Hab. Fort Stedman, 


6 Mr, Hdward Meyrick on a collection 


8. Botyodes asialis, Gn. 
Hab. Koni. 


9. Omiodes bianoralis, Walk. 
Hab. Koni. 


10. Omiodes vulgalis, Gn. 
Hab. Koni. 


11. Agrotera effertalis, Walk. 
Hab. oni. 


12. Conogethes rigidalis, Snell. 
Hab. Fort Stedman. 


13. Conogethes plagiferalis, Walk. 
Hab. Fort Stedman. 


14. Conogethes boteralis, Walk. 
Hab. Mone, Fort Stedman. 


15. Conogethes amyntalis, Walk. 
JTab. Koni. 


Loxocorys, n. g. 

Face oblique, forming a rounded-conical projection beneath; a 
small erect scaletuft on crown between antenne ; ocelli distinct ; 
tongue developed. Antenne 2, in ¢ stout, filiform, minutely 
ciliated (1). Labial palpi rather long, curved, ascending, shortly 
rough-scaled beneath, terminal joint moderate, loosely scaled, 
pointed. Maxillary palpi rather long, filiform, Abdomen in ¢ 
with moderate anal tuft. Forewings with vein 7 from near 8, 
9 and 10 out of 8. Hindwings 1; veins 3, 4, 5 approximated at 
base, 7 out of 6 near origin, anastomosing with 8 to middle. 


16. Loxocorys sericea, Butl. (Scopula). 
Hab. Fort Stedman. 
17. Notarcha quaternalis, Z. 
Hab. Koni. 


18. Phlyctenia itemalesalis, Walk. 
Hab. Fort Stedman, 


of Lepidoptera from Upper Burma. 7 


19. Phlyctenia ferrugalis, Hb. 
Hab. Mone. 


20. Hyalobathra dialychna, sp. n. 

“@. 20-21 mm. Head, palpi, antenne, thorax, and abdomen 
pale brownish-ochreous ; palpi 12, base white; antennal cilia- 
tions 14. Forewings with apex rectangular ; brownish-ochreous, 
posterior half suffused with reddish-brown ; lines indistinct, 
blackish-grey ; first curved, indented in middle ; a cloudy blackish- 
grey blotch in middle of dise ; second line forming an oblique 


blackish mark on costa at 2, obtusely bent above middle, below 


middle again bent inwards to beneath discal blotch, and thence 
again rectangularly bent to inner margin at 2 ; a bright ochreous- 
yellow suffused blotch extending along apical third of costa ; cilia 
white, with a blackish-grey basal line. Hindwings pale brownish- 
ochreous, towards costa whitish-ochreous ; traces of a grey discal 
blotch and some blackish dots indicating second line as in fore- 
wings, but very indistinct ; an indistinct fine waved blackish-grey 
subterminal line ; cilia as in forewings. 


Hab. Koni; two specimens. 
21. Isocentris illectalis, Walk. 
Hab. (oni. 
22. Pyrausta ablactalis, Walk. 
Hab. Fort Stedman. 
23. Pyrausta miniosalis, Gn. 
Hab. Fort Stedman. 
24. Pyrausta celatalis, Walk. 
Hab, Fort Stedman. 
25. Pyrausta extinctalis, Christ. 
Hab. Koni. 
26. Pyrausta abruptalis, Walk. 
Hab. Fort Stedman. 


27. Acharana otrealis, Walk. 
Hab. Yoni, Fort Stedman. 


28. Titanio comparalis, Hb. 
Hab, Mone. 


8 Mr. Edward Meyrick on a collection 


29. Titanio fessalis, Swinh. 
Hab. Koni. 


Labial and maxillary palpi unusually long; frontal 
plate short, rounded; hindwings with ves 4 and 5 
stalked. 


30. Hellula undalis, F. 
Hab. Koni. 


31. Metasia zanclogramma, sp. n. 

9. 14-16 mm. Head, antenne, thorax, and abdomen ochreous- 
whitish, shoulders irrorated or suffused with dark fuscous. Palpi 
white, terminal joint and apex of second dark fuscous. Legs 
whitish, anterior pair banded with dark fuscous. Forewings 
elongate-triangular, costa hardly arched, apex roundpointed, hind- 
margin sinuate ; pale greyish-ochreous, more or less irrorated with 
fuscous ; costa suffused with dark fuscous on basal half; lines dark 
fuscous ; first from + of costa to + of inner margin, curved ; two 
subquadrate spots outlined with dark fuscous in disc; a small 
blackish spot on costa beyond middle; second line rising from 
posterior extremity of a small longitudinal black mark on costa at 
3. preceded and followed by a clearer whitish-ochreous spot, running 
nearly straight to anal angle, slightly indented in middle, obsoletely 
continued upwards to lower margin of second discal spot, thence 
again distinct to % of inner margin ; a cloudy dark fuscous line along 
hindmargin from apex to near anal angle : cilia rather light fuscous, 
base whitish-ochreous. Hindwings with colour, second and _ hind- 
marginal lines as in forewings, but second line without costal spot ; 
a small dark fuscous discal spot, connected with second line ; cilia 
fuscous-whitish, with a fuscous subbasal line. 


Hab. ‘Koni; two specimens. 
Very like some of the other small obscure species of 


the genus, but distinguished from all by the dark costal 
mark from which the second line rises. 


METASIODES, n. g. 


Characters of Metasia, but face without prominence, 
labial palpi ascending, terminal joint short, obtuse. 


32. Metasiodes heliaula, sp. n. 


69. 15—18 mm. Head ochreous-whitish, with two fuscous 
spots before antennee ; frontal prominence hardly perceptible. Palpi 


of Lepidoptera from Upper Burma. 9 


white, terminal joint and apex of second dark fuscous. Antenne 
whitish-ochreous, ciliations in ¢ 3. Thorax and abdomen ochreous- 
yellow. Legs whitish, anterior pair banded with dark fuscous. 
Forewings elongate-triangular, costa almost straight, apex round- 
pointed, hindmargin sinuate ; bright yellow, more or less ferru- 
ginous-tinged ; eight small blackish spots on costa, the sixth double 
and tending to form a semicircular ring ; lines ferruginous, rather 
irregular ; first at 1; two quadrate spots outlined with dark fuscous 
in disc, separated by a quadrate semitransparent whitish spot ; 
second line waved, running from eighth costal spot near apex to 
anal angle, thence obsoletely continued to beneath second discal spot, 
and thence again distinct to } of inner margin; a thick dark 
fuscous line along upper 3 of hindmargin, preceded by a ferruginous 
suffusion ; cilia yellowish, with blackish spots at apex and middle 
of hindmargin. Hindwings yellow ; a small dark fuscous discal 
spot ; second line as in forewings, but rising from 2 of costa; a 
ferruginous apical patch, bordered by a thick dark fuscous line 
along upper half of hindmargin: cilia as in forewings, sometimes 
with a cloudy dark grey subbasal line on upper half of hindmargin. 


Hab. Koni, Fort Stedman ; five specimens. 


33. Nacoleia contingens, Moore. 
Hab. (Koni. 


34. Sameodes cancellalis, Z. 


Hab. Soni. 


35. Diasemia grammalis, Dbld. 
Hab. Koni. 


36. Bocchoris inspersalis, Z. 


Hab. Koni. 


37. Stegothyris diagonalis, Gn. 
Hab. Koni, Mone, Fort Stedman. 


38. Cnaphalocrocis medinalis, Gn. 


Hab. Mone. 


39. Dolichosticha venilialis, Walk. 
Hab. Fort Stedman 


10 Mr. Edward Meyrick on a collection 


40. Dolichosticha marisalis, Walk. 
Hab. Koni. 


41. Dolichosticha perinephes, Meyr. 
Hab. Tabet. 


42, Pagyda salvalis, Walk. 
Hab. Koni. 


43, Wilodes fuividorsalis, Hb. 
Hab. Fort Stedman. 


44, Nuusinoe onychinalis, Gn. 
Hab. Fort Stedman. 


45. Nausinoe geometrals, Gn. 


Hab. Fort Stedman. 


46, Siriocauta testulalis, Hb. 
Hab; \ Wont. 


47, Cataclysta mesorphna, sp. n. 

¢. 12mm. Forewings dark fuscous ; a moderate orange fascia 
at 1, not reaching costa, preceded and followed by obscure pale 
leaden lines; apical area orange, indented by a long leaden- 
metallic wedge-shaped projection from costa beyond 3, and en- 
closing a leaden-metallic spot on anal angle, and a silvery-white 
narrow wedge-shaped fascia, margined with dark grey, from costa 
before apex to hind margin below middle : cilia dark fuscous-grey. 
Hindwings with vein 8 absent ; dark fuscous, sprinkled with white 
in dise ; a longitudinal orange streak near inner margin ; a slender 
whitish curved transverse line beyond 2; four rather large round 
black spots on hindmargin, separated by orange hindmarginal 
dots surmounted by golden-metallic scales, first and fourth spots 
including central golden-metallic spots ; cilia fuscous, base darker, 

Hab. Koni; one specimen. 


48. Nymphula bifurcalis, Pryer. 
Hab. Fort Stedman. 


49. Nymphula turbata, Butl. 
Hab, Fort Stedman. 


of Lepidoptera from Upper Burma. ie 


50. Nymphula myina, Meyr. 
Hab. Koni. 


51. Nymphula fluctuosalis, Z. 
Hab. (oni. 


52. Nymphula depunctalis, Gn. 
Hab. ‘Koni. 


53. Mixophyla erminea, Moore. 
Hab. Fort Stedman. 


54. Orambostenia angustifimbrialis, Swinh. 
Hab. Fort Stedman. 


55. Donacaula chlorosema, sp. n. 

¢. 17—20mm. Head, antennz, thorax, and abdomen white ; 
antenne subdentate, ciliations $. Labial palpil$,fuscous. Maxil- 
lary palpi fuscous, apex white. Legs fuscous, posterior tibie 
white. Forewings elongate-triangular, hindmargin rather strongly 
rounded beneath ; 11 running into 12; white; markings pale 
ochreous-yellow, faint and cloudy ; two or three small spots in 
dise near base ; first line at 4, bent above middle, partially inter- 
rupted ; two spots on costa beyond middle, one at ~, and an erect 
mark from inner margin before anal angle, with some faint 
scattered scales in disc between these: cilia white. Hindwings 
white ; a small pale ochreous-yellow spot at anal angle; cilia 
white. 


Hab. Koni; two specimens. 


56. Schenobius punctellus, Z. 
Hab, Koni. 


57. Scirpophaga wanthogastrella, Walk. 
Hab. Fort Stedman 


PYRALIDIDZ. 
58. Hereulia psamathopis, sp. n. 


¢d. 27—31 mm. Head, thorax, and abdomen light brownish- 
ochreous, crown more yellow-ochreous. Antennal ciliations 13. 
Forewings with costa slightly sinuate, apex obtuse, hindmargin 
rounded, oblique ; light brownish-ochreous, irrorated with fuscous ; 


12 Mr. Edward Meyrick on a collection 


first line faintly darker, almost obsolete ; second line thick, very 
cloudy, fuscous, from # of costa to before } of inner margin, 
somewhat sinuate: cilia light brownish-ochreous, with cloudy 
fuscous lines. Hindwings with colour as in forewings, but lighter, 
hairs in dise towards base reddish ; second line as in forewings, 
but more distinct, somewhat curved ; cilia as in forewings. 


Hab. Koni; two specimens. 
59, Pyralis vibicalis, Ld. 
Hab. Koni. 


60. Pyralis pictalis, Curt. 
Hab. Tabet, Koni. 


Prosaris, n. g. 


Face with projecting tuft of scales ; ocelli distinct ; tongue 
developed. Antenne 2, in ¢ serrulate, ciliated (2). Labial palpi 
long, porrected, curved downwards, clothed beneath throughout 
with very long dense loosely-appressed hairs, terminal joint 
moderately long. Maxillary palpi rather short, thick, triangularly 
dilated with scales. Abdomen in ¢ with moderate anal tuft. 
Anterior femora in ¢ with tuft of hairs beneath; middle and 
posterior tibize and first joint of tarsi in ¢ clothed with long 
rough hairs. Forewings with vein 1 shortly furcate, 7 and 8 out of 
9, Hindwings in ¢ above with an erect triangular tuft of scales 
below middle, beneath with a large bladderlike swelling on vein 8 
towards base ; 7 out of 6, anastomosing very shortly with 8. 


61. Prosaris pernigralis, Rag. (?) 

g. 21 mm. Head reddish ochreous, face mixed with dark 
fuscous. Palpi dark reddish-fuscous. Thorax dark fuscous mixed 
with ochreous, purplish-tinged. Forewings triangular, costa nearly 
straight, hardly sinuate in middle, apex obtuse, hindmargin bowed 
rather oblique ; deep purplish-reddish-fuscous, irrorated with 
blackish ; hindmarginal area lighter and greyer ; a suffused deep 
red patch towards costa near apex, surrounding a small clear 
whitish-ochreous spot on costa: cilia deep purple-reddish, mixed 
with blackish. Hindwings with colour and cilia as in forewings ; 
discal scaletuft blackish, followed by a deep reddish suffusion, 
including a suffused whitish-ochreous dot; a suffused blackish 
shade at 3, parallel to hindmargin. 


Hab, Koni; one specimen. 


of Lepidoptera from Upper Burma. 13 


M. Ragonot described his pernigralis from a single 9 ; 
hence he was not acquainted with the full generic 
characters, and his specific description also differs in 
some particulars, which are probably sexual; I have, 
therefore, described my ¢ in full, but I have httle doubt 
that it is truly identical with his species. 


62. Huboloma nummosalis, Rag. 
Hab. Koni. 


M. Ragonot’s figure is poor; the ocelli are present ; 
the face has a projecting tuft of scales; im the forewings 
vein 6 rises out of 7, but this character is very probably 
inconstant. 

SICULODID. 
63. Striglina idalialis, Walk. 

Hab. Mone. 


64. Siculodes subrosealis, Leech. 
Hab. Fort Stedman. 


PHYCITIDA:: 
65. Heteroglypta ephippella, Rag. 
Hab. Koni, Mone. 
66. Myelois robusta, Moore. 
Hab. Koni. 
67. Rhodophea duplicella, Rag. 
Hab. (oni. 


68. Canthelea gratella, Walk. 
Hab. Soni. 


69. Piesmopoda steniella, Rag. 
Hab. (Yoni. 
70. Dera spurcella, Rag. 
Hab. Koni. 


71. Htiella zinckenella, Tr. 
Hab. (oni. 


72. Critowia subconcinnella, Rag. 


Hab. Koni. 


14 Mr. Edward Meyrick on a collection 


GALLERIAD. 


73. Lamoria planalis, Walk. 
Hab. Koni. 


74. Lamoria rufivena, Walk. 
Hab. Fort Stedman. 


CRAMBIDA. 
70. Aquita torrentella, Meyr. (?) 


Hab. Koni; the single specimen differs somewhat from 
those originally described from Australia, but I cannot 
venture to separate it specifically. 


76, Surattha invectalis, Walk. 
Hab. Mone. 


77. Ancylolonia indica, Feld. 
fab, Koni. 


18. Ptychopseustis aneenella, Snell. 
Hlab. Koni. 


TINHINA. 
GELECHIAD. 
1. Anacampsis seutata, sp. n. 


g. 12 mm. Head and thorax ochreous, face and _ palpi 
whitish-ochreous. Antenne fuscous-whitish. Forewings elongate, 
narrow, costa gently arched, apex round-pointed, hind margin 
oblique, hardly rounded ; dark slaty-fuscous ; a moderately broad 
whitish-ochreous streak along inner margin from base to 3, pointed 
posteriorly ; a moderate triangular ochreous-white spot on costa 
about +; some whitish dots round apical margin: cilia dark slaty- 
fuscous. Hindwings and cilia whitish-grey, slightly purplish- 
tinged. 


Hab. Fort Stedman; one specimen. 


HIERANGELA, 0. &. 


Head smooth; ocelli absent; tongue developed. Antenne $, 
in ¢ serrulate, simple, basal joint slender, without pecten. Labial 
palpi very long, recurved, smooth, slender, second joint somewhat 
thickened, terminal joint longer than second, acute. Maxillary 


of Lepidoptera from Upper Burma. 15 


palpi rudimentary. Posterior tivize clothed with hairs. Forewings 
with vein 1b furcate, 2 from before angle, 8 and 9 out of 7, 7 to 
costa (?), 11 from beyond middle. Hindwings ¢, narrow, parallel- 
sided, apex strongly and acutely produced, hindmargin beneath 
apex straight, oblique, cilia 3; veins 3 and 4 from a point, 6 and 
7 from a point. 


2. Hierangela erythrogramma, sp. n. 

g. 14 mm. Head yellow, with red central-streak on crown, 
face white. Palpi whitish, second joint red above towards apex. 
Antenne whitish-ochreous, towards base reddish above. Thorax 
yellow, with four longitudinal red stripes. Abdomen pale grey. 
Forewings very elongate, broadest near base, thence gradually 
narrowed to apex, acute; bright yellow ; a crimson-red costal 
streak from base to ?, paler posteriorly, leaving extreme costal 
edge whitish ; a longitudinal median crimson-red streak from base 
to costa before apex, interrupted at 4: space between this and 
costal streak suffused with fuscous ; a small fuscous spot on lower 
margin of median streak before middle ; a transverse crimson-red 
spot from inner margin near base, meeting median streak ; a small 
red spot on inner margin beyond this ; a crimson-red streak along 
inner margin from before middle to apex, interrupted above anal 
angle and below apex : cilia grey, on costa pale ochreous, with a 
blackish apical hook and a small golden-metallic subbasal spot at 
apex, base beneath apex reddish. Hindwings and cilia grey, 
towards anal angle whitish-grey. 

Hab. Fort Stedman; one specimen. 


8. Cladodes arotraea, sp. n. 

g. 11—14 mm. Head and thorax whitish-ochreous, with three 
fuscous stripes. Palpi whitish-ochreous, with a dark fuscous line 
on each side of anterior edge. Antennz whitish-ochreous, spotted 
with fuscous. Forewings elongate, narrow, costa gently arched, 
apex pointed, hindmargin almost straight, oblique; pale whitish- 
ochreous, all veins suffusedly margined with dark fuscous ; a round 
black dot in dise at 2, a second nearly beneath it on fold, and a 
third in disc at 3; a small suffused blackish apical spot ; cilia pale 
whitish-ochreous, with an indistinct fuscous line. Hindwings pale 
grey ; cilia grey-whitish. 

Hab. ‘Koni; two specimens. 


Onepata, Walk. 


Head smooth; ocelli almost concealed; tongue developed. 
Antenne #, in ¢ simple, basal joint moderate, without pecten. 


16 Mr. Edward Meyrick on a collection 


Labial palpi very long, recurved, smooth, second joint rather 
thickened above middle, terminal joint as long as second, slender, 
acute. Maxillary palpi rudimentary. Posterior tibiz clothed with 
hairs above. Forewings with vein 1b furcate, 2 almost from angle, 
3 and 4 stalked, 5 absent, 7 to costa, 8 and 9 out of 7, 11 from 
beyond middle. Hindwings somewhat over 1, trapezoidal, hind- 
margin not indented, cilia # ; veins 3 and 4 stalked, 5 absent, 6 and 
7 stalked. 

Near Cladodes; Walker’s description is of course 
inadequate. 


4. Onebala blandiella, Walk. 


Hab. Mone; one specimen. Walker’s type is from 
Ceylon, and differs slightly in marking. 


TorRODORA, n. g. 

Head smooth, sidetufts somewhat spreading ; ocelli absent ; 
tongue developed. Antenne 2,in ¢ serrate, ciliated (3-1), basal 
joint rather long, without pecten. Labial palpi long, recurved, 
second joint much thickened with appressed scales, somewhat 
rough beneath, terminal joint as long as second, slender, acute. 
Maxillary palpi very short. Posterior tibiz clothed with rough 
hairs, posterior tarsi sometimes with basal joint rough-haired 
above. Forewings with vein 1b furcate, 2 and 3 stalked from 
angle, 7 and 8 stalked, 7 to apex or just below, 9 out of 7 or 
in 2 sometimes separate, 11 from beyond middle. Hindwings over 
1, trapezoidal, hindmargin more or less sinuate, cilia }-2 ; veins 
3 and 4 from a point, 5 approximated to 4 at base, 6 and 7 stalked. 

Allied to Lecithocera. 'The three species described are 
very similar, but certainly distinct. T'ype, 7’. characteris. 


5. Torodora characteris, sp. n. 

¢o 9. 18—22 mm. Head and thorax dark fuscous. Palpi 
whitish-ochreous, second joint dark fuscous except apex. Antenne 
whitish-ochreous, sharply serrate, ciliations 1. Abdomen fuscous. 
Posterior tarsi with basal joint hairy. Forewings elongate, costa 
gently arched, apex obtuse, hindmargin sinuate, hardly oblique ; 
7 to apex ; rather dark fuscous ; a rather irregular black spot or 
small blotch on submedian fold before + of wing, connected with 
inner margin by an indistinct darker suffusion ; a small transverse 
black spot in disc beyond middle ; a faintly indicated slightly bent 
pale transverse line about 4: cilia fuscous, base pale ochreous, 
Hindwings rather light fuscous ; cilia as in forewings. 


Hab. oni; two specimens. 


of Lepidoptera from Upper Burina. 17 


6. Torodora parallactis, sp. n. 

d. 24mm. Differs from 7. characteris as follows: (terminal 
joint of palpi broken) antenne yellowish-tinged, less serrate, 
ciliations ?; posterior tarsi not rough-haired ; forewings with 
vein 7 to below apex, anterior blotch triangular, more distinctly 
connected with inner margin, followed by some whitish-ochreous 
scales, posterior spot absent, replaced by two transversely placed 
whitish-ochreous dots ; hindwings paler, cilia more ochreous. 


Hab. Mone; one specimen. 


7. Torodora ancylota, sp. un. 

2. 19mm. Head and thorax whitish-ochreous, tinged with 
pale brown-reddish. (Palpi and antenne broken.) Forewings 
elongate, costa slightly arched, apex strongly produced, pointed, 
hindmargin hence concave, rather oblique ; 7 to below apex, 9 out 
of 7 ; ochreous-fuscous, irrorated with dark fuscous; extreme 
costal edge yellowish ; a small black spot at base of costa ; an 
erect black bar from inner margin at 4, reaching 2 across wing, 
dilated on submedian fold ; a small black spot on costa at 2; two 
dark fuscous dots transversely placed in disc beyond middle ; a paler 
bent transverse line, preceded by a darker suffusion, faintly indi- 
cated at +: cilia pale fuscous, base whitish-ochreous. Hindwings 
with apex more pointed, hindmargin more sinuate than in the 
other species; light fuscous ; cilia as in forewings. 


Hab. Fort Stedman; one specimen. 
The peculiar shape of wing easily separates this species 
from the others. 


8. Lecithocera luticornella, Z. 
Hab. Koni; one specimen. 


9. Lecithocera pachyntis, sp. n. 

dg. 13—14mm. Head fuscous, sides of crown ochreous-yellow. 
Palpi pale yellowish-ochreous, sometimes partially suffused with 
fuscous. Antenne as long as forewings, lower half thickened with 
scales, much more strongly near base, ochreous-yellowish, upper 
half annulated with dark fuscous. Thorax and abdomen ochreous- 
fuscous. Forewings elongate, costa slightly arched, apex obtuse, 
hindmargin obliquely rounded ; vein 9 separate ; fuscous ; a black 
dot in dise at 1, and another at 2: cilia light ochreous, with two 
fuscous lines. Hindwings rather light fuscous ; cilia pale ochreous, 


Hab. Soni; two specimens. 
TRANS. ENT. SOC. LOND. 1894.—PaRT I. (MARCH.) B 


Ts. Mr. Edward Meyrick on a collection 


ZALITHIA, 0. &. 


Head smooth; ocelli present; tongue developed. Antenne Z, 
in ¢—(?), basal joint slender, without pecten. Labial palpi long, 
recurved, smooth, second joint rather thickened, terminal joint 
slender, acute. Maxillary palpi very short. Posterior tibizw with 
somewhat rough scales. Forewings with vein 1b furcate, 2 and 3 
stalked, 7 and 8 stalked, 7 to costa, 11 from middle. Hindwings 
over 1, oblong-ovate, cilia 3; veins 3 and 4 from a point, 5 parallel 
to 4, 6 and 7 closely approximated at base, 

10. Zalithia wranopis, sp. n. : 

9. 15mm. Head dark fuscous, with bright blue reflections, 
face brassy-ochreous. Palpi ochreous-yellow, anterior edge of 
terminal joint fuscous. Antenne dark fuscous, spotted with 
whitish-yellowish. Thorax dark fuscous, with three blue lines. 
Abdomen dark fuscous, beneath pale whitish-ochreous. Forewings 
elongate, rather narrow, costa slightly arched, apex rounded, hind- 
margin rather obliquely rounded; orange-ferruginous, apical 2 
coppery-blackish ; markings bright metallic green-blue ; a streak 
along anterior half of costa ; a streak along submedian fold from 
base to middle of wing; a rather narrow fascia separating the 
ferruginous and black portions, interrupted below middle and not 
reaching inner margin ; an irregular apical fascia, broken into spots 
on lower part of hindmargin: cilia fuscous, basal half orange- 
ferruginous round apex, with a black subbasal line. Hindwings 
dark fuscous, darker and somewhat coppery-tinged on posterior 
half ; cilia blackish-grey, terminal half whitish except towards apex 
and anal angle. 


Hab. Koni; one specimen. 


TipHa, Walk. 

Head smooth ; ocelli present; tongue developed. Antennzx 
with more or less strong thickening or nodosity near base, rough- 
scaled above. Labial palpi moderately long, curved, ascending, 
second joint with appressed scales, dilated aud somewhat tufted at 
apex, terminal joint very short (?), loosely scaled. Maxillary palpi 
very short. Posterior tibia rough-haired. Forewings with cell 
extremely narrow, 1b furcate, 2 and 3 stalked from angle, 4 absent, 
5 absent, 8 and 9 out of 7, 7 to costa, 11 from 4+. Hindwings over 
1, elongate-ovate, cilia }; in ¢ with membranous thickening along 
basal third of costa, with a tuft of long hairs lying along it on upper 
surface ; veins 3 and 4 stalked from angle, 5 absent, in © 6 and 7 


7 


of Lepidoptera from Upper Burma. 19 


stalked, in ¢ 6 absent and cell narrowed and pointed posteriorly, so 
that 7 rises from a point with stalk of 3 and 4. 


I do not possess Walker’s typical species, and the 
specimens of the following species are partly damaged ; 
hence the above generic characters are not quite complete, 
but sufficiently so -to show the great peculiarity of the 
structure. The genus is distinctly allied to Zalithia. 


11. Tipha helioclina, sp. n. 

d6 2. 20—22 mm. Head shining pale ochreous. Thorax metallic 
leaden-grey. Forewings elongate, narrow, costa gently arched, apex 
obtuse, hindmargin obliquely rounded ; bright orange ; a dark 
leaden-metallic streak along inner margin from ;{ to base, thence 
along costa to 1, whence it crosses the wing in a rather strong curve 
to middle of inner margin ; a large bronzy-purplish-fuscous patch 
occupying apical area, its anterior edge forming a very strong 
rounded angulation which reaches to middle of wing ; cilia brassy- 
metallic. Hindwings and cilia rather dark purplish-fuscous. 


Hab. Fort Stedman; two specimens. 


XYLORYCTIDZ. 


ProcHorRYCcTIs, n. &. 

Head with appressed scales, sidetufts loosely spreading; ocelli 
present ; tongue developed. Antenne ?,in ¢@ bipectinated, to- 
wards apex simple, basal joint stout, without pecten. Labial palpi 
long, curved, ascending, with appressed scales, terminal joint 
shorter than second, acute. Maxillary palpi rudimentary. Posterior 
tibize clothed with long hairs. Forewings with vein 1b furcate, 
2 from 4, 7 and 8 stalked, 7 to hindmargin, 9 absent, 11 from beyond 
middle. Hindwings 1, trapezoidal-ovate, hindmargin sinuate, cilia $ ; 
veins 3 and 4 short-stalked, 6 and 7 approximated towards base. 


Nearly allied to Cryptophasa. 


12. Ptochoryctis eremopa, sp. n. 

¢. 17 mm. Head, palpi, antenne, thorax, and abdomen 
whitish-ochreous; antennal pectinations blackish. Forewings 
elongate, costa slightly arched, apex obtuse, hindmargin slightly 
rounded, rather strongly oblique ; whitish-ochreous, on posterior 
half thinly sprinkled with ochreous-brown ; inner margin indis- 
tinctly suffused with ochreous-fuscous : cilia ochreous-whitish, with 
two cloudy ochreous-brownish lines. Hindwings light grey ; cilia 
as in forewings. 

Hab. WKoni; one specimen. 


20 Mr. Edward Meyrick on a collection 


TRICHERNIS, n. &. 
Head with appressed scales, sidetufts loosely spreading and 
projecting between antenne ; ocelli present, tongue developed. 
Antenne ¢, in ¢ with long fine cilia (5), basal joint moderate, 
without pecten. Labial palpi long, curved, ascending, second 
joint thickened with appressed scales, terminal joint shorter than 
second, slender, acute. Maxillary palpi rudimentary. Posterior 
tibize clothed with long rough hairs. Forewings with vein 1b 
furcate, 2 from angle, 7 and 8 stalked, 7 to hindmargin, 11 from 
middle. Hindwings 1, oblong-ovate, cilia ? ; veins 3 and 4 stalked 
5 approximated to 3 at base, 6 and 7 stalked. 


13. Tricheriis centrias, sp. n. 

gd. 15mm. Head and thorax pale ochreous-yellowish. Palpi 
whitish-ochreous, second joint dark fuscous externally except at 
apex. Abdomen whitish-ochreous. Forewings elongate, moderate, 
costa gently arched, apex obtuse, hindmargin straight, rather 
oblique ; pale ochreous-yellowish ; a round black discal dot beyond 
middle: cilia pale ochreous-yellowish. Hindwings and cilia 
whitish-ochreous. 


Hab. oni; one specimen. 


DEPRESSARIAD&. 
14. Pheosaces torrida, sp. n. 

3 ¢. 22—25 mm. Head and palpi ochreous-yellow, terminal 
joint of palpi in ¢ 3, stout, in @ 3, slender. Thorax light 
ochreous-fuscous, in ¢ paler and more yellowish. Abdomen 
fuscous, margins yellowish. . Forewings elongate-oblong, costa 
moderately arched, apex obtuse, hindmargin nearly straight, rather 
oblique ; in ¢ unicolorous light ochreous-yellowish ; in ? rather 
light ochreous-brown, apex and costa more or less suffused with 
ochreous-yellow : cilia ochreous-yellowish. Hindwings rather 
dark fuscous, in ¢ lighter and somewhat suffused with ochreous- 
yellowish ; cilia ochreous-yellowish, in @ base suffused with 
fuscous. 


Hab. Soni; three specimens. 


Homosaczs, n. g. 


Head with appressed scales, sidetufts loosely spreading ; ocelli 
present ; tongue developed. Antenne +, in ¢ stout, filiform, 
simple, basal joint moderate, without pecten. Labial palpi long, 
curved, ascending, with appressed scales, terminal joint shorter 


than second, acute. Maxillary palpi rudimentary. Posterior 


of Lepidoptera from Upper Burma. 21 


tibie clothed with long rough hairs. Forewings with vein 1b 
furcate, 2 and 3 stalked, 7 and 8 stalked, 7 to hindmargin, 11 from 


before middle. Hindwings 1, elongate-ovate, cilia 2; veins 3 and 
4 from a point. 


Alhied to Phcosaces. 


15. Homosaces anthocoma, sp. n. 


29. 15-17 mm. Head and palpi yellow-ochreous, palpi 
externally infuscated. Antenne, thorax, and abdomen dark 
fuscous. Forewings elongate, moderate, costa gently arched, apex 
obtuse, hindmargin rather obliquely rounded ; dark fuscous, faintly 
purplish-tinged ; cilia dark fuscous, Hindwings and cilia dark 
fuscous. 


Hub. Soni; five specimens. 


PERIACMA, n. g. 

Head with appressed scales, sidetufts loosely spreading ; ocelli 
concealed ; tongue developed. Antennze almost 1, in ¢ filiform, 
simple, basal joint moderately long, without pecten. Labial palpi 
long, curved, ascending, thickened with appressed scales, second 
joint short, terminal joint thrice as long as second, pointed. 
Maxillary palpi rudimentary. Posterior tibiz clothed with rough 
hairs. Forewings with vein 1b furcate, 2 from near angle, 7 and 
8 stalked, 7 to apex, 1b from middle. Hindwings under 1, very 
elongate-ovate, cilia 1 ; veins 3 and 4 from a point. 


Specially characterised by the peculiar structure of the 
labial palpi. Type, P. ferialis. 


16. Periacma ferialis, sp. n. 

@. 13—14mm. Head, palpi, and thorax light orange ; second 
joint of palpi mixed with dark fuscous externally, terminal joint 
with a dark fuscous apical spot. Antennze ochreous-yellowish, 
spotted with dark fuscous. Abdomen grey, anal tuft whitish- 
ochreous. Forewings elongate, costa gently arched, apex round- 
pointed, hindmargin very obliquely rounded ; bright orange ; 
markings dark slaty-purplish ; a streak along submedian fold from 
near base to middle of wing, tending to coalesce with a spot above 
its middle, and an oblique bar from inner margin near base ; a 
narrow slightly curved fascia from 2 of costa to anal angle ; an 
apical spot, sometimes ill-marked ; cilia orange, becoming pale 
whitish-ochreous towards anal angle. Hindwings grey; cilia 
whitish-ochreous, towards base greyish-tinged, 


Hab. Koni; two specimens. 


22 Mr. Edward Meyrick on a collection 


17. Periacma orthiodes, sp. n. 

g. 15 mm. Head, palpi, and thorax rather deep ochreous- 
yellow. Antenne grey. Abdomen ochreous-yellowish. Forewings 
elongate, moderate, costa gently arched, apex obtuse, hindmargin 
nearly straight, oblique; orange-yellow ; markings rather dark 
purplish-fuscous ; an oblique spot from inner margin near base; a 
streak along submedian fold from near base to middle of wing, 
with a spot above its middle ; a narrow slightly curved fascia from 
2 of costa to anal angle ; a hindmarginal fascia, moderate at apex, 
narrowed to a point at anal angle; cilia orange-yellow, tips whitish- 
yellowish, at anal angle purplish-tinged. Hindwings grey; cilia 
yellowish, towards anal angle greyish. 

Hab. Yoni; one specimen. Very similar to the 
preceding, but certainly distinct by the different form of 
wing, unspotted palpi, grey antenne, and _ yellow 
abdomen. 


18. Periacma chlorodesma, sp. un. 


&. 18 mm. Head orange-yellow, face and palpi whitish- 
ochreous. Antenne dark grey, base whitish-ochreous. Thorax 
rather dark purplish-grey, Abdomen grey, anal tuft whitish- 
ochreous. Forewings elongate, costa moderately arched, apex 
obtuse, hindmargin oblique, slightly rounded ; rather dark fuscous, 
slightly purplish-tinged ; a whitish-ochreous streak along anterior 
half of costa, meeting a narrow straight ochreous-whitish fascia 
which runs from costa beyond middle to inner margin before anal 
angle ; a suffused ochreous-yellow streak along apical portion of 
costa and upper half of hindmargin ; cilia ochreous-yellow, at anal 
angle dark fuscous. Hindwings grey; cilia light ochreous-yellowish. 


Hab. Koni; one specimen. 


GCOPHORIDA. 


19. Psecadia hilarella, Walk. 
Hab. Fort Stedman. 


BLASTOBASID®, 


ARCTOSCELIS, n. @. 


Head smooth ; ocelli absent ; tongue developed. Antennse 4, in 
g serrulate, shortly ciliated, basal joint moderate, without pecten. 
Labial palpi long, curved, ascending, second joint with appressed 
scales, somewhat rough beneath towards apex, terminal joint 


of Lepidoptera from Upper Burma. 23 


shorter than second, slightly roughened in front, acute. Maxillary 
palpi obsolete. Posterior tibiz and basal joint of tarsi with 
extremely long spreading erect spatulate hair-scales. Forewings 
with vein 1b furcate, 2 from 4, 7 and 8 stalked, 7 to hindmargin, 
11 from before middle. Hindwings 3, lanceolate, cilia 2; veins 
3and 4 stalked, 5 absent. 


20. Arctoscelis epinyctia, sp. n. 

g. 11 mm. Head ochreous-white, crown sprinkled with 
fuscous. Palpi whitish, externally mixed with fuscous. Antenne 
ochreous-whitish: Thorax dark fuscous, somewhat mixed with 
whitish. Forewings elongate, rather narrow, costa moderately 
arched, apex pointed, hindmargin extremely obliquely rounded ; 
fuscous, closely irrorated with dark fuscous ; a white dot in disc 
before =: cilia rather dark fuscous. Hindwings light fuscous, 
closely irrorated with dark fuscous ; cilia rather dark fuscous, 


Hab. Koni; one specimen. 


ELACHISTIDA. 
21. Cosmopteryx asiatica, Stt. 


Hab. Fort Stedman. 


PLACOPTILA, n. &. 


Head smooth ; ocelli absent ; tongue developed. Antenne almost 
1, in ¢ shortly ciliated, basal joint elongate, without pecten. 
Labial palpi very long, recurved, smooth, slender, terminal joint 
longer than second, acute. Maxillary palpi rudimentary. Abdomen 
in ¢ with claspers very large, exposed, their base covered above 
by a flat plate of expanded scales. Posterior tibize smooth-scaled, 
with whorls of scales at origin of spurs. Forewings with vein 1b 
furcate, 2 from 4, 6 absent, 7 to costa, 11 from middle. Huind- 
wings 3, narrow-lanceolate, cilia 4 ; veins 6 and 7 stalked. 


Nearly allied to Cosmoptery”. 


22. Placoptila electrica, sp. n. 

g@. 11—12 mm. Head, palpi, and thorax dark silvery-grey ; 
eyes crimson. Antennz and abdomen dark grey. Forewings 
elongate, very narrow, pointed ; blackish ; markings violet-silvery- 
metallic ; base suffused with dark silvery-grey ; a straight slender 
fascia at 4, dilated below middle ; two small marginal opposite spots 


at 2; a small spot on anal angle, and another beyond it on costa ; 


24 Mr. Edward Meyrick on a collection 


space between these and preceding pair sometimes distinctly 
bronzy ; cilia blackish, at apex with a white spot on tips. 
Hab. Koni; four specimens. 


HYPONOMEUTID-A. 
23. Cerace stipatana, Walk. 
IIab. WKoni. I believe this genus is usually referred 


to the Tortricina, but it properly belongs here, and is 
allied to Oeta. 


Comocritis, n. ¢. 

Head with appressed scales, sidetufts loosely spreading, pro- 
jecting between antenne; ocelli present; tongue developed. 
Antenne 2, in ¢ shortly bipectinated, with streak of rough scales 
on back near base, basal joint large, with dense pecten. Labial 
palpi moderate, curved, ascending, second joint loosely scaled, 
terminal joint less than half second, pointed. Maxillary palpi 
very short, filiform. Posterior tibiz clothed with rough scales. 
Forewings with vein 1b furcate,2 from near angle, 7 to costa, 
11 from middle, with a subhyaline groove on lower surface beneath 
12 towards base. Hindwings 1, elongate-ovate, cilia } ; veins 3 and 
4 from a point, 5 and 6 somewhat approximated at base. 


Allied to Oeta, but not closely. 


24. Comocritis olympia, sp. 0. 

dé. 30mm. Head, palpi, and thorax white. Abdomen grey. 
Forewings elongate, moderate, costa rather strongly arched, apex 
rounded, hindmargin obliquely rounded ; rather light ochreous- 
grey, densely mixed with blackish-grey ; a white basal fascia ; a 
broad white costal streak throughout, suddenly narrowed near base, 
lower edge cloudy and shaded off with blue- whitish, interrupted by 
a light ochreous spot on costa at > ; alongitudinal yellow-ochreous 
patch, marked with blackish lines on veins, extending through 
lower part of disc from near base to }; a crescentic white spot in 
disc at 3, and a second, slightly ockreous-tinged, at 3; a triangular 
white spot on anal angle: cilia white, with a light fuscous patch 
on lower part of hindmargin. Hindwings grey, apex white ; cilia 
grey, round apex white. 

Hab. ‘Koni; one specimen. 


PLUTELLIDZ. 


25. Plutella cruciferarum, Z. 
Hab, Koni. 


of Lepidoptera from Upper Burma. 25 


GRACILARIAD. 


26. Gracilaria chalcanthes, sp. n. 

6. 12—13mm. Head shining whitish-ochreous, mixed with 
ochreous, face silvery-white. Palpi white. Antenne white, 
annulated with dark fuscous. Thorax pale ochreous-yellowish, 
Abdomen pale grey. Anterior and middle legs dark fuscous, tarsi 
white ; posterior legs white, femora with a dark fuscous_ band, 
Forewings elongate, very narrow, parallel-sided, pointed ; ochreous, 
towards costa tinged, or costal ? wholly suffused with light brassy- 
yellow; margins marked with scattered black dots, mostly minute ; 
a black dot in dise before +, and another beyond middle : cilia pale 
ochreous-greyish, round apex ochreous, with rows of black points 
on terminal half. Hindwings rather dark grey ; cilia light grey. 


Hab, Koni; two specimens. . 


ANTIOLOPHA, 0. 2. 


Head smooth; no ocelli; tongue developed. Antenne 12, 
basal joint moderate, without pecten, Labial palpi moderately 
long, arched, subascending, second joint smooth, terminal joint as 
long as second, acute, with a triangular tuft of scales in front above 
middle. Maxillary palpi moderately long, slender, arched, por- 
rected, acute. Middle tibiz thickened with scales; posterior 
tibize rough-scaled towards apex. Forewings with vein 1b furcate, 
2 from ¢, 7 to costa, 8 absent, 11 from 4+. Hindwings 2, linear- 
lanceolate, cilia 5 ; veins 5 and 6 stalked. 

Nearly allied to Graciluria, from which it differs essen- 
tially by the tufted terminal joint of palpi. 


27. Antiolopha hemiconis, sp. un. 


?. 8—) mm. Head and thorax white. Palpi white, apex of 
second and tuft of terminal joint black. Antenne white, annu- 
lated with fuscous, Abdomen grey. Legs ochreous, anterior and 
middle tibiz suffused with blackish, tarsi white with cloudy 
ochreous bands and apex of joints blackish. Forewings elongate, 
very narrow, parallel-sided, pointed ; light reddish-brown ; anterior 
half of inner margin suffused with ochreous-whitish ; a triangular 
blotch on middle of costa faintly outlined with blackish scales, and 
more or less distinctly tinted with whitish : cilia grey-whitish, on 
hindmargin with basal half pale brown-reddish, and four rows of 
black points on terminal half. Hindwings rather dark grey ; 
cilia light grey. 

Hab, Koni; two specimens. 


26 Mr. Edward Meyrick on a collection 


TINEID. 


28. Adela satrapodes, sp. n. 


9. 16mm. Head ferruginous, face indigo-blue. Palpi and 
antennx violet-black. Thorax metallic indigo-blue. Forewings 
moderate, posteriorly dilated, costa moderately arched, apex 
obtuse, hindmargin very obliquely rounded ; deep shining indigo- 
blue; a broad bright orange fascia before middle, finely black- 
margined ; some scattered black scales beyond this, tending to 
form a transverse parallel line : cilia indigo-blue, tips round apex 
violet. Hindwings deep purple ; cilia bronzy-fuscous, 


Hab. Fort Stedman; two specimens. 


29. Melasina invariella, Walk. (Torna). 
Hab. Koni. 


30. Melasina ochrocoma, sp. n. 


a2. 22—26 mm. Head pale yellow-ochreous, face fuscous. 
Palpi dark fuscous, apex whitish ochreous. Antenne fuscous, 
pectinations in ¢ 6, in @ 2. Thorax and abdomen fuscous. 
Forewings rather elongate, somewhat dilated posteriorly, apex 
obtuse, hindmargin rather oblique ; light fuscous, mixed with 
whitish-fuscous and darker fuscous, tending to form faint 
strigule ; a darker fuscous suffusion tending to form a_ basal 
patch, a transverse spot from inner margin beyond middle, and a 
fascia from costa beyond middle to anal angle, but all very faint 
and sometimes obsolete: cilia whitish-fuscous, with a cloudy 
fuscous line. Hindwings fuscous, in @ paler; cilia as in fore- 
wings. 

Hab. Joni, Mone; six specimens. 


Tt belongs to the group called Alavona by Walker, 
which I do not think sufficiently distinct to be separated 
from Melasina; from its nearest allies it is separated 
by the yellowish head. 


Tatsizima, Walk. 


Head rough-haired ; ocelli present ; tongue obsolete. Antennze 
over 1, broadly and flatly compressed, joints closely set, simple in 
both sexes, basal joint short, without pecten. Labial palpi rather 
long, curved, ascending, second joint with dense more or less 
roughly projecting scales or hairs beneath, with some long bristles 
near apex, terminal joint shorter than second, with appressed 


of Lepidoptera from Upper Burma. 27 


scales, pointed. Maxillary palpi obsolete. Posterior tibiew shortly 
rough-scaled. Forewings with vein 1b furcate, 2 almost from 
angle, 7 to apex, 11 from before middle. Hindwings 1, elongate- 
ovate, cilia 2; veins tolerably parallel. 


31. Thisizima ceratella, Walk. 
Hab. Koni. 


32. Thisizima antiphanes, sp. n. 


¢. 13 mm. Head, palpi, and antenne pale yellow-ochreous. 
Thorax and abdomen dark fuscous. Forewings elongate, costa 
moderately arched, apex rounded, hindmargin obliquely rounded ; 
dark fuscous, with coppery-purple reflections ; a moderate straight 
yellowish-white fascia from + of costa to 2 of inner margin, some- 
what widened downwards ; a moderate yellowish-white spot on 
costa before apex : cilia dark fuscous. Hindwings and cilia dark 
fuscous. 


Hab. Mone; one specimen. 


33. Scardia tholerodes, sp. n. 


g. 16—20 mm. Head light ochreous-yellowish. Palpi light 
yellowish, mixed with dark fuscous, second joint with short rough 
projecting scales. Antenne whitish-fuscous, towards base darker. 
Thorax fuscous mixed with dark fuscous and whitish-ochreous. 
Forewings elongate, costa moderately arched, apex rounded, hind- 
margin very obliquely rounded ; 7 and 8 stalked, 9 absent ; whitish- 
ochreous, with scattered strigule of mixed fuscous and dark fuscous 
scales; the confluence of these forms a suffused basal patch, an 
oblique cloudy fascia before middle, and a fascia from beyond 
middle of costa to anal angle: cilia whitish-ochreous mixed with 
dark fuscous. Hindwings pale grey, indistinctly irrorated with 
darker ; cilia whitish-ochreous, with a cloudy grey line. 


Hab. ‘Koni; three specimens. 


34. Blabophanes monachella, Hb. (longella, Walk.). 
Hab. (Koni. 


It appears to me that the head of Walker’s longella is 
not yellow as alleged, and that it does not in any way 
differ from monachella, which I have also recorded from 
the Hawaiian Islands. 


28 Mr. Edward Meyrick on «@ collection 


385. Tinea platyntis, sp. n. 

od ?. 15—20 mm. Head light ochreous-orange. Palpi dark 
fuscous mixed with ochreous. Antennze whitish-ochreous. Thorax 
light ochreous, anteriorly purplish-tinged, Abdomen light ochreous. 
Forewings elongate, moderate, costa moderately arched, apex round- 
pointed, hindmargin very obliquely rounded ; yellow-ochreous ; 
costal edge infuscated towards base: cilia yellow-ochreous. Hind- 
wings bronzy-grey ; cilia whitish-grey. 

Hab. Mone, Koni; three specimens. Also from 
Mooltan in the Punjab. 


36. Tinea liomorpha, sp. n. 


¢d. 14mm. Head light ochreous-yellowish. Palpi fuscous, 
Antenne whitish-ochreous. Thorax fuscous-purplish. Abdomen 
light greyish-ochreous. Forewings elongate, rather narrow, costa 
moderately arched, apex roundpointed, hindmargin very obliquely 
rounded ; purplish-fuscous: cilia purplish-fuscous. Hindwings 
with veins 3 and 4 from a point ; light fuscous, with brassy re- 
flections ; cilia pale whitish-fuscous, 


Hab, Mone; oue specimen. 


The neuration of the hindwings is exceptional, but 
perhaps not constant. 


SARIDOSCELIS, n. g. 


Head rough-haired, face smooth; ocelli present; tongue 
developed. Antenne 3, basal joint moderate, with pecten. Labial 
palpi rather long, slightly curved, porrected, slender, anterior edge 
slightly rough, terminal joint as long as second, pointed. Maxillary 
palpi short, porrected, filiform. Anterior tibiz and tarsi clothed 
with long rough scales above ; posterior tibie smooth. Forewings 
with vein 1b furcate, 2 from near angle, 7 to hindmargin, 10 absent, 
11 from before middle. Hindwings 1, elongate-ovate, cilia + ; veins 
tolerably parallel. 


Allied to Calantica. 


37. Saridoscelis sphenias, sp. n. 

Q. 12mm. Head, palpi, antenne, and thorax white, Fore- 
wings elongate, costa moderately arched, apex tolerably acute, 
hindmargin sinuate, oblique ; white, partially finely sprinkled with 
pale fuscous ; a few scattered black scales ; two oblique wedge- 
shaped fuscous spots, anteriorly margined with dark fuscous, on 


of Lepidoptera from Upper Burma. 29 


inner margin in middle and before anal angle, emitting from apex 
two partially obsolete and hardly traceable very oblique dark 
fuscous lines, first apparently sharply angulated near costa, where 
it forms a short distinct oblique mark at 2, second running to hind- 
margin beneath apex; a short black interrupted dash before apex : 
cilia white, with two lines of fuscous or dark fuscous points. Hind 
wings and cilia grey. 


Hab. \koni; one specimen. 


38. Crobylophora daricella, Meyr. 
Hab. Mone; four specimens. 


There is no doubt as to the identity of this minute 


Australian species, whose occurrence somewhat surprised 
me. 


wae 
Pile ‘ ‘pot 
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Cha, Ui 
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eeu.) 


Il. Further observations on the Tea-bugs (Helopeltis) of 
India. By Cuartes Owen Warernovse, F.W.S. 


[Read Dee. 6th, 1893.] 


In the year 1886, I read before this Society some 
descriptions and observations on the genus LHelopeltis 
(Trans. Ent. Soc. Lond., 1886, p. 457), and in 1888 I 
supplemented this by the description of another species 
(Trans. Ent. Soc. Lond., 1888, p. 207). Until quite 
recently I had only seen a single female example of 
Helopeltis theivora. Iam glad to say the British Museum 
has now received a fair series of this species, with both 
sexes, and I can therefore give a description of the male. 


Helopeltis theivora. 

¢. Black. Front of the head very pale yellow on each side. 
Antenne black, or nearly so, the basal joint obscure fusco-testa- 
ceous mottled with brown, very pale yellow at the extreme base. 
Temora nearly black, mottled with fuscous-testaceous, with a 
pale ring at the base. Tibiz nearly black, mottled with fuscous- 
testaceous. Scutellar horn about as long as the length of the 
pronotum, very distinctly curved, with the usual cup-shaped 
enlargement at the apex. Some examples show a little yellow at 
the base. Length 23 lines. 


From this description it will be seen that the male 
differs chiefly from the female in having the pronotum 
black. The antennz and legs are also darker, and the 
whole insect is more slender than the female. The 
scutellar horn is a trifle shorter and a little less curved 
than in the female. 

Many of the females have the dark brown at the 
base of the pronotum extending across the whole base. 


The following is an extract from the letter from Mr. 
Frank Austen, who sent the specimens. It is dated 
from Sylhet, July 1893 :— 


‘The other day I sent you, per parcel post, a small 
box, containing a bottle in which are specimens of what, 
in the tea districts at any rate, is known as mosquito 
blight. It is to be found in many of the low-lying 


TRANS. ENT. SOC. LOND. 1894.—Parr i. (MARCH.) 


32 Mr. C. O. Waterhouse on the Tea-bugs of India. 


gardens such as this, and from the damage it does is one 
of the, if not the most, serious pests the tea bush has to 
contend against. It attacks the young shoots and 
appears to puncture them so that they shrivel up and 
iurn quite black. As yet there is no known cure. 
The method we adopt here is to employ a large 
force of small children, catching the insects by hand. 
This at best can only check it at the beginning of the 
season, for when acres and acres of tea are black with 
it, the thousands of insects brought in every day seem 
to make no difference. It is first noticeable about the 
beginning of the rains, i.v., the early part of June, and 
continues to increase until the end; the bushes attacked 
by it becoming gradually blacker, until after about the 
end of August or middle of September, they yield little 
or no leaf at al]. There is apparently no other plant it 
attacks, even in the jungle, and as far as one can see 
there is nothing that preys upon it. Wet dull weather 
is especially favourable to its propagation. It does not 
appear to fly much, and in the cold weather, after the 
bushes are pruned, there is not a sign of it. We always 
burn all the prunings, but it is doubtful whether much 
good is gained by so doing. It seems to appear 
spontaneously, first of alla bush here and a bush there, 
often acres apart, is attacked by it, then it gradually 
spreads.” 

Referring to Wood-Mason’s suggestion (‘‘ Report on 
the Tea-bug of Assam,” 1884, p. 18) that the indigenous 
tea-plant is not subject to the attacks of Helopeltis, Mr. 
Austen writes, “This is quite a fallacy, as this garden 
Maguracherra, consisting of some 450 acres, is composed, 
roughly speaking, of half hybrid and half indigenous, and 
last year the indigenous flats were badly blighted as well 
as the hybrid.” . . . “There was also another idea 
that heavy pruning eradicated the Helopeltis ; but last 
cold weather we cut back 50 acres of hybrid tea to a 
standard of i8 inches (the average height of a tea-bush 
is, say, 3 feet to 3 feet 6 inches), and this year this part 
of the garden was the first to get blighted, and is by far 
the most blighted part of the garden ; there could have 
been very few leaf-buds left on the bushes. Also, each 
bush had its roots carefully forked round, and then the 
whole place was hoed te a considerable depth.””—Letter, 
Nov. 28th, 1893. 


O88t") 


III. Notes on some Lepidoptera received from the 
neighbourhood of Alexandria. By Gerorae T, 
Bernune-Baker, F'.L.8. 


[Read Dec. 6th, 1893. ] 
PLATE I, 


Last year I had handed to me a collection of Lepidoptera 
from Alexandria for examination and determination, and 
during the present summer I had another collection also 
sent me, so that the two lots will give us some idea of 
the insects from this neighbourhood, and I therefore 
make no apology for cataloging the whole collection in 
detail, following the order of Staudinger’s list. The only 
thing that strikes me as worthy of special note, is the 
entire absence of any species belonging peculiarly to the 
Ethiopian region ; with sucha splendid migratory channel 
as the river Nile, this is not what I should have 
expected. 


1. Anthocharis belia, Cr. 
Two small but otherwise quite typical specimens. 
2. Colias edusa, F., and var. Helice, Hb. 
The type species not rare, and, as usual, a few of the 
white variety appear with it. 
3. Deudorix livia, Klug. 


Common. I have some dozen specimens in_ both 
Sexes. 


4, Thestor ballus, F.} 
One specimen quite typical, but small. 


5. Lycena boetica, L. 
Not uncommon, the blue suffusion of the 9 is un- 
usually bright in some of the specimens. 
6. Lycena egyptiaca, sp.n. (PI. [, fig. 1.) 
¢. Upper side, primaries and secondaries dull violet-blue, very 
TRANS. ENT. SOC. LOND. 1894,—PART I. (MARCH.) C 


34 Mr. George T. Bethune-Baker’s notes 


shghtly lustrous, outer margins very narrowly black. Secondaries 
with a blackish spot between the first and second median nervules, 
and another at the anal angle ; tail blackish. Underside, primaries 
and secondaries ochreous-cream colour, with transverse irregular 
fasciz margined with white, as follows :—Primaries, two across the 
cell followed by two across the centre of the wing, reaching almost 
to the inner margin ; between the two submarginai fascie is a short 
one from the costa to the central disco-cellular branch. Secondaries 
crossed by about eight whitemargined fasciz ; some much broken 
and transfused. There are two brilliant metallic green spots, 
pupilled with bluish-black and margined with yellow, at the anal 
angle. Margins of both primaries and secondaries finely fuscous, 
Exp. alar. 23 mm. 

2. Brownish, basal and discal area irridescent-blue, the under- 
side pattern showing through by transparency, and being decidedly 
intensified in the primaries; the black anal spot is composed 
of two confluent ones, not one only, as inthe ¢. The markings of 
the underside are similar to those of the ¢. Exp. alar, 23-24 mm. 

The wings of both sexes are somewhat transparent, 
but more so in the 9 than in the d. 


7. Lycena lysimon, Hb. 
Several typical specimens. 


8. Vanessa atalanta, L. 


Two specimens, one with the white dot in the red 
band. 


9, Vanessa cardut, L. 
One specimen. 


10. Danais chrysippus, L. 
Common. Most of the specimens fine. 
11. Hesperia mathias, Fab. (Thraz, F.) 
Three typical specimens. 


HETEROCERA. 


12. Acherontia atropos, L. 
One specimen. 


13. Sphinx convolvuli, L. 


The few specimens of this insect are most unusually 
small. 


on some Lepidoptera from Alexandria, 39 


14. Deilephila livornica, Esp. 
Specimens decidedly below the average size. 


15. Deilephila celerio, L. 
Typical, but possibly rather darker than usual. 


16. Deilephila nerii, L. 
Common. ‘The specimens sent are fine and of beautiful 
colouration. 


17. Harias insulana, B. 


Very common and very variable, ranging from beautiful 
uniform green to yellow and straw colour. 


18. Nola squalida, Steyr. 
One specimen only. 


19. Cletthara hittora, sp.n. (PI. I., fig. 2.) 


Primaries ochreous-grey, basal third dark brownish-grey bor- 
dered externally by a lighter band, which is edged by a perpen- 
dicular wavy line of black with an internal edge of whitish ; the 
black line extends from costa straight across to the inner margin ; 
the central area is occupied by a patch of pale reddish-brown, bor- 
dering which, on the posterior edge, is a trace of a very interrupted 
dark line, rising at the centre of the costa and being produced 
outward round this reddish-brown patch, and then receding to the 
internal margin; there is a submarginal band of pale reddish-brown 
shading, edged internally by a toothed interrupted blackish line 
arising in a small apical black patch. Posterior margin finely 
blackish. Fringes brownish-grey with darker dividing line. 
Secondaries grey subhyaline with a dark border. Fringes whitish 
with darker dividing line. Exp. alar. 17 mm. 

There is one specimen, which may be a 9, but the body is 
absent, in which the primaries are all dark grey; all the markings 
of the ¢ are reproduced, with the exception of the reddish-brown 
central area, but the pattern is very much intensified and darkened, 
and each of the dark transverse lines are accompanied by a pale 
border. Exp, alar. 18 mm. 


20. Detopeia pulchella, L. 
Common. 


36 Mr. George T. Bethune-Baker’s notes 


21. Ocnogyna loewit, L., var. Olathrata, Ld. 
Not uncommon. 


22. Cossus L-nigrum, sp.n. (Pl. 1, fig. 3.) 


Primaries ashen-grey, basal area with scarcely any marks or 
reticulations ; on the submedian vein is a short dark rich brown 
dash beginning near the centre, and from the end near the anal 
angle rises a thinner dash up to the lower median branch, forming 
an L-shaped sign, at the top of which is a small dark brown V ; 
the posterior half of the wings is reticulated and marked in the 
manner usual to the genus, but to a less extent than ordinary, 
whilst from a point on the costa about a third from the apex 
rises a dark brown waved stripe extending across the apical area, 
but curved, and reaching down to the anal angle; beyond this is 
another short line just in front of the apex, going across and 
touching the posterior margin. Costa darkly dotted. Fringes ash- 
grey. Thorax as primaries. Secondaries dark uniform brownish- 
grey without markings. Fringes grey. Abdomen paler than 
secondaries. Exp. alar. 36 to 39 mm. 


This species is, perhaps, nearest T’erebra, though not 
very near any of the genus that I have seen ; it can, how- 
ever, be immediately separated from Terebra by only 
being about half its size, and by the markless and 
uniform secondaries, 


23, Psyche (subgenus Manatha) hampsoni, sp. n. 
Primaries, veins la and b anastomosing ; vein 6 present in both 
wings, but not emitted in primaries from lower angle of cell, as in 
Viciella, Schiff., the type of the subgenus JJegalophanes ; veins 
4and 5 and 8 and 9 stalked. The shape of the wing is contorted 
and the costa excised. Nospine on the fore tibia. Wings are of 


a uniform luteous grey, head paler, thorax darker than ground 
colour. 
24. Orgyia dubia, Tausch., var. Judaea, Stgr. 
Not uncommon. 


25. Bombyx serrula, Gn. 
Not uncommon. 


26. Bombyx undata, Klug. 


One specimen, which I refer somewhat doubtfully to 
this species. 


on some Lepidoptera from Alewandria, 37 


27. Megasoma repanda, Gn. 
Common. 


28. Megasoma acacix, Klue. 


Of this beautiful species described by Klug I have 
three fine specimens, and also larvee well preserved. 


29. Bryophila fraudatricula, Hb., var. Pallida, nov. var. 
| (PI. I, fig. 4.) 

Primaries pale greenish-grey, the black transverse lines and 
black dash in the centre of the median area as in the parent 
species, but the short dash just above the anal angle is absent. 
Orbicular and reniform stigmata moderately distinct, outlined in 
blackish, the latter being filled in with pale grey. Both are much 
more distinct than in the original form. Secondaries grey, paler 
than in Praudatricula. Just beyond the centre the wing is 
crossed by a very distinct blackish line, following the course of 
the posterior margin, and extending from the costa to the inner 
margin. In the discal cell there is a dark grey spot. 


This is probably a good localform. I have three speci- 
mens all smaller than the type form, which agree closely 
inter se, but in one the median black dash is absent. It 
is evidently closely allied to Mraudatricula, so I have 
thought it better to make it a variety of that insect 
(from which it can be immediately separated by its very 
pale colour) rather than make it a new species. 


30. Agrotis pronuba, L. 
Not uncommon and quite typical. 


31. Agrotis puta, Ub. 
Common, the @ being unusually dark. 


32, Agrotis desertorum, B. 
One beautifully pale grey specimen. 


38. Agrotis aleaandriensis, sp.n. (PI. L, fig. 5.) 

g. Primaries yellowish-grey, first transverse line from discal 
cell to inner margin fawn colour, but rather indefinite, preceding 
which are three small black spots—one on the costa, one on the 
median vein, and a third on the submedian, the latter being nearer 
the base than the preceding ones ; there is a dark dot close to the 
base on the costa and median vein, The second transverse line 


08 Mr. George IT. Bethune-Bakevr’s notes 

beyond the reniform stigma is also fawn colour, extending from 
subcostal to submedian vein, but rather broken, the area between 
the first and second lines is tinged with fawn colour, beyond the 
second line is a curved row of dark dots from subcostal to 
submedian vein. Orbicular stigma almost obsolete. Reniform 
stigma outlined in fawn colour, with a similar coloured crescentic 
continuation below the extremity of the discal cell ; directly above 
this stigma the costa is darkly dotted. Posterior margin darkly 
dotted ; fringes yellowish-grey with paler extremities. Secondaries 
silvery grey. Fringes entirely snow-white. Exp. alar. 42 mm. 

@. Primaries uniform brownish-grey, slightly lustrous, no 
marking at all visible except a small darker spot representing the 
orbicular stigma; the reniform stigma is finely outlined and 
pupilled with darker grey. Posterior margin darkly dotted. 
Fringes whitish with dark extremities. Secondaries silvery-grey, 
brownish towards outer margin, slightly lustrous. Fringes silvery- 
grey. Exp. alar. 40 mm. 


I have also one specimen, probably the same species, of 
an uniform pale ochreous-fawn colour, the first and 
second line as in type, but the former extended up to the 
costa and rather broken, the latter being followed 
immediately by a curved row of very fine dark minute 
dots, from which to the posterior margin the ground 
colour is rather darker, interrupted only by a doubly curved 
paler transverse line from the costa to the anal angle. 

This species will follow Desertorwm, B. 


34. Agrotis ml, sp.n. (Pl. 1, fig. 6.) 

3. Primaries grey, the first line close to the base, rising in a 
black costal dot,isonly just traceable.; the second line, pale and waved, 
rises from a black costal spot, extends to the inner margin, though 
somewhat indistinct there, and is bordered externally by a fine 
blackish edge ; in the median area is a small narrow oval blackish 
loop adjoining the second line, and filled in with pale ochreous. 
Third line very dark grey, curved outwardly, rising ina black costal 
spot and reaching to the inner margin; this line consists of a series 
of minute distinct crescentic marks joined at the tips, and is edged 
by a fine line of slightly paler ground colour. Posterior margin 
finely edged with black, and preceded by a band of dark grey 
shading ; just in front of the apex there is also a patch of this dark 
grey shading. Orbicular stigma slightly ochreous-grey, very finely 
encircled, and minutely pupilled with blackish ; reniform stigma 


on sone Lepidoptera from Alexandria. 39 


dark grey edged with black. Between these two stigmata isa trace 
of an indefinite perpendicular band of dark grey shading right 
across the wing. Fringes grey with a dark dividing line. Thorax 
grey, abdomen paler grey. Secondaries and fringes white. Exp. 
alar. 38—39 mm. 

@. Primaries pale greyish-cinnamon-brown, first line close to 
base indistinct reddish-brown, second line waved reddish-brown, 
third line very indistinct, followed by a curved row of white dots, 
from whence the ground colour of the wing becomes abruptly 
darker up to the hindmargin, which is finely edged by a light line, 
and preceding which is a series of dark grey shadings between each 
nervule. Orbicular stigma distinctly encircled with reddish-brown, 
as is the reniform, the latter being filled in with greyish. Between 
these two stigmata there is a perpendicular band of dark reddish- 
brown shading right across the centre of the wing, and a similar 
coloured patch is in front of the apex. Fringes brownish-grey. 
Secondaries brownish-grey, darker near the hindmargin. Fringes 
pale grey. Thorax as primaries, abdomen as secondaries. Exp. 
alar. 39 mm. 


30. Agrotis suffusa, Hb. 
Not uncommon and quite typical. 


36. Mamestra afra, sp.n. (PI. L., fig. 7.) 

Primaries pale ochreous-grey, the first line close to the base is 
black, toothed, extending from the costa to the submedian vein, and is 
followed by a whitish stripe; second line in front of the orbicular 
stigma is composed of three crescentic dark marks starting from a 
dark grey costal spot, the central and inner crescents being preceded 
by a whitish edging. Orbicular stigma white, beneath which is a 
small dark crescent touching the middle of the second line and so 
making the letter q below the stigma. Reniform stigma margined 
with black, except on the costal side, with a basal inner margin of 
white and filled in with dark grey, except at the apical corner, 
where the colour is ochreous-grey. Subterminal line, consisting of 
a very strongly serrated fine dark line, bordered externally by 
white, is curved out beyond the reniform stigma, but terminates 
on the inner margin nearer the base than is that stigma. The 
terminal line close to posterior margin, strongly serrated, is white, 
edged internally by a dark shading, and with a dark patch opposite 
the reniform stigma. The hindmargin is conspicuously edged with 
dark crescentic marks between each vein. Fringes grey tessellated 
with white, and having a dark dividing line; on the costa are seven 


40 Mr. George T’. Bethune-Baker’s notes 


dark grey spots, four near the base and three between the two 
stigmata. 

Secondaries white, cell marked by a dark median vein and closed 
by a distinct dark crescent ; there is a broad dark grey band from 
the apex to the inner angle. Posterior margin edged by a fine dark 
line, which is preceded by a broad whitish band. Fringes white. 
Antenne finely serrated. Exp. alar. 33 mm. 


Tins species is allied to Stigmosa, Chr., but can be 
at once distinguished by the white secondaries with the 
crescentic termination of the cell. ‘The primaries are 
much greyer in colour, the transverse lines more pro- 
nounced, whilst there is no trace of the triangular spot in 
front of the apex. 


Qr 


37. Mamestra abbas, sp.n. (PI. 1., fig. 8,a¢, b?.) 

¢d. Primaries uniform brownish-grey, first two transversa lines 
slightly paler, and finely margined on each side with dark brown, 
the third pale line darkly margined internally only ; subterminal 
line fine, indistinct and interrupted, with three dark arrow-shaped 
marks emitted towards the base. Hindmargin darkly dotted. 
Orbicular stigma pale ochreous-grey, pupilled with the ground 
colour, and partly encircled with dark brown, beneath this is a 
distinct darkly outlined q-shaped sign. Reniform stigma pale 
brownish-grey outlined with dark brown and pupilled with paler 
grey, which pupil is margined with dark grey. Fringes brownish- 
grey with a paler dividing line. Secondaries brownish-grey, 
darker around the posterior margin. Fringes pale grey. Exp. 
alar. 35 mm. 

2. Primaries, ground-colour very pale ochreous-grey, pattern as 
in ¢. Secondaries very pale ochreous with a broad dark posterior 
margin. Exp. alar. 38 to 40 mm. 

This species will come next to Trifoliv; it is easily 
recognized by its quite uniform and duller colouration 
and markings, by its much darker secondaries, and by 
its exceedingly pale ¢. 


38. Heliophobus marsdeni, sp. un. (PI. I., fig. 9.) 
¢. Primaries brownish-ochreous, basal area much paler, the 
first transverse line in front of the orbicular stigma is dark brown, 
interrupted and edged internally with pale ochreous ; beyond the 
reniform stigma is a pale ochreous transverse dentated and inter- 
rupted line edged internally with dark brown; area between these 
lines darker than rest of wing: subterminal line pale ochreous, 


on some Lepidoptera from Alexandria. 41 


interrupted and edged internally with dark brown; posterior 
margin darkly dotted, orbicular stigma distinct, pale ochre, finely 
outlined with dark brown. Reniform stigma indistinct, grey, with 
a pale dash on the basal edge ; costa with three pale ochreous spots. 
Fringes lustrous ochreous-brown, Abdomen greyish white. 
Secondaries very pale grey, slightly darker on posterior margin. 
Exp. alar. 30 to 32 mm. Antenne very strongly ciliate, almost 
plumose. 

@. Similar to g, but the colours each intensified, 7.e., the dark 
areas are darker, the light are lighter ; all the markings much more 
pronounced, and the reniform stigma distinct. Antenne simple. 
Exp. alar. 28 mm. 


The ¢ differs somewhat in colouration, some being 
quite pale ochreous with dark central areas, others dark 
greyish ochre, and much more uniform in general tone; 
the pattern is however apparently quite constant. The 
primaries of ? are decidedly narrower than the ¢. 


This species will follow Orana of Lucas, but is easily 
recognizable by its more variegated appearance, visible 
in even the greyest and most uniform specimens; also by 
the basal line, the pale line beyond the reniform stigma, 
and the dark reniform itself. There is no pale patch at 
the apex as in Orana. 


The secondaries are very much paler, and the abdomen 
is whitish-grey instead of brown as in Lucas’s species. 
I have named this insect after Mr. Marsden of Alex- 
andria, through whose kindness I have received these 
collections. 


39. Miana trilinea, sp.n. (PI. 1., fig. 10.) 

Primaries ashen-grey, first line near base dark brown, strongly 
dentate on median vein; second dark brown line touching orbi- 
cular stigma internally ; third line just beyond the reniform, like- 
wise dark brown ; the area between the first and second lines is 
much paler grey. Just beyond the third line is a band of paler 
shading ; orbicular and reniform stigmata pale grey darkly out- 
lined. Posterior margin finely edged with dark brown. Fringes 
lustrous grey. Seccndaries pale grey, slightly lustrous, darker near 
posterior margin. Fringes paler grey. Thorax as_ primaries, 
Abdomen as secondaries. Exp. alar. 21} to 234 mm. 


40. Hriopus latreillii, Dup. 
A common insect of the ordinary type. 


49 Mr. George T. Bethune-Baker’s notes 


41. Prodenia littoralis, B. 
Several specimens quite typical. 


42. Sesamia cretica, Ld. 
Two nice specimens. 


43. Caradrina exiqua, Hb. 
Not uncommon. 


44. Caradrina mediterranee, sp.u. (PI. L., fig. 11.) 

Primaries ashen-fawn colour with the least trace of a greyish 
subterminal transverse curved line. Just in front of the posterior 
margin is an indistinct row of very pale ochreous-white spots, the 
middle spots having their inner margin dusted with golden-brown. 
Orbicular stigma obsolete, reniform stigma small dark brown 
encircled with pale ochreous. Costa with four indistinct black 
spots. Fringes lustrous greyish-fawn colour, Secondaries white, 
lustrous, very slightly greyish near posterior margin. Fringes 
lustrous greyish, Exp.alar. ¢ 30mm.; 9 25 mm. 


This species is perhaps nearest Quwadripunctata, but at 
once separable by the almost entire absence ef markings 
and by its much paler and cleaner appearance. From 
Flava of Fallow it is recognizable by the same 
characteristics. 


45. Cucullia chamomille, Schiff. 


I have four specimens of this genus, which are doubtful, 
but appear to be a very dwarfed race of this species, and 
I therefore place them provisionally under this name. 


46. Plusia chalcites, Esp. 


Two ordinary specimens. 


47. Plusia gamma, L. 
Not uncommon. 
48. Heliothis peltigera, Schiff. 
Not rare. 


49. Heliothis nubigera, H. 8. 
One specimen only. 


on some Lepidoptera from Alexandria. 43 


50. Heliothis armigera, Z. 
One specimen only. 
51. Hrastria scitula, Rbr. 
One specimen. 
52. Thalpochares phoenissa, Ld. 
Two typical specimens. 
53. Thalpochares ostrina, Hb. 


Similar to European specimens. 


54. Thalpochares parva, Hb. 
T'wo specimens quite typical. 
5d. Cerocala scapulosa, Hb., var. Insana, H. 8. 
One specimen. 


56. Pericyma albidentaria, Frr. 


I have two specimens of this insect, which form a 
sort of ‘ transit” to the var. Squalens, Ld. 


57. Leucanitis stolida. 
One specimen of the usual form. 


58. Grammodes bifasciata, Petag. 
Not uncommon. 


59. Pseudophia wlunaris, Hb. 
4 common insect. 


60. Pseudophia benenotata, Warren. 


A common species. In the National collection is the 
hitherto unique type which Warren described. It is a 
g, and as I have several 9 before me it may be advisable 
to describe this sex. 

@. Primaries pale ochreous-grey, with perhaps the least trace 
of pink ; both first and second transverse lines paler than ground 
colour, not darker asin the ¢; the subterminal pale line is much 
more distinct, and the shading on each side of it is paler than in 
the opposite sex. Secondaries slightly ochreous-grey with broad 
dark borders and very pale fringes. 'Thorax and abdomen as 
primaries, Exp. alar. 26 to 30 mm. 


AA, Mr. George IT’. Bethune-Baker’s notes 


Specimens vary somewhat in depth of colour, some 
being paler and others rather darker than the type. 


61. Spintherops exsiccata, Ld. 
Several specimens, calling for no remark. 


62. Hypena ravalis, H.S. 
Not uncommon. 


63. Hypena lividalis, Hb. 
Three ordinary specimens. 


64. Hypena obsitalis, Hb. 


A common insect. 


65. Arrarde parva, sp. nu. 

Primaries pale ochreous-brown, basal half decidedly darker than 
exterior half; the subterminal waved line is paler and bordered 
internally by a fine line of darker ochreous, and externally by a 
broadish band of the same coloured shading. Apex with a small 
patch of blackish scales, a few similar coloured scales being 
scattered near the anal angle. Costa near apex lightly spotted. 
Fringes ochreous-brown. Secondaries brownish-grey, darker near 
the outer margin, with pale grey fringes. Thorax as primaries. 
Abdomen rather darker than secondaries. Exp. alar, 14 mm. 


I have but one specimen of this insect, but it so 
evidently a distinct species that I do not hesitate to 
describe it, though from a single specimen. 


66. Nemoria faustinata, Mall. 
Two specimens of usual form. 


67. Acidalia coenosaria, Ld. 
One specimen. 


I have two other species of this genus, which I believe 
to be new, but I do not think it wise to describe them 
from single specimens, and I shall therefore await other 
specimens before naming them. 


68, Macaria aestimaria, Hb., var. Sareptanaria, Stgr. 
wo very handsome forms of this variety. 


on some Lepidoptera from Aleaandiia. 45 


69. Fidonia megiaria, Obr., var. Obscuraria, n. var. 
(Pl. ls ties, £2.) 

The specimens obtained around Alexandria are very 
dark indeed, and appear to be quite a local race; it is a 
common insect in the district. The ground colour is 
uniform dark greyish brown, the pattern being quite 
similar to that of Oberthur’s species, but very considerably 
darker. Out of ali the specimens before me the lghtest 
is very much darker in every respect than the darkest 
Algerian form that Ihaveseen. The Heyptian specimens 
are a very handsome variety of it. 


70. Sterrha sacraria, L., and var. Sanguinaria, Esp., 

and ab. Oranaria, Lucas. 

I have this species of all shades from Alexandria, from 
the palest forms up to var. Sanguinaria, also I have the 
duller and darker insect described by Lucas under the 
name Oranaria. 

I have no doubt whatever that Staudinger is right in 
his 1872 catalogue, where he says, on page 176, ‘an 
sequ. ab.” Sacraria being the following species. 

1 have preserved larvee of both forms before me which 
are quite similar, and I have also two very rich dark 
forms infinitely darker than Oranaria, the oblique stripe 
being very dark ash-grey, thus showing that the species 
varies very extensively indeed. 


71. Cidaria fluviata, Hb. 


Not uncommon. 


72. Hupithecia ultimaria, B. 
Two specimens, decidedly smaller than usual, 


73. Hupithecia distinctaria, H.S. 
Two pretty forms of this wide-spread species. 


74, Hupithecia (Gymnocelis) pumilata, Hb. 
T'wo or three ordinary specimens. 


75. Mnesivena quadripunctata, sp.n. (Pl. L, fig. 13.) 
Primaries brownish-grey, with a transverse pale line near the 
base from the costa to the inner margin, the line being composed 
of three small crescents ; the submarginal pale line, frequently and 
sharply serrated, recedes slightly from the apex to the subcostal 


46 Mr. George T. Bethune-Baker’s notes 


vein, where it curves outwards, receding again to the submedian 
vein, whence it recedes rapidly and considerably to the inner 
margin, and has a very fine dark edging. In the central area of 
the wing are four distinct whitish spots, around each of which are 
sprinkled, in fresh specimens, some dark brown rough scales, Hind- 
margin finely and darkly edged. Fringes pale grey. Secondaries 
whitish, with a fine submarginal dark grey stripe a little distance 
from the hindmargin, but following its course. Hindmargin finely 
and darkly edged. Fringes whitish. Antenne ¢ ciliate; 9 simple. 
Exp. alar. 23 mm. 


76. Mnesivena bella, sp.n. (PI. I, fig. 14.) 


Primaries pale-grey, irrorated with pale chestnut roughish 
scales ; basal area dark chestnut-brown with a patch of superim- 
posed rosy scales ; in the central area beyond the middle, near the 
inner margin, is another patch of rosy scales, having a dark 
chestnut basal border, which is edged by a white line, the inner 
margin beneath this patch being chestnut colour. Posterior margin 
broadly but sparingly irrorated with rosy scales, which also extend 
along the inner margin from the anal angle to the rosy patch just 
mentioned. From the costa near the apex, a fine indistinct 
transverse white line extends half across the wing, forming the 
internal border to the posterior rosy scales. Fringes greyish, 
irrorated with pink. Antenne ¢ ciliate. Secondaries whitish, 
with whitish fringes. Exp. alar. 19 to 23 mm. 


In specimens that have flown much the rosy scales get 
almost entirely rubbed off. 


77. Hypotia syrtalis, Rag. 
The specimens are typical. 


78. Hypotia bilinea, sp.n. (PI. I., fig. 15.) 

Primaries greyish-straw colour, central area bounded internally 
by an oblique dark brownish transverse line, and externally by a 
similar dark curved transverse line. In this area, beyond the 
centre, and near the costa, is a dark brownish-grey spot. Just 
before the posterior margin is a fine dark line, which in flown 
specimens is very indistinct. Secondaries very slightly paler than 
primaries, with a dark brownish-grey curved stripe beyond the 
centre, extending from the upper margin to the anal angle, and 
another similar stripe just in front of the posterior margin. 
Fringes ochreous-grey. Thorax as primaries, abdomen as secon- 
daries. Exp. alar. 16-18 mm. 


on some Lepidoptera from Alewandria. 47 


(on Lolissagra, so. 0.6 (PI 1.; fie. 16:) 

Primaries dark ashen-grey irrorated more or less with blackish 
scales. Across the centre of the wing is a transverse whitish 
stripe, the upper half of which is waved, the lower half being 
straight and edged externally with very dark brown. The second 
whitish transverse line near the posterior margin is very irregularly 
toothed and waved. Submarginal area with a broadish band cf 
whitish shading. Between these two lines in the central area on the 
internal margin and near the costa is a very distinct white spot 
darkly encircled. Posterior margin very distinctly dotted with 
black. Fringes shining brownish-grey. Secondaries dark brownish- 
grey with white fringes, which have a very dark and broad 
dividing line. Exp. alar. 235 mm. 


I have two specimens, one being paler than the type. 
80. Hellula undalis, F. 
Common and quite typical. 
81. Odontia dentalis, Schiff. 
Two small specimens. 
_ 82. Anthophilodes baphialis, Ld. 
Fairly common. 
83. Botys ferrugalis, Hb. 
Several specimens. 
84. Huwrychreon nudalis, Hb. 
Two specimens, just like those from Algeria. 
85. Orobena isatidalis, Dup. 
A common species. 
86. Margarodes unionalis, Hb, 
Two specimens. 
87. Metasia emiralis, Obr. 


Several specimens, which are somewhat yellower and 
the dark marks darker than the Algerian form. 


8&8. Synclera traducalis, L. 


Two specimens. 


48 Mr. George T. Bethune-Baker’s notes 


89. Duponchelia fovealis, Li. 
One or two ordinary specimens. 


90. Nymphula fusco-marginata, sp.n. (PI. 1, fig. 17.) 


Primaries greyish-brown, with a short interrupted dark dash 
from the roots just beneath the discal cell’; beyond the centre is a 
smallish dark brown circular sign, the circle being incomplete on 
the posterior side, beneath which is a trace of a cinnamon-brown 
transverse dash extending to the inner margin. Submarginal line 
white, edged externally with black, from where up to the posterior 
margin is a band of pale cinnamon-brown. Fringes greyish lustrous, 
with a dark dividing line. Secondaries whitish, with a large black 
spot on the upper part of the discal cell, whence to the inner 
marginruns a waved black stripe ; beyond this isa dark grey stripe, 
followed by a line of the ground colour, after which is a broad 
band of very dark grey, extending from the costa to the inner 
margin just over the anal angle; this is followed by a broadish 
stripe of the whitish ground colour edged externally bya fine 
dark line, the posterior margin itself being broadly bordered with 
pale cinnamon-brown, margin itself very finely black. Fringes 
whitish, with a fine black dividing line. Thorax as primaries, 
abdomen brownish. Exp. alar. 16 mm. 


9]. Schoenobius dodatellus, Walker. 


One specimen. 


92, Chilo dubia, sp.n. (Pl. L, fig. 18.) 

Primaries pale brownish-ochreous ; all along the median area 
are very sparsely scattered a few dark-brown scales, which also are 
extended in lines between all the nervules on the hind margin. 
Secondaries whitish, somewhat lustrous, with white fringes. 3 ¢ 
specimens. Exp. alar, 283—32 mm. 

In some specimens there is an indistinct darkish~ spot 
closing the discal cell. 


93. Ancylolomia palpella, Schiff. 
Not uncommon. 
04, Crambus alexandriensis, sp.n. (Pl. L., fig. 19.) 
@. Primaries pale-grey irrorated all over with greyish-brown 


scales, interspersed with patches of black irrorations ; about one 
third from the base is a whitish < shaped mark between the discal 


on some Lepidoptera from Alexandria. 49 


cell and the inner margin. The subterminal white transverse line 
is very sharply angulated outwards near the apex, whence it recedes 
in a wavy curved line to the inner margin; the central area is 
occupied by a patch of black scales edged externally with white, 
over which on the costa is another small blackish patch also, and 
in the cell is a small blackish dash over the < shaped white mark. 
The hind marginal area is dark grey, the posterior margin itself 
being edged with a fine metallic-greenish line. Fringes whitish, 
brown at extremities, and having a brown dividing line. Secon- 
daries greyish-white, slightly lustrous, with white fringes. Thorax 
greyish, abdomen paler grey. Exp. alar. 24 mm. 

The 9? isdarker with less marking, much less black scaling, and 
having the > shaped mark and the subterminal line only just traced 
out. Exp. alar. 22 mm. 


This species is not very near any Palearctic Crambus, and 
though | am acquainted with nearly all the species from 
our region, I cannot now place it in its correct position. 
Perhaps when I get more specimens (I have only one pair 
before me), I may be able to settle this point. 


95. Crambus.afra, sp.n. (PI. I., fig. 20.) 

Primaries pale fawn colour, the first very oblique and sharply 
dentate rich brown line rises beyond the centre of the costa and 
recedes sharply to a point about one third from the base on the 
inner margin, the subterminal line is the same rich reddish-brown 
colour, is sharply angulated outwards near the costa, and is broadly 
edged externally by the pale ground colour, the rest of the 
marginal area being dark mauvish-grey ; from the roots to the first 
line the whole of the median area is of a rich dark brown, broadly 
edged all round by the pale ground colour. Posterior margin 
darkly dotted. Fringes brownish-grey tesselated with fawn 
colour. Secondaries brownish-grey with whitish fringes, which 
have a fine dark dividing line. Thorax brown, patagiw fawn 
colour. Abdomen greyish. Exp. alar. 19 mm. 


This pretty species is nearest to Jucuindellus, H. 8, 
but is decidedly smaller, and can also be distinguished 
by the very dark and serrated lines, which are quite 
diferent in shape to those found in that species. 


96. Hromene ramburiella, Dup. 


One poor specimen. 
TRANS, ENT. SOC. LOND, 1894,—pParr I. (MARCH.) D 


50 Mr. George 'l'. Bethune-Baker’s notes 


97. Hvromene ocellea, Hw. 

Not uncommon apparently, one or two specimens 

having been in each of the two collections received. 
98. Myelois rhodochrella, H.S., and var. Hellenica, Stgr. 

One specimen of the type, and also one of its beautiful 

variety. 
99. Ematheudes punctella, Tr. 
One or two nice specimens. 


109, Melissoblaptes bipunctanus, Z. 
These specimens are fine, but call for no special remark. 


101. Crocidosema plebejana, Z. 
One specimen. 


102. Hapsifera luridella, Z. 
Several specimens. 


103. Scardia-mediterranee, sp.n. (PI. I. fig. 21.) 

Primaries slaty grey, a broadish longitudinal pale ochreous 
stripe extends from the base along the median area just beneath 
the cell, over which patches of dark brown rough scales are more 
or less scattered. Beyond the centre and near the costa is a short 
dash of the same pale cchreous, in the middle of which is a distinct 
spot of these dark brown superimposed scales. Costa palely spotted 
near the apex. Posterior margin darkly edged. Fringes dark grey. 
Secondaries brownish-grey, paler at the base. Fringes brownish- 
grey, tibize dark grey frequently encircled with pale ochreous. 
Hindlegs whitish. Exp. alar. 19 to 20 mm. 


Specimens vary slightly duter se, some having the 
costa palely spotted all along, and having a few pale spots 
in the subcostal area, which. gives such specimens a more 
variegated appearance. 


104. Tinea tapetzella, L. 
Two specimens. 
100. Agdistis tamaricis, Z. 
Two or three ordinary specimens. 


106. Alucita monodactyla, L. 
Two specimens. 


on some Lepidoptera from Alexandria. 


EXPLANATION OF Prare I. 


Lycena egyptiaca, sp. n. 

Cletthara littora, sp. n. 

Cossus L-nigrum, sp. 0. 

Bryophila fraudatricula, Hb., var, Pallida, nov. var, 
Agrotis alewandriensis, sp. n. 

Agrotis nili, sp. n. 

Mamestra afra, sp. n. 

da& Ob Mamestra abbas, sp. n. 

Heliophobus marsdeni, sp. u. 

Miana trilinea, sp. 0. 

Caradrina mediterranea, sp. 0. 

Fidonia megiaria, Obr., var. Obscuraria, nov. var. 
Mnesixena quadripunctata, sp. n. 

Mnesizena bella, sp. n. 

Hypotia bilinea, sp. n. 

Talis afra, sp. n. 

Nymphula fusco-marginata, sp. n. 

Chilo dubia, sp. n. 

Crambus alexandriensis, sp. n. 

Crambus afra,sp. n. 


Scardia mediterranée, sp. n. 


51 


vnrfns 
or 
jel) 
ow 


IV. The Rhynchophorous Coleoptera of Japan. Part III. 
Scolytide. By Watter F. H. Branprorp, M.A., 
EZ.s: 

[Read Dec. 6th, 1893.] 


The Rhynchophora collected by Mr. George Lewis in 
the Japanese Archipelago during the years 1880 and 
1881 have been described in part by Dr. Sharp in the 
‘Transactions’ of this Society for 1889 and 1891. In 
this paper I deal with the Scolytide of that collection, 
four species of which I have already described in my 
paper on the Scolyto-platypini. 

To the present time our knowledge of Japanese 
Scolytids rests upon the materials brought together by 
Mr. Lewis up to 1872, before which date but one species, 
Genyocerus adustipennis, Motsch., was known as Japanese. 
Nothing resembling that insect exists in this collection, 
and I have nothing to say about it. The species of the 
earlier collection were submitted, the Yomicini to 
Hichhoff, the remainder to Chapuis, and the results are 
given in the ‘ Annales de la Société entomologique de 
Belgique,’ 1874, pp. 195—203, in a paper entitled 
“ Scolytides recueillis an Japon par M. G. Lewis.” They 
distinguished 18 species, of which one occurred also in 
Kurope; the rest were new. In 1878 Hichhoff in his 
“ Ratio Tomicinorum ” added six species (one Huropean), 
and fully described those he had previously diagnosed. 

Nothing else has been written on the subject, and I 
have not found it necessary to redescribe the species of 
Chapuis and WHichhoff, and have merely indicated 
localities, etc., and some points of distinction between 
them and new species. ‘The original descriptions, and in 
the case of TVomicini the later ones of Hichhoff, are 
sufficient to identify them by. ; 

The number of species known to exist in Japan before 
my examination of this collection was 25; I raise it to 
104 by the addition of 79 species, of which 71 are new. 


TRANS. ENT, SOC. LOND, 1894.—PART I. (MARCH.) 


54 Mr. Walter F. H. Blandford on the 


Provided that I have correctly identified all the five 
new species described by Hichhoff in 1878, this collection 
contains all known Japanese species except Hylastes 
attenuatus, Xyleborus badius, and Genyocerus adusti- 
penis, and all types peculiar to Japan except of that 
insect and the five referred to. 

The number of species, 104,* is perhaps a little above 
the mark, because I have been obliged to describe under 
separate names three male Xylebori which cannot be 
referred to their respective females. Ina few cases I 
may have subdivided a species into two, but it is likely 
that these are counterbalanced by others where I have 
included distinct species as varieties. Students of the 
iiuropean forms know that species closely resembling 
each other in appearance may differ in habits, food- 
plants, and the form of their galleries. In dealing with 
an exotic collection one has to do without the assist- 
ance of such facts. 

They are divided into 25 genera, of which three are 
new, Hyorrhynchus, Spherotrypes and Acanthotomicus. 
The two first are quite distinct; Spherotrypes is also 
found in India. <Acanthotomicus is a separation from 
Tomicus, Latr. (1807). I have restored Taphrorychus 
apatoides, Wichh., to Dryocetes, and do not include any 
Japanese species in the former genus. 

So complete a collection testifies both to the ability of 
Mr. Lewis as a collector and to the richness of Japan in 
this family, for, though it is probably surpassed in this 
respect by many tropical countries, its 104 species compare 
very favourably with the 130 or so described from Europe 
and the rather larger number from North America. 
There must be many others to discover. Not a few 
species are unique, others have occurred here and there 
as single specimens, and 7 out of the 18 first described 
have not reappeared. ‘I'he total number existing in the 
islands may be expected to exceed 150, 

The best represented genera are [Hylesinus (6 species), 
Phleosinus (7 species), Scolytus (6 species), Dryocetes 
(S species), and Xyleborus (29 species exclusive of males). 


‘The number varies from that given in the ‘ Proc, Ent. Soc.,’ 
1393, p. xxxix, as I have received, since then, additional specimens 
from Mr. Lewis’s unmounted Coleoptera, and a small number from 
Colonel von Schénfeldt. 


Rhynchophorous Coleoptera of Japan. 55 


Oryphalus and Pityophthorus with one species each, and 
Tomicus with two, are poorly represented, and are likely 
to be augmented by future collectors. The Platypini 
consist of nine species in three genera. 

In origin the Japanese Scolytide are partly Palearctic 
and partly Oriental. From the former region come 
Hylastes attenuatus and glabratus, Myclophilus piniperda 
and minor, Crypturgus pusillus, Tomicus cembre, Dryo- 
cates autographus, and Urypedendrow quercus; Xyle- 
borus adumbratus, m., and sobrinus, Hichh., are little 
more than varieties of X. pfeili and saxesent respectively. 
Little is known of N. Asiatic Scolytide and further 
identifications cannot be made. The only Oriental species 
I have recognised are X. obliquecauda, Motsch., which 
occurs in Ceylon, and X. badius, Kichh.; but in many 
other Xylebori, and in the genera Spheerot trypes, Cos- 
moderes, Crossotarsus, and Diapus, we have character- 
istically Oriental forms. There are no American species 
among them, though in one or two genera, as NScolytus, 
[have not sufficient material from America to institute 
a comparison; and none of the few Hawaiian species 
described by Dr. Sharp are to be found. 

Though our knowledge of Asiatic Scolytide is so im- 
perfect as to render such conjectures hazardous, I am 
inclined to think that Oriental forms predominate. ’ There 
is no tendency of the Palearctic species to be confined 
to the northern islands of the Archipelago. 

In describing these small insects it is important to 
obtain all measurements exactly, whether of the total 
length or of the relative dimensions. Hstimation with the 
eye of the comparative width and length of a prothorax 
leads to error, as do rough measurements of length. 

Chapuis gives the length of Phlwosinus perlatus as 
2mm., whereas the type measures 3 mm., that 1s, 50 per 
cent. longer. Such an error in the measurement of a 
species an inch long would be gross, and it is hardly less 
so when made about a smallinsect. Exceptin a few genera, 
as Scolytus, the range of size does not exceed one fourth 
of the average length, and may be much smaller. 

I have taken all measurements with a micrometer eye- 
piece, and a mechanical microscope stage fitted with a 
scale reading to 1-10th mm. ‘This method is quite exact, 
and speedy in practice. 


56 Mr. Walter F. H. Blandford on the 


A large proportion of the type-specimens, both of 
genera and species in this family, are in Chapuis’ collection 
in the Brussels Museum, which I have visited for the 
purpose of examining them. I desire here to warmly 
acknowledge the hospitable reception I have met with 
from M. G. Severin, Curator of the Articulata, and the 
admirable facilities he has afforded me for tke study of 
that collection. 


SCOLYTINI. 


Hynastes, Hr. 


I have added three species to those described by 
Chapuis and changed one of his names. 


TABLE OF SPECIES: 
1. Third tarsal joint not wider than the preceding joints ; meso- 
sternum not prominent in front 
Elalastes 2s) stirs) © igh 4 loanngen ee 
wider than preceding joints ; mesosternum 
prominent (Hylurgops, Lec.) . 5 


2, Mostrumcarimate.. 7) 4: 2eys.e ss 5 | enallelus. lien 24 
NO CANINANS wk) vem tie wees lewkcai at teers te een 

3. Thorax not transverse, uniformly narrowed toapex .. . 4 
transverse, constricted atapex. . . . . . ambiquus. 


4. Interstices with a single row of bristles from base to apex. 
attenuatus, Er. 
with an irregular double row from base to middle, 
thence single . . plumbeus. 
. Punctures of thorax of two sizes. . . . . énterstitialis, Ch. 
uniform . . 1! < . . glabratus. Zett. 


qn 


Hylastes parallelus, Chap. 
Chap., Scol. Jap., p. 196. 


Common; Yokohama, Bukenji, Kiushiu (Higo, &c.) 


Hylastes atienuatus, Hr, 
One example, Hiogo (von Schdnfeldt). 


Rhynchophorous Coleoptera of Japan. 57 


Hylastes plumbeus, n. n. 
H. obscurus, Chap., Scol. Jap., p. 197. 


Common; taken with H. parallelus, also at Nikko, 
Kobe, &e. 


Somewhat variable, in colour from black to obscure 
brown, in the width of the rostrum and prothorax, and 
in the depth of the elytral striz. I have changed the 
name, owing to the priority of Hylastes (Hylastinus) 


obscurus, Marsh, (trifolii, Mil.) 


Hylasies ambiguus, sp. 0. 

Oblongus, subnitidus, piceo-brunneus, capite granulato, fronte 
haud carinata, punctata ; prothorace transverso, antice constricto, 
fortiter punctato, linea media elevata levi; elytris basi singulis 
subtiliter rotundatis, prothorace latioribus et duplo longioribus, 
striato-punctatis, striis aequaliter impressis, interstitiis rugosis 
versus apicem tuberculatis, squamulis brevibus vestitis et uniseriatim 
setosis ; tarsorum articulo 30 antecedentibus haud latiore. Long. 
2°7 mm. 


Fujisan, one specimen, in bad condition, apparently 
dead when taken. 


Suggesting in appearance the species of the sub-genus 
Hylurgops, Lec., but without the prominent mesosternum and 
evidently bilobed 3rd tarsal joint. Reddish-brown, somewhat shin- 
ing. Head black, finely granulate on vertex with a shining central 
line, rostrum wide, not carinate, front strongly punctured ; 
antennal club with first joint large, equal to the two succeeding 
joints. Prothorax transverse, constricted towards apex above and 
at sides, strongly and closely punctured, inconspicuously pubescent, 
with a fine central raised line from base to anterior constriction. 
Elytra wider than prothorax at base, and exactly twice as long, 
basal angles prominent rounded, sides slightly rounded to near 
apex, thence somewhat oblique and feebly sinuate, apex rather 
obtuse in middle ; above brown, lighter behind, striato-punctate, 
striae not deep, their punctures strong, interstices rugose at base, 
with a single row of fine tubercles from middle to apex, covered 
with short scale-like hairs and a single series of scattered sete. 
Underside brown, punctured ; middle coxw rather widely separated. 


Very like H. opacus, Er., but quite distinct in the 
shape of the thorax. 


58 Mr. Walter F. H. Blandford on tie 


Hylastes interstitialis, Chap. 
Chap., Scol. Jap., p. 196. 


A few more examples taken at Subashiri, Kiga, and 
Nagasaki, 


ITylastes glabratus, Zett. 

Nikko ; two specimens. 

This European species is very similar to I. interstitialis, 
but the finer uniform punctuation of the prothorax is 
quite different from the much coarser and variolose . 
punctuation of interstitialis, in which the coarse punctures 
are mixed with finer ones. J. pinifew, Fitch, which I 
consider distinct from HH. glabratus, has the thoracic 
punctures of two sizes, but not variolose or confluent. 


Myetorniuus, Hichh. 


Tylurgus, Latr. .« . « Blastophagus, Wichh. 


Both the Huropean species are found in Japan. 


Myelophilus piniperda, Fabr. 
Already recorded by Chapuis (Scol. Jap., p. 197). 
Taken at Nagasaki and Oyayama, both in Kuushiu. 
Some specimens measure as much as 5‘ mm. 


Myeloplilus minor, Eart. 


This species has been hitherto inserted in the Japanese 
list by error; Chapuis mentioned it (Scol. Jap., p. 198) 
as a species likely to occur, and stated that it was 
brought from China by Mr. Lewis, who has since then 
taken one specimen near Nagasaki. 


HyorRHYNCHUS, nov. gen. 

Caput rostratum, rostello lateraliter marginato, Oculi bipartiti. 
Antenne sub carina rostrali inter partes oculorum inferiores 
et mandibulas inserte, scapo recto, funiculo 7-articulato, 
articulo lo magno, 20 obconico, ceteris transversis latitudine 
crescentibus, clava magna oblonga subcompressa, 3-articulata, 


Tthynchophorous Coleoptera of Japan. 5G 


suturis rectis notata, pilosa. Prothorax subdepressus, lateribus 
determinatis, non tamen marginatis, basi immarginata. Wlytra ad 
basim singulatim convexa, thoracem superantia, Cox antice 
magn globose, a processu prosterni angusto separate. Pedes 
longi, tibiis subcompressis, leviter dilatatis, ad apicem oblique 
truncatis, margine exteriore inermi, ‘T'arsi articulo lo brevi, 2o 
paullo longiore incrassato, 30 brevi, subtus producto et profunde 
emarginato, 40 minimo, So magno ceteris conjunctim acquali, 


flead shortly rostrate, the sides of the rostrum carinate, differing 
according to sex in the single species. Eyes divided, their seg- 
ments distant, placed above and below the base of the rostrum. 
Antennge inserted below the rostral carina between the lower half 
of the eye and the base of the mandible, short, the club nearly 
equal in length to scape and funiculus together. Maxilla with 
outer border rounded, inner edge convex and narrowly produced at 
tip, set with straight flat spines. Submentum produced into an 
angle at either side and broadly emarginate between, witha second 
emargination in middle at base of mentum, which is nearly twice 
as long as broad, narrow at base and becoming dilated to middle 
with sides, thence parallel ; labial palpi long, with joint 1 as long as 
broad, 2 transverse, 3 longer than broad. Prothorax flattened 
above, its sides strongly inflexed to anterior cox, forming an 
angle with pronotum which is not sharp or margined. Anterior 
cox not situated near front border of prosternum, very large, 
prominent and globose, separated by a narrow process ; meso- 
sternum not strongly depressed, middle and posterior cox 
prominent, remote ; metasternum rather short with wide parallel 
episterna, First two abdominal segments little longer than two 
following, which are together equal to fifth, Legs long, tibime quite 
simple, flattened, slightly curved, obliquely excised at apex, with 
inner angle shortly spinose; fourth tarsal joint inserted near base 
of third, which is produced underneath into a long split lobe, but 
not laterally widened. 


The one species of this genus has, at least in the male, 
more the appearance of a Curculionid or an Anthribid, 
than a Scolytid, owing to the prominent rostrum, the 
general shape of its body, the vestiture, and the unarmed 
tibie. It is, however, a true Scolytid, and either one 
of the Hylesini, or not remote from them. I know, 
however, of no described genus with which it is 
closely allied. 


60 Mr. Walter F. H. Blandford on the 


Hyorrhynchus lewisi, sp. n. 

Oblongo-ovatus, opacus, niger, pube sericea fusco-cinerea dense 
vestitus, antennis tarsisque ferrugineis ; prothorace transverso, 
basi bisinuata, lateribus postice parcius, antice fortius rotundatis, 
supra post apicem longitudinaliter impresso, dense granulato ; 
elytris post medium dilatatis, supra tenuiter striatis, fuliginosis, 
vitta transversa angulata et apice cinereis, interstitiis leniter 
convexis ad basim granulatis. Long. 3°5—5°3 mm. 

Mas. Capite fortiter longitudinaliter;sulcato, rostro longiore 
ad apicem in angulos prominentes lateraliter producto.’ 

Fem. Capite subconvexo, rostro breviore; prothoracis lateribus 
antice muricatis. 


Several examples taken at Sapporo, in 1883, by a 
native collector. 


Head in the male produced obliquely forward into a flattened 
rostrum, the carinate sides of which form a sharp prominent and 
backwardly directed angle; upper divisions of the eyes rounded 
triangular, situated on either side of front, which is deeply furrowed 
between them, and finely carinate on the inner margin of each 
eye-segment ; lower divisions of the eyes hidden from above by 
the lateral carina. Head in the female with a very short rostrum, 
narrowly carinate and not produced at sides, impressed over mouth 
and with a short median carina; in both sexes black, rugosely 
punctured with short close-lying hairs. Antenne ferruginous. 
Prothorax narrowed in front, widest at base, with sides rounded; 
surface black, pilese, with short cinereous hairs, closely granulate 
and muricate at sides in the female, with a weak impression on 
either side towards the base. Scutellum rounded, pubescent. 
Elytra wider than prothorax, and more than twice as long, dilated 
behind middle, separately rounded at base, and overlapping thorax, 
inflexed below humeral angles to afford room for middle femora, 
sides nearly straight to behind middle, thence strongly rounded ; 
surface covered with fine close-lying hairs, smoky-black except on 
humeral angles, apex, and an oblique angulated vitta, where they 
are cinereous, With fine incised strize meeting at apex, as in 
Hylesinus, interstices subconvex, granulate at base. Underside black 
with fine close punctuation, pubescent. Legs piceous with tarsi 
lighter. 


The sexual differences in the rostrum are not found 
in any other Scolytid known to me, and suggest those 
of the Brenthidx, with which this insect has no affinity. 


Rhynechophorous Coleoptera of Japan. 61 


SPHHROTRYPES, NOv. gen. 


Caput oblongum, in rostellum haud productum. Oculi bipartiii. 
Antenne -lateraliter inter mandibulas et partem inferiorem 
oculorum inserts, breves, funiculo 7-articulato, articulis latitu- 
dine crescentibus, clava ovata, rotundata, compressa, 3-articulata, 
vittis setarum transversis annulata, suturis transversis. Pro- 
thorax lateraliter marginatus. Mesosternum brevissimum, inflexum; 
metasternum breve. Coxe antice et mediz late distantes. Tibi 
antic ad apicem spinula unica extus producta armatze, posteriores 
spinose. Tarsorum articuli 1-3 equales, 30 bilobo. 


Head flattened, oblong but not rostrate; eyes bipartite, the 
divisions flat, subtriangular, granulate and widely separated, united 
posteriorly by a fine carina. Antennz inserted at upper angle of 
lower division of eye, short, their scape curved, slightly clubbed ; 
funiculus with 1st joint large, globose, 2nd obconical, 3—7 trans- 
verse, increasing in width ; club rounded oval, flat, with transverse 
sutures, Ist two joints transverse, together equal in length to 3rd, 
with transverse bands of short hairs, three on 1st joint, two on each 
succeeding joint, which give the club the appearance of being 
multi-articulate ; apex of 3rd joint spongy, covered with short thick 
serrate hairs. Mandibles stout, prominent, not toothed. Maxille 
very hairy externally, inner margin convex, set with short flat 
spines ; maxillary palpi short with joints equal in length, hairy. 
Submentum inconspicuous, produced into a short pointed process 
on either side of mentum, which is oblong, slightly narrowed at 
base and in middle of sides, with apex truncate ; ligula small, ovate 
acuminate, inserted near apex of mentum; labial palpi short, 
joints 1 and 3 as long as broad, 2 transverse, densely hairy. 
Prothorax bordered at sides, excised in front to anterior cox, 
which are widely separated by a transverse prosternal process. 
Mesosternum very short, inflexed so that anterior and middle coxa 
meet when the thorax is depressed. Metasternum barely larger 
than 1st abdominal segment, its episterna moderately wide, dilated 
anteriorly. Posterior coxee separated by a broad rounded process 
of Ist abdominal segment, which is slightly longer than the 2nd, 
3rd and 4th successively shorter, 5th about equal to Ist. Anterior 
tibie slightly dilated towards apex, outer margin almost simple 
except for a strong outwardly directed spine at apex and a smaller 
spine at inner angle; hinder tibiee slightly curved, spined exter- 
nally. Tarsi with 3rd joint bilobed, its processes long and narrow. 


In spite of the divided eyes, I think this very distinct 


62 Mr. Walter I’. H. Blandford on the 


genus may be placed at present in the Hylesini, near 
Dendrosinus, which resembles it in shape, and in the 
structure of the antennal club. ‘The peculiarities of the 
underside are due to its globose form, and consequent 
compression. ‘The elytra are separately curved at base, 


but are not elevated above the base of the prothorax. 
Spheerotrypes pila, sp. n. 

Brevissime ovatus, fere globosus, niger, elytris piceis, antennis 
tarsisque ferrugineis; capite supra os transverse impresso, 
reticulato punctato, fronte pilosa; prothorace valde transverso, 
anterius fortiter angustato, basi postice obtuse producta, sub- 
tilissime marginata, supra post apicem transverse impresso, dense 
punctato, subnitido, parcius squamoso, linea media elevata laevi : 
elytris striato-punctatis, punctis obsoletis, interstitiis planis, 
squamatis, squamis pro maxima parte fuscis, interdum praecipue 
versus apicem cinereis. Long. 2-4—3 mm. 


Hitoyoshi, several examples; it has occurred in the 
thin bark of a camellia. 

Very short oval, exceedingly convex. Head with front 
flat in female, impressed in male, punctured, and thinly hairy, 
the hairs ascending on to middle of vertex, which is smooth 
at the sides, finely reticulate and scantily punctured. Prothorax 
nearly double as wide as long, its base bordered and produced 
backwards to form an obtuse angle, and slightly concave on either 
side, basal angles acute, sides rounded and strongly narrowed from 
base to apex; dorsum separated throughout from flanks by a fine 
ridge, convex, transversely impressed behind apex, with close rugose 
punctuation and a narrow elevated line from base to middle, some- 
what shining with a scanty covering of scales, chiefly at apex and 
on sides, its anterior border fringed with short hairs. Scutellum 
oblong, rugose. Elytra rather wider than prothorax and _ less 
than twice as long, conjointly emarginate at base, basal borders 
slightly rounded, crenate, not overlapping base of thorax, basal 
angles very broadly rounded, sides rounded from base to apex; 
above dull brown covered with fuscous scales and with a dusty 
appearance, due to scattered cinereous scales; striate, the strix 
rather deep, with obsolete and scattered punctures, interstices quite 
flat, rugose, more strongly at base. Underside black, punctured. 
thinly covered with scales. Legs blackish with tarsi lighter. 


[The following species, though not from Japan, is so 
closely allied that it may well be described here: 


Tthynchophorous Coleoptera of Japan. 63 


Spherotrypes globulus, sp. n. 

Brevissime ovatus, niger, elytris piceis, antennis et tarsis 
rufescentibus ; capite reticulato, punctato, fronte squamulosa, sub- 
tuberculata, obsolete carinata; prothorace transverso, anterius 
fortiter angustato, basi postice acutius producta, marginata, post 
apicem constricto, dense et rugose punctato, linea media elevata 
obsolescente, opaco, densius squamoso; elytris striato-punctatis, 
interstitiis planis, rugosis, squamis fuscis passim obscure dilu- 
tioribus obtectis. Long. 2-4—5 mm. 

Inpia, Belgaum ; taken by Mr. H. F. Andrewes. 

Differs from the preceding as follows :— 


Front of head tuberculate, more strongly covered with short 
scales with a few longer ones at sides, and usually distinctly 
carinate over the mouth, sometimes throughout less impressed in 
the male. The carina is variable, and a trace of it exists in 
S. pila. Prothorax more strongly angulate behind, its central line 
nearly obsolete, the surface quite dull with closer punctures and 
more scales. Scales of the elytra brown, occasionally a little lighter, 
but not evidently pale and cinereous, as in 5. pila, except afew at 
the apex of the suture. | 


Hytesinus, Fabr. 


The six Japanese species are all new. They may be 
thus distinguished :— 


1. Prothorax not closely covered with scales . . . . eae 


and elytra closely covered with chenuered scales 
scutulatus. 


2, Alternate interstices of elytra elevated towards apex . . . 9 
Interstices similar towards apex . ...-. +--+, 4 

3. Elytra glabrous, sizeunder4mm.... .- + - ~- costatus. 
hairy atapex,size6mm. .,.... +--+ « nobilis. 

A) Kilytra umicolorous black 2. . 2 « «°s * 4 2 © 4 « 5 


with a transverse vitta of light scales . . . céugulatus. 

5. Form broadly ovate, elytra less convex longitudinally than 
HbdoMen, ol ate as) a) se aa Cate an weklecOnate 

oblong ovate, elytra not less convex than abdomen tvstis. 


Hylesinus costatus, sp. 0. 
Oblongus, subnitidus, fere glaber, niger, elytris picinis, antennis 
tarsisque ferrugineis; prothorace transverso, basi vix producta, 
lateribus rotundatis, granulato-punctato, linea media obsolete 


64. Mr. Walter F. H. Blandford on the 


elevata, lateraliter versus apicem muricato; elytris prothorace 
plus quam duplo longioribus, subcylindricis, striato-punctatis, 
interstitiis usque ad declivitatem transverse rugosis, lo, 30, 50, 7o 
in declivitate elevatis, seriatim tuberculatis, ceteris punctatis. 
Long. 3°4 mm. 


Junsal; one specimen. 


Oblong, black, with elytra obscurely piceous. Head with 
labrum pitchy, separated by a transverse depression from front, 
which is flattened, shining, strongly punctured and glabrous ; 
vertex finely reticulate. Antenne ferruginous, club small, pointed, 
sutures transverse. Prothorax transverse, base biconcave, 
depressed, scarcely produced in middle, sides strongly rounded ; 
surface convex, finely reticulate and with strong asperate punctua- 
tion somewhat weaker at base, with traces of a central elevation ; 
sides tuberculate before apex. Scutellum small, punctured. 
Elytra wider than prothorax and two and a half times as long, 
their bases convex, overlapping thorax, sides parallel to middle, 
then rounded to apex, surface convex cylindrical, strongly 
declivous behind, with strong punctured striw, the punctures 
round and distinct ; all interstices transversely rugose and punc- 
tured to middle, 1, 3, 5, 7 and 9 after middle with a series of 
transverse asperities which become strongly tuberculate on the 
apical declivity, where the interstices are elevated ; alternate 
interstices not asperate behind middle, multipunctate. Underside 
black, punctured, shortly pubescent, metasternum with a deep 
longitudinal impression, its episterna not very narrow ; abdomen 
scarcely convex longitudinally, its terminal segment rugose. 
Tibia spined externally. 

In the flatness of the abdomen this species differs 
from the majority of the genus, except the sub-genus 
Pteleobius, Bedel, to which it is not related. 


Hylesinus nobilis, sp. n. 
Oblongo-ovalis, obscurus, niger, antennis tarsisque piceis; pro- © 
thorace transverso, basi producta, lateribus versus apicem sub- 
angustatis, .aequaliter granulato-exasperato, brevissime  piloso ; 
elytris prothorace plusquam duplo longioribus, ad basim angustatis, 
lateribus anterius subtiliter, posterius fortiter rotundatis, apice 
emarginato, profunde striato-punctatis, interstitiis rugulose tuber- 
culatis versus apicem breviter pilosis, 10, 30, 50 subelevatis. Long. 
6°2 mm. 


Sapporo ; one specimen. 


Rhynchophorous Coleoptera of Japan. 65 


Oblong-oval, black, dull. Head finely aciculate with scattered 
punctures, front flattened, shortly pubescent, antenne pitchy- 
brown with club blackish, longer than funiculus, bluntly pointed, 
its sutures oblique. Prothorax transverse, base strongly produced 
behind and biconcave, sides strongly rounded behind, contracted 
in front and sinuate, apex nearly straight in middle ; surface 
slightly impressed in middle behind apex, not distinctly impressed 
before base, uniformly and densely granulate, with short pube- 
scence. Elytra wider than prothorax in middle and more than 
twice as long, humeral angles very obtuse, sides rounded at base, 
becoming wider, thence nearly straight, gradually and strongly 
rounded towards apices which are separately rounded ; surface 
convex, more strongly towards apex, with deep indistinctly punc- 
tured striz ; interstices in front strongly tuberculate, with a few 
scattered hairs, posteriorly with rugosities weaker, hidden by short 
fuscous hairlike scales, and with a single row of erect sete ; Ist, 
3rd and 5th elevated at apical declivity, the two latter conjoined. 
Underside punctured and finely pubescent, abdominal segments not 
very convex. Legs black, with tarsi lighter, anterior tibiz dis- 
tinctly spined on outer side of apex. 


The largest species in the genus. 


Hylesinus laticollis, sp. n. 


Ovalis, convexus, obscurus, breviter pilosus, niger, antennis 
ferrugineis, prothorace fere duplo latiore quam longiore, basi valde 
producta, lateraliter fortiter rotundato, supra rugose exasperato, 
lateribus antice muricatis, linea media obsoleta laevi, ante basim 
utrinque oblique impresso ; elytris prothorace plus quam duplo 
longioribus, versus apicem oblique declivibus, striatis, striis ad 
medium uniseriatim punctatis, postice laevibus ; interstitiis 
similibus, antice tuberculatis, postice squamulis brevibus fuscis 
vestitis, margine laterali breviter setoso ; abdomine valde convexo 
ad apicem setoso. Long. 4—4:2 mm; lat. 2°2—2°3 mm. 


Three examples, taken by a native collector at Sapporo 
in 1883. 


Broad oval, convex, black. Head strongly punctured, front 
flattened, shortly hairy ; in two specimens, probably males, im- 
pressed over mouth with a fine central carina, and with stronger 
pubescence ; in the other, probably a female, with impression and 
carina obsolete and hairs scantier; epistoma shortly produced over 
mandibles, vertex reticulate. Antenne ferruginous, club long, 


TRANS. ENT. SOC. LOND. 1894,—PaRT I. (MARCH.) 


65 Mr. Walter F. H. Blandford on the 


stout, with transverse sutures. Prothorax very transverse, 
narrowed from base to apex, with sides rounded, base strongly 
produced in middle, surface asperately punctured, with short 
bristles, closer at sides, with au indistinct smooth central line, 
and an oblique impression on either side before and parallel to 
base ; sides muricate towards apex. Scutellum rounded, rugose. 
Elytra wider than prothorax and two and a half times longer, 
widest in middle, base of each strongly rounded and crenate ; sides 
slightly rounded to middle, gradually more strongly towards apex, 
which is obtuse; surface obliquely and not strongly declivous 
to apex, with deep rather wide strix, obsoletely punctured to 
middle, thence smooth, interstices coarsely tuberculate at base, the 
tubercles becoming finer, and being replaced behind the middle 
by short fuscous scales, which give the elytra a brownish tinge ; 
the two outer interstices and lateral margin with a close covering 
of short bristles. Underneath coarsely punctured, shortly hairy ; 
abdomen more convex longitudinally than elytra, first two seg- 
ments much longer than last three, fourth and fifth bristly. Spines 
of anterior tibiz nearly obsolete. 


Hylesinus tristis, sp. n. 

Oblongo-ovalis, obscurus, niger, antennis tarsisque ferrugineis ; 
prothorace transverso, basi brevius producta, lateribus rotundatis, 
versus apicem muricatis, supra asperate punctato, utrinque ante 
basim impresso; elytris prothorace sesqui amplius longioribus, 
lateribus ad medium subrectis, postice rotundatis, apice subtiliter 
emarginato, striatis, striis ad basim modo obsolete punctatis, inter- 
stitiis transverse asperatis, postice squamulis cinereis inconspicuis 
vestitis, margine lateraliin versura solum distincte setoso. Long. 
3—3°5 mm. ; lat. 1:5—1°9 mm. 

Mas. Fronte impressa, interstitiis post medium seriatim 
tuberculatis. 

Several specimens taken at Ichiuchi, May Ist, 188], 
and a few at Junsai. 

Similar in sculpture and appearance to H. luticollis, but smaller, 
more oblong-oval, less strongly convex, the elytra more declivous 
behind, as convex as ventral surface. Front in male broadly im- 
pressed between eyes with a short central carina, pubescent ; in 
female, narrowly impressed over mouth, flat between eyes, with 
pubescence thin. Prothorax less transverse than in H. laticollis, 
with sides less narrowed to apex, and base less strongly produced 
in middle, its sculpture similar. Elytra narrower, subparallel to 
middle, less obtuse at apex, interstices less coarsely tuberculate at 
base, alternate interstices behind middle with an indistinct row 


Rhynchophorous Coleoptera of Japan. 67 


of stronger tubercles in the male ; lateral set shorter throughout, 
inconspicuous before middle of elytra. Anterior tibie distinctly 
spinose on outer margin. 


Hylesinus cingulatus, sp. n. 

Oblongo-ovalis, obscurus, niger, antennis tarsisque rufescen- 
tibus, sat dense et breviter pilosus ; prothorace transverso, basi 
bisinuata, haud producta, lateribus aequaliter rotundatis, antice 
tuberculatis, supra dense granulato-exasperato ; elytris ad medium 
lateribus subparallelis, at subsinuatis, postice rotundatis, supra 
striato-punctatis, squamosis, vitta fusco-cinerea transversa latera- 
liter versus apicem curvata notatis, interstitiis ad basin tuberculatis, 
postice subrugosis. Long. 2°3—2:°8 mm. 


Lake Junsai; five specimens. 


About the size and shape of small examples of H. frazini. 
Black, dull. Head closely granulate, front hairy, impressed in 
male, subconvex in female. Antenne ferruginous, club rather 
broad, obtusely pointed, with dark pubescence. Prothorax with 
base bisinuate, not produced as in the two last species, sides rounded 
at base, thence narrower and straighter to apex ; above uniformly 
convex, thinly hairy, without scales, granulate, the asperities 
stronger towards the sides, which are distinctly tuberculate in 
front. Elytra as wide as prothorax, and two and a half times 
longer, separately rounded at base, overlapping thorax, sides 
subparallel, feebly sinuate to behind middle, thence rounded ; 
surface convex, obliquely declivous behind, striate, the strive punc- 
tured to behind middle, interstices tuberculate at base, then rugose, 
covered with short hairs and scales, blackish except on middle of 
elytra, where they form a transverse cinereous-brown vitta, curved 
backwards at sides so as to cover the apices of the 5th to the 
marginal interstices, and continued narrowly along apical margin, 
forming an irregular oval, Underside punctured, with rather 
dense cinereous pubescence ; abdomen not strongly convex. Legs 
ferruginous or pitchy, with tarsi lighter, front tibie hairy, spined 
externally. 


Hylesinus scutulatus, sp. n. 

Oblongus, niger, squamulis cinereis fuscisque  tesselatus, 
antennis pedibusque nigrescentibus ; prothorace vix latiore quam 
longiore, basi bisinuata, lateribus rotundatis versus apicem tuber- 
culatis; elytris prothorace latioribus et illo duplo longioribus, 
cylindricis, striato-punctatis, interstitiis ad basim subtuberculatis, 
per totum uniseriatim sctosis. Jong, 2°7 mm, 


68 Mr. Walter F. H. Blandford on the 


One or two examples taken at Kiga, Subashin, 
Nagasaki, Omori, and Oyama. 


Oblong, cylindrical, black, tesselated with close-lying grey and 
brownish scales. Head granulate with front flattened, hairy, im- 
pressed and more densely hairy in the male. Antenne black, club 
rather short, acuminate oval, its basal joint large. Prothorax 
rather broader than long, its base bisinuate, not produced, sides 
uniformly rounded, tuberculate in front, surface regularly convex, 
front and sides with short erect bristles. Elytra cylindrical, wider 
at base than thorax, and a little more than twice as long, base 
crenate, sides subparallel to behind middle, apex strongly declivous 
and convex; with rather fine punctured strie, interstices flat 
with a single row of sete throughout, and with one or two 
tubercles at base. Underside strongly punctured with short scale- 
like hairs, metathoracic episterna narrow, abdomen not convex 
longitudinally. Legs black, with tarsi lighter. 


Two specimens are larger and broader than the rest, 
the scales are pale ashy-grey and yellowish, whereas in 
the others they are light and dark brown. I believe the 
two former to be females and not specifically distinct. 
This species may be referred to Bedel’s sub-genus 
Pteleobius, and is nearest to H. vittatus, Fabr., of the 
European fauna, but is larger, more elongate, with the 
strize deeper and less clearly punctured, and the inter- 
stices more convex and distinctly setose. 


Puia@osinus, Chap. 


In the generic diagnosis given by Chapuis (Syn. Scol., 
p. 93), the third tarsal joint is stated to be simple. This 
is correct for the European P. aubei, Perr., and P. 
thuye, Perr., but in the Japanese species it is bilobed, 
though M. Chapuis has made no mention of this point 
in the descriptions of P. lewisi and perlatus. It is also 
bilobed in the American species, according to Leconte, 
except in his Chetophleus hystrix, which can hardly be 
separated from the genus upon the characters given, and 
in which the 3rd tarsal jot “is emarginate rather than 
bilobed.’? This difference in structure is not here of 
generic value, and the genus is sufficiently recognisable 
by the structure of the antenne. 


Rhynchophorous Coleoptera of Japan. 69 


There are seven Japanese species, of which five 
are new. 


TABLE OF SPECIES. 
1. Head with a median carina immediately over mouth . . . 2 
notcarinate overmouth . . . : 5) 
2. Elytra closely covered with scales, muteraticgs altaenntely lighter 
anddarker . . 3 


thinly covered with hairs or scales, unicolorous . . 4 
3. Prothorax abruptly contracted in front, reddish-brown 
pulchellus. 


gradually contracted in front, black . . . dubius. 

4, Elytra black, interstices similar, sizesmall . . . . minutus. 
red-brown, 2nd interstice depressed and narrowed at apex 

perlatus, Ch. 

5. Interstices without seriate bristles, Ist and 35rd tuberculate 
towards apex. . rudis. 

uniformly granulate, with seriate bristles. . . 6 

Bristles on 2nd interstice ceasing before apex, front subcarinate 
between eyes. . serzatus. 

on interstices, similar throughout. . . . lewis, Ch. 


i=z) 


Phieosinus pulchellus, sp. u. 


Ovalis, rufo-testaceus, antennis pedibusque ferrugineis ; capite 
nigro, punctulato, breviter piloso, vix rostrato, supra os fortiter 
carinato ; prothorace transverso, antice abrupte contracto, supra 
convexo, linea media subelevata, rugose punctato et squamis 
adpressis haud dense vestito ; elytris post medium subdilatatis, 
fortiter striato-punctatis, interstitiis planis, 20, 40, 60 muticis, 
squamis brunneis, ceteris versus apicem subtuberculatis, squamis 
flavo-testaceis vestitis. Long. 2°5 mm. 


One specimen, Wada Toge. 


Oval, convex, red-brown, elytra closely squamose with alternate 
lighter and darker stripes. Head impressed over mouth (probably 
a male character) with a strong frontal carina, punctured, 
and shortly hairy. Prothorax nearly one-half broader than long, 
strongly narrowed in front, sides behind nearly straight, then very 
convex at contraction, becoming straighter in front ; base narrowly 
impressed, surface convex, slightly depressed at apex, and obscurely 
elevated in middle from base to apical depression, densely punc- 
tured and sprinkled with close-lying yellow scales. Elytra wider than 
prothorax and twice as long; basal margin crenate, everted at sides, 
humeral prominences obliterated sides subsinuate to beyond 


70 Mr. Walter F. H. Blandford on the 


middle, thence rounded ; surface slightly dilated at posterior third, 
then strongly convex to apex, closely squamose, the scales cinnamon- 
brown on the 2nd, 4thand Gth interstices, yellow on the rest of the 
elytra ; striate, the stric rather wide, with distinct punctures, inter- 
stices flat, eranalate at base, 2nd widened at base and on summit 
of declivity, 1st, 3rd and outer interstices inconspicuously 
tuberculate towards apex. Underside ferruginous, scantily pube- 
scent, strongly punctuate on metathorax and two first abdominal 
segments, Legs ferruginous, anterior tibize with strong out- 
wardly directed spines at outer apical angle. 


Phleosinus dubius, sp. n. 

Ovalis, niger, subopacus, antennis tarsisque ferrugineis ; capite 
punctato-granulato, brevissime piloso, haud rostrato, fronte supra 
os deplanata, in medio carinata; prothorace transverso, antice 
gradatim contracto, supra convexo, linea media passim ele- 
vata, rugose punctato, parce squamoso; elytris subcylindricis, 
lateribus post medium subrectis, striatis, striis obsolete punctatis, 
interstitiis squamosis, squamis fuscis, in interstitiis lo, 30, 50 non- 
nihil dilutioribus, his versus apicem tuberculatis. Long. 2°2 mm. 


One specimen, Kurigahara. 


Very similar to P. pulchellus, with the appearance of being a 
colour variety, but differing in certain other points. Entirely 
black with only the antennee and tarsi reddish ; head less strongly 
impressed over mouth, and frontal carina less acute. Prothorax 
narrower, flatter throughout with the sides more gradually sloped 
towards the apex, and not strongly rounded in the middle, so 
that the apical constriction is less marked; punctuation less 
strong and interstices covered with fine reticulation, not evident 
in P. pulchellus. Wlytra more oblique at base, with borders 
straighter, not evidently dilated behind the middle, the sides 
consequently not subsinuate, but straight to near the apex ; 
humeral elevations small, distinct, black, covered behind the 
middle (the base is probably rubbed in the single specimen), 
with fuscous scales, which are obscurely lighter on the alternate 
interstices, as in 2. pulchellus. Stris finer, not evidently punctured, 
the 2nd interstice narrower and straighter throughout. Apical spines 
of the anterior tibiz longer, curved backwards, and connate for 
the greater part of their length, 

The two preceding species resemble small species of 
Hylesinus rather than see osinus, owing to the thickness 
of their scaly covering; but they present no characters 
to separate them from the latter genus. 


Rhynchophorous Coleoptera of Japan. 71 


Phiwosinus minutus, sp. n. 


Ovalis, niger, subnitidus, antennis pedibusque infuscatis ; capite 
granulato, fronte subconcava, pilis flavis circumdata, supra os 
carinata; prothorace transverso, antice contracto, lateribus in 
medio fortiter rotundatis, supra subtilissime reticulato et disperse 
punctato, punctis asperatis, elytris cylindrico-convexis, striato- 
punctatis, interstitiis planis subrugosis, uniseriatim  subtiliter 
tuberculatis et squamulis erectis vestitis, lo, 30, 50 in margine 
apicali spinula unica armatis. Long. 1-5 mm. 


One example, Ichiuchi. 


Black ; head not rostrate, closely granulate, front impressed, 
concave, strongly punctured round margin, with a few longish hairs, 
carinate above mouth. Prothorax transverse, strongly contracted 
towards apex, base bisinuate, its angles broadly rounded, sides 
slightly rounded, more strongly at contraction, surface convex with- 
out median elevation; scantily pubescent, closely reticulate and asper- 
ately punctured, the punctures strongest towards apex, sparser over 
sides behind middle. Scutellum piceous, shining, punctiform. Elytra 
scarcely wider than prothorax and less than twice as long ; basal 
margins conjointly nearly straight, crenate at sides only ; sides 
straight to middle, thence uniformly rounded; above shortly 
cylindrical and very strongly rounded at apex, pitchy-black with 
punctured strie, the punctures not close, interstices flat, shining, 
subrugose, with irregular rows of erect scales and minute tuber- 
cles, Ist, 3rd, 5th and 7th with one or two stronger tubercles 
towards apex and a sharp pointed spine just above lower border of 
elytra. Legs piceous with tarsi lighter. Third joint scarcely 
bilobed. 


The smallest species as yet in the genus, with a 
deceptive resemblance to Phleophthorus rhododactylus, 
Marsh. (spartii, auct.), but separable by the point of 
attachment of the antennz, the broader and anteriorly 
contracted thorax, which is finely reticulate as_ well 
as punctured, and the apical spines of the elytral 
interstices. 


Phleosinus perlatus, Chap. 
Chap., Scol. Jap., p. 198. 


Originally from Hiogo ; one or two fresh specimens are 
without indication of locality. The only sexual character 


72 Mr. Walter F. H. Blandford on the 


is that the front is impressed and concave in the males, 
flat in the females. The length, inaccurately given 
by Chapuis, is 2°5—3 mm. 


Phleosinus seriatus, sp. u. 


Oblongo-ovalis, subopacus, breviter pilosus, niger, elytris rufo- 
brunneis vel nigris, antennis tarsisque ferrugineis ; capite sub- 
rostrato, fortiter punctato, fere glabro, supra os impresso ; pro- 
thorace antice constricto subtransverso, supra convexo dense ac 
fortiter punctato, linea media abbreviata levi nitida; elytris 
pube brevissima vestitis, striato-punctatis, punctis rotundis sat 
frequentibus, interstitiis passim convexis, rugosis, serie unica 
setarum ornatis, lo, 30 versus apicem tuberculatis, setosis, 20, 40 
muticis, in declivitate haud setosis, margine laterali per totum 
spinulis minutis armato. Long. 2.4 mm. 

Mas. Fronte media impressa, et linea elevata laevi inter 
oculos ornata. 

Fem. Fronte deplanata, linea media laevi haud elevata. 

A pair, Higo. 

Oblong-oval, black, or with elytra reddish-brown; head 
subrostrate, strongly punctured, with an arcuate impression 
above mouth anda second smooth impression above that in male, 
at which ends the short frontal median shining line, which is 
elevate though scarcely carinate in the male, flat in the female ; 
antenne ferruginous. Prothorax transverse, its basal angles 
rounded, sides strongly contracted towards apex, in the male almost 
angulated, above convex without central elevation, but with a 
median smooth line, which does not reach either border, covered 
with short close hairs, densely and strongly punctured and not 
asperate. Elytra with basal borders rounded, scarcely everted at 
sides, sides straight in front, rounded posteriorly and slightly 
sinuate at apex of 4th interstice when seen from above ; surface 
subdilated behind middle, thence convexly declivous, covered with 
short hairs, strie weak with large round close punctures ; inter- 
stices with traces of tuberculation throughout. evident at base and 
on alternate interstices towards apex, where the 2nd interstice is 
depressed as in P. perlatus, more strongly in the male than in the 
female, and a single series of bristles, absent on alternate inter- 
stices, towards the apex ; marginal interstice with a row of short 
spines throughout, stronger towards apex. Legs pitchy with knees 
and tarsi reddish. Underside black, punctured and covered with 
squamous hairs. 


Rhynchophorous Coleoptera of Japan. 73 


Very similar to P. perlatus, Chap., from which it may 
be distinguished by its smaller size, the absence of a 
carina on the epistoma, the frontal median elevation not 
reaching to the mouth, by the absence of a longitudinal 
elevation from base to apex of the thorax, by the 
shallower and evidently punctured elytral striz and the 
distinct rows of bristles on the interstices, which are 
alternate towards the apex; a similar row of scale-hke 
bristles occurring in P. perlatus cannot be distin- 
guished from the general covering with a lens. ‘The 
alternation of these bristles will also differentiate it 
from P,. lewist. 


Phleeosinus lewisi, Chap. 


Chap., Scol. Jap., p. 198. 
Numerous specimens, Kashiwagi, Chiuzenji, Kobe and 
Nowata. 


Varies in size from 1°8 to 2°3 mm. 


Phleosinus rudis, sp. n. 


Ovatus, subnitidus, niger, elytrorum apice nonnunquam dilu- 
tiore antennis tarsisque ferrugineis ; capite vix rostrato, oculis 
exacte planis, vix emarginatis ; prothorace transverso, basi in medio 
producta, lateribus rotundatis, versus apicem leviter sinuatis, supra 
convexo, dense et fortiter punctato, parcius piloso; elytris 
oblique declivibus supra breviter pilosis, striato-punctatis, inter- 
stitiis planis granulatis, versus apicem uniseriatim tuberculatis. 
Long. 2°7—3 mm. 

Mas. Interstitiis lo, 30 in declivitate tuberculis magnis, 
discretis ornatis, 2, 4 inermibus. 

Fem. Interstitiis lo, 30 in declivitate subtilius tuberculatis, 
20 subtilissime, 


Kashiwage and Kobe. 


Black, oval ; head strongly punctured, front flattened in female, 
impressed in male, almost glabrous, except for a few hairs over 
mouth and a few on vertex in female, eyes perfectly flat. Pro- 
thorax transverse, its base produced in middle towards scutellum, 
slightly contracted towards apex with sides rounded from base ; 
convex, somewhat shining, strongly and closely punctured without 
asperation, usually with a median shining longitudinal line. 


74 Mr. Walter F. H. Blandford on the 


Scutellum round, dull, punctured. Elytra scarcely wider than 
prothorax and rather less than twice as long, their basal borders 
rounded, crenate, slightly everted, sides straight at base, rounded 
from middle to apex ; above gradually declivous almost from base, 
black, with apex sometimes piceous, with short scanty pubescence, 
without scales or setose hairs, strix rather fine, indistinctly punctured 
at base, nearly smooth towards apex, interstices granulate to 
middle, then 2nd and 4th in male smooth, multi-punctate, with one 
or two tubercles before apex, in the female with a few fine 
tubercles throughout ; Ist in male with about 5, 3rd with about 7 
strong spinous tubercles, which do not unite to form a crest, in 
female with smaller tubercles : outer interstices towards apex with 
rows of tubercles in both sexes. Underside black, punctured, 
pubescent ; metasternum rather prominent. Legs black with tarsi 
lighter. 


The-largest Japanese species and the only one allied to 
the Huropean species in appearance and sexual characters. 


Potye@rapuus, Er. 


Of all the genera of the family, this one, considering 
its small extent, is the most difficult to deal with. Till 
recently it included but two species, P. poligraphus, Lin., 
and rufipennis, Kirby, but Thomson has added three 
others found in Hurope, of which I have seen but one, 
P. grandiclava, or rather an insect forwarded to me by 
Herr Reitter as P. poligraphus, var. grandiclava. I have 
also had before me an Indian species, and there are 
apparently three distinct forms from Japan. Well- 
marked differences are wanting amongst all these insects, 
which are almost identical in structure and sculpture ; 
and the difference of appearance which can be seen in 
comparison of examples vanishes in the most elusive way 
when an attempt is made to define and embody it in a 
description. 

It is possible that the structure of the male organs will 
be of assistance here, as Lindeman has shown that they 
are often distinct in closely-allied Scolytids. But for 
satisfactory examination of these, series of a certain length 
are required, and have not been forthcoming. I can 
therefore but echo the words of Leconte who, in treating 
of the almost equally difficult genus Dendroctonus, wrote : 
“Tf I have failed to indicate more strongly the differences 


Rhynchophorous Coleoptera of Japan. 75 


between these species it is because they are not dis- 
tinguished by any prominent or definite characters ; and 
the student, who may have difficulty in identifying the 
species as here defined, would have almost equal difficulty 


if the specimens in my collection were before him.” 


Polygraphus oblongus, sp. 0. 

Oblongus, subnitidus, niger, elytris apice rufescentibus, squamis 
flavo-cinereis vestitus ; clypeo emarginato, oculis subconvexis, 
antennarum clava oblonga infuscata acuminata ; prothorace antice 
fortiter constricto, linea media subelevata, punctis subaciculatis 
sat densis notato; elytris versus apicem subdilatatis, tenuissime 
striatis, ad basim granulatis. Long. 3°2 mm. 

Mas. Fronte subconvexa, breviter pilosa. 

Fem. Fronte convexa, bituberculata, brevissime pubescente. 


Four examples, Chiuzenji and Subashiri. 


Oblong, black, with elytra becoming gradually reddish towards 
apex. Head with front slightly convex in male, rugosely punctured 
and rather dull, pubescence short, in female convex, shining, very 
shortly pubescent, with two distinct tubercles in middle, below 
them with slight impression, clypeus emarginate in middle, eyes 
feebly convex, more distinctly in female, antennal club rather large, 
infuscate and evidently acuminate at inner side of apex, more 
strongly in male than in female. Prothorax nearly half as: broad 
again as long, strongly constricted towards apex with sides more 
convex behind constriction than at base; surface rather shining, 
with close subaciculate punctuation and thin squamous covering ; 
median line slightly elevated, variable in length and distinctness. 
Elytra more than half as long again as prothorax, one-third longer 
than wide, slightly but discernibly dilated towards apex, dull, 
scales close, cinereous with a yellow tinge, striz faint but dis- 
tinguishable throughout, weaker at apex in female than in male. 
Legs dark, ferruginous, with tarsi lighter ; all tibize spined at apex. 


When compared with LP. poligraphus, this species 
differs in its larger size, infuscate elytral club, thinner 
frontal pubescence, shorter and more constricted pro- 
thorax. The elytral apex is lighter in colour, and this 
does not appear to be due to immaturity. The punc- 
tuation of the head in the female is more distinct. 


Polygraphus proximus, sp. n. 


Antecedenti omnibus simillimus, sed brevior, elytris ad apicem 


76 Mr. Walter F. H. Blandford on the 


distinctius rufescentibus, non dilatatis, striis minus perspicuendis. 
Long. vix 3 mm. 


Two examples, Sapporo. 


The two specimens I have included under this title 
differ principally in shape from the preceding species. 
They are distinctly shorter, particularly in the elytra, 
which are not more than one-fourth longer than wide, 
and are not dilated towards the apex, which is slightly 
flattened. The head is similar, the antennal club less 
deeply infuscate. Prothorax transverse and strongly 
constricted, the median line well-marked and elevated at 
the base, more so than in the specimens of P. oblongus, 
where it is nearly obsolete, rather more shining and less 
closely punctured, the punctures subaciculate. Elytra 
more abruptly and distinctly reddish towards apex, their 
striz less distinct. Posterior tibiz feebly spined. 


One example is rubbed bare, and its appearance is 
materially altered, the pubescence on the other is not 
intact, but appears to be a little less close than in P. 
oblongus. Both are females. 


Polygraphus miser, sp. n. 

P, oblongo similis sed minor, angustior, antennarum clava vix 
acuminata, prothorace anterius minus angustato, post apicem haud 
constricto; elytris ad basim magis infuscatis, tenuiter striatis, 
squamis subtilioribus ; pedibus ferrugineis. Long. 2°4 mm. 


Four specimens, Nikko. 


Hardly to be distinguished from P. oblongus except by 
its smaller size, which is quite noticeable when the 
specimens from each locality are compared side by side. 


In shape narrower, with the prothorax less transverse and con- 
stricted in front, and the elytra proportionately shorter. Antenne 
entirely testaceous, with the club not acuminate; prothorax more 
shining, less squamous and more sparingly punctured, the punctures 
but feebly aciculate ; one example shows a median raised line, the 
other three not. Hlytra with basal half darker and more abruptly 
contrasted with the shorter and lighter apica] half, with rather 
thinner squamous covering, and more distinct striz. Head in the 
female duller, more punctured and less convex, front bituberculate; 
in the male narrower. 


Rhynchophorous Coleoptera of Japan. 77 


Scotytus, Geoff. 


One species has been described by Chapuis. I add 
five more, and there is possibly one other. They present 
the uniformity in appearance characteristic of this widely 
distributed genus. ‘The species were principally obtained 
from birch. 

TABLE OF SPECIES. 
1. Second abdominal segment unarmed . . . ...... 2 
with a median process . . claviger. 
2. Abdomen concave, its Srd and 4th segments with a tubercle 
onapicalmargin . . . ce ee 
not concave, 3rd and 4th eeamene simple 24 4 8 
3. Apex of elytra finely serrate near suture (size 44 mm.) . agnatus. 


SITE Oira/ Moa 7. ara eegets eee epee Mee ee 
4, Alternate interstices of elytra with an preatilan double row 
OL Punehures, 6 fh) Soa)  fnontalis: 


All interstices with a single row of punctures. . . . . . 5 

5. Elytra brown, strongly punctured inrows . . . . . aratus. 
black, finely punctured, with oblique aciculate 

scratches tee ed ee kts) a SAPONICUa OL: 


Scolytus esuriens, sp. n. 

Nitidus, niger, elytrorum lateribus et apice nonnunquam 
rufescentibus, antennis pedibusque ferrugineo-piceis ; prothorace 
longitudine vix latiore, punctato, linea media levi; elytris 
lateribus subrotundatis posterius vix angustatis, apice conjunc- 
tim subtruncatis, fortiter striato-punctatis, striis impressis, 
interstitiis planis subtiliter uniseriatim, 30 saltem_biseriatim, 
punctatis ; abdomine concavo, segmento lo margine prominulo, 
3o et 4o in margine medio tuberculo parvo ornatis. Long. 
38—5°5 mm. 

Mas. Fronte deplanata breviter pilosa. 

Fem. Fronte subconvexa parcissime pilosa, vertice subnitido, 
discrete punctato, Long. 3°8—5:'5 mm. 

Several specimens taken at Junsai, and single 
examples at Miyanoshita and Chiuzenji. 

Nearly allied to S. geoffroyi, Goetze. 

Black with sides and apex of elytra more or less reddish. 
Front of head flattened in male and not impressed, with short 
pubescence, subconvex in female, impressed over mouth, and 
thinly hairy ; vertex in female convex, with a median impressed 
line shining and diffusely punctured. Prothorax with a shallow 
impression on either side, more strongly punctured than in 


78 Mr. Walter F. H. Blandford on tle 


S. geoffroyi, the central impunctate line sometimes obsolete. 
Elytra not evidently narrowed behind, their apices conjointly 
rounded, nearly truncate, with striz deeper than in 8. geoffroyi 
and more strongly punctured, interstices somewhat less flat, finely 
punctured in rows, the 3rd and in larger specimens the 5th or 7th 
with irregular double rows ; punctuation of outer interstices strong 
at base. Abdomen with apical margin of Ist segment prominent, 
thickened in middle and shining, 2nd segment concave, 8rd and 
4th with a small median tubercle on margin, 5th with a longitu- 
dinal impression near apex. Metasternum shining, diffusely 
punctured. 


Variable in size, in the width and number of punctures 
of interstices, which are sometimes partly striate. 

One example from Junsai is smaller, with the elytra 
longer and narrower, and separately rounded at apex, 
the 8rd interstice has a double row of punctures at the 
base only, and the outer interstices are strigose with 
indistinct punctures. ‘he underside does not differ, 
though the tubercles are inconspicuous. It appears to 
be an ill-developed example. 


Scolytus agnatus, sp. n. 

Nitidus, niger, elytris apice rufescentibus, antennis pedibusque 
piceo-ferrugineis ; prothorace latitudine et longitudine fere equali 
in disco subtiliter, in lateribus fortius punctato ; elytris lateraliter 
subrotundatis, postice angustatis, in apice singulatim rotundatis, 
margine apicali subtiliter serrato, punctato-striatis, striis non 
impressis, interstitiis angustis subrugosis, singnlariter seriatim 
quam striis vix subtilius punctatis ; abdomine deplanato, segmentis 
30 et 4o muticis, Long. 4°6 mm. 

Fem. Fronte subconvexa, glabra, vertice obscuro, dense ac 


rugose punctato. 


Two specimens taken at Junsail in company with the 
last species. 

Very like S. esuriens, and readily confounded with it. 

Black, with apex of elytra reddish. Head in the female (I have 
not seen the male) quite glabrous, its vertex dull, rugosely and 
closely punctured. Prothorax similar, but without lateral impres- 
sions. Elytra larger, more evidently narrowed towards the apices, 
which are separately slightly rounded, the whole posterior margin 
finely toothed. Punctures of the elytral striz stronger, somewhat 
irregular and not distinctly impressed, interstices narrow, punc- 


Rhynchophorous Coleoptera of Japen. 79 


tured nearly as strongly as the striz in irregular series, which are 
not double, except at base ; somewhat rugose, and with slight 
longitudinal impressions. Punctures of the sides of the elytra 
much stronger than in S, eswriens. Abdomen not concave, margin 
of the Ist segment not prominent but sloped upwards, 3rd and 
4th unarmed, the 5th transversely impressed, Metasternum closely 
and rugosely punctured. 
Scolytus frontalis, sp. n. 

Oblongus, nitidus, niger, elytris rufo-brunneis, antennis pedi- 
busque ferrugineis ; capite (in maribus saltem) oblongo, fronte 
impressa, strigose aciculata, pilis longis crispatis circumdata ; 
prothorace haud transverso sat fortiter punctato, linea media 
laevi, punctis in disco paullo subtilioribus, magis diseretis ; elytris 
prothorace fere dimidio longioribus, lateribus subrotundatis, 
postice angustatis, subtiliter punctato-striatis, striis non impressis, 
interstitiis irregulariter vix subtilius seriato-punctatis, lo, 30, 50 
biseriatim, subrugosis ; abdomine subconvexo, margine segmenti 
Imi haud prominulo, cum ceteris mutico. Long. 3°6—4°3 mm. 


Three examples taken at Fukushima. 


Resembling S. prun?, Ratz., in appearance, but with the pro. 
thorax more strongly punctured, and the elytral striz: much less 
regular. Black with anterior margin of thorax and elytra red- 
brown. Head (in the male, to which sex the specimens belong) 
oblong, produced in front, and impressed, with long aciculate 
scratches from mouth to vertex, margined with long curled yellow 
hairs, median line finely carinate, epistoma subcircularly emargi- 
nate. Prothorax about as long as broad with moderately strong 
oval punctures, closer and deeper at sides. Elytra as wide in 
middle as prothorax, and nearly one-half longer, narrowed behind, 
rather abruptly rounded at apex and feebly emarginate at suture ; 
surface little depressed round scutellum, with fine irregular rows 
of punctures, interstices narrow and flat, with slight transverse or 
oblique wrinkles, their punctures little finer than and not readily 
distinguishable from those of striz, forming an irregular double 
row on Ist, 3rd and 5th interstices, and a single row on remainder, 
except at base where they are irregular. Underside piceous, lighter 
at sides, thinly pubescent ; abdominal segments simple, margin of 
1st not prominent. ; 


Scolytus aratus, sp. n. 


Nitidus, niger, elytris piceo-brunneis, basi cum pedibus rufescente; 
capite convexo, subtiliter aciculato, parcissime piloso; prothorace non 


80 Mr. Walter F. H. Blandford on the 


transverso, fortiter punctato, punctis ovatis in medio disco magis 
discretis, subtilioribus ; elytris prothorace dimidio longioribus, 
lateribus subrotundatis, postice angustatis, margine apicali sub- 
truncato, fortiter lineato-punctatis et substriatis, interstitiis 
angustis uniseriatim punctatis, punctis vix subtilioribus, seriebus 
nonnunquam irregulariter impressis ; abdomine convexo, mutico, 
piloso. Long. 5 mm. 

Mas. (?). Abdominis segmento 5o pilis longis ornato. 

Fem. (?), Abdominis segmento 50 breviter piloso. 


Junsai, two specimens. 


Allied to S. carpini, Ratz., but with stronger thoracic punctua- 
tion, and elytral striz more impressed with coarser punctures. 
Black with elytra pitchy-brown. Head convex, finely aciculate 
and glabrous, without sexual differences; vertex dull, closely punc- 
tured. Prothorax as long as broad, with strong oval punctures, 
scattered on disc, closer, deeper and rugose at sides and apex. 
Elytra as wide as thorax and one-half longer, with lateral 
margins slightly rounded to behind middle, then narrowed ; apical 
margin nearly truncate, feebly excised at suture, and slightly 
irregular, but not serrate as in S. agnatus ; surface with regular 
rows of strong punctures, deeper and closer at base, slightly 
impressed throughout, interstices narrow, with a single series of 
punctures rather weaker than those of strive, and sometimes im- 
pressed, first and second with a few oblique wrinkles. Abdomen 
convex, covered with short hairs; last segment in male (?) emar- 
ginate at apex and impressed, with a fringe of long hairs. 


Smaller than any other Japanese species except 
S. japonicus, which is readily distinguished by its black 
colour, and the absence of striate impressions on the 
elytra, the punctures of which are fine and obscured by 
oblique wrinkles. 


Scolytus japonicus, Chap. 
Chap., Scol. Jap., p. 199. 


A single further specimen of this species, the repre- 
sentative of the European S. rugulosus, taken at Junsai. 
It attacks the plum-tree. 


Scolytus claviger, sp. n. 
Subelongatus, depressus, niger, elytris piceis, antennis pedi- 
busque rufescentibus ; prothoracis disco parce punctato, punctis 
versus basim subtilioribus ; elytris regulariter striato-punctatis, 


Rhynchophorous Coleoptera of Japan. 81 


striis vix impressis, interstitiis paullo subtilius uniseriatim vel 
irregulariter biseriatim punctatis, depressione apicali ad latera 
serie tuberculorum intus marginata. Long. 3°7 mm. 

Mas. Fronte impressa, longius pilosa ; abdominis segmento 20 
processu oblongo deplanato versus apicem incrassato et recurvato 
armato. 

Fem. Fronte subconvexa, parce pilosa; abdominis segmento 
20 tuberculo brevi conico armato. 

A pair taken at Kiga. 

Elongate and depressed, black with elytra piceous. Head with 
front strongly impressed in male and margined with long curled 
hairs, in female subconvex and strigose with scanty pubescence ; 
vertex strongly punctured in both sexes. Prothorax as long as 
broad, with sides nearly parallel behind middle, not strongly 
contracted in front, its punctuation rather strong, not close, weak 
over base only. Elytra as wide as prothorax and one-half longer, 
very slightly rounded at sides and narrowed behind, apex serrate 
and feebly emarginate ; surface with regular impressed rows of 
punctures, interstices narrow with subimpressed rows of rather 
weaker punctures, tending to become double on alternate inter- 
stices. Apical impression extending along sides of elytra and 
bordered within, above outer angles, by six or seven small 
tubercles. Underside black, abdomen reddish at sides, its first 
ventral segment prominently bordered, second in male with an 
oblong process, flattened vertically, its tip thickened and recurved 
upwards; in female with a short pointed tubercle, and rather 
dense pubescence. | 


Alhed to S. multistriatus, but differing in the structure 
of the abdomen, and the stronger punctuation, the thora- 
cic punctures being more scanty. The apical segments 
of the abdomen, nearly horizontal in the male example, 
are unfortunately missing in the female, which prevents 
comparison with another specimen from Nishi, which 
appears distinct, but cannot be separated by any definite 
characters ; it 1s rather larger, more convex, the thorax 
is more evidently narrowed in front, with the sides 
rounded behind ; the elytra are darker and broader; the 
abdomen is nearly vertical, armed as in the female type. 
This insect does not show any differences in sculpture 
that are of specific value, though the thorax is more 
finely punctured behind the middle. It must be sepa- 
rated, if at all, on further examples of both sexes. 


TRANS. ENT. SOC. LOND. 1894,—PART I. (MARCH.) F 


82 Mr. Walter F. H. Blandford on the 


TOMICINI. 


Cryprureus, Er. 

C. pusillus, Gyll, has ie been recorded from 
Japan by Hichhotf (Rat. Tom., p. 74), who examined 
specimens taken by Hiller in Nipon, and could discover 
no difference between them and Huropean examples. 
-Mr. Lewis has taken a series on Fujisan and at Subashiri 
in which the pubescence on the elytra is scanty, fine 
and less distinguishable than in Kuropean specimens, but 
which have no other well-marked distinguishing fea- 
tures. ‘The prothorax is rather strongly narrowed 
behind, but its shape is variable in Huropean examples. 


Crypaauus, Er. 


At present the Japanese Cryphali are represented 
by a single specimen only in Mr. Lewis’s collection. As 
the genus is very common in the Oriental region, and 
there are some ten Huropean species, it is probably much 
more numerous in Japan than it appears to be at present. 


Cryphalus exiquus, sp. n. 


Oblongo-ovalis, convexus, opacus, niger, squamosus ; prothorace 
semi-orbiculato, margine antico bituberculato, anterius tuberculis 
discretis in plaga postice producta exasperato, posterius rugose 
punctato ; elytris latitudine longioribus, lineato-punctatis, inter- 
stitiis squamosis setis erectis raris ornatis. Long. 1:3 mm. 


One example, Fukushima. 
One of the smallest species in the genus. 


Oblong-oval, convex, dull black with a covering of grey scales. 
Head with front subconvex, reticulate, punctured at sides, im- 
pressed over mouth and with an obtuse median elevation separated 
above from vertex by a sharp transverse shining carina; eyes 
oblong-oval, anteriorly emarginate ; antenne testaceous with club 
deeply infuscate, roundish oval, its basal joint shining, with- 
superior apical border rounded, fringed, remaining joints pilose 
with border less strongly rounded. Prothorax narrowed towards 
apex, rather broader than long, its base bisinuate with narrowly 
elevated margin, basal angles rounded when seen from above, 
sides and apex rounded throughout, the latter more strongly, its 


Rhynchophorous Coleoptera of Japan. 85 


margin with two prominent tubercles in middle; surface uni- 
formly convex, not gibbous, anteriorly with scattered asperate 
elevations, forming a patch angulately produced behind but not 
reaching very near base, interstices and remainder of surface finely 
reticulate and rugosely punctured, with thin covering of scales 
and hairs. Scutellum very small, triangular. Elytra as wide as 
prothorax, and not quite half as long again, slightly and separately 
rounded at base, humeral angles rather obtuse, but not rounded, 
shoulders finely elevated, sides nearly straight and subparallel to 
posterior third, then broadly rounded to apex ; surface transversely 
convex, subcylindrical to behind middle, then obliquely decli- 
vous and convex to apex, with distinct rows of punctures, the 
outer ones obsolete towards apex, interstices narrow, finely 
multipunctate and squamous, with asingle series of scattered erect 
setee. Legs fuscous with tarsi testaceous, 


This species can be distinguished from other described 
Cryphali, in which the eyes are emarginate and the elytra 
not rounded from base to apex, by the two tubercles 
on the anterior margin of the prothorax. Amone the 
European species it is most like U. abietis, Ratz, but is 
much smaller, with the elytral stris not impressed, and 
the antennal club and legs darker. The transverse 
carina which borders the vertex of the head in front 
occurs in some Ceylonese species in my hands, and is 
probably a sexual character. As with other Oriental 
Oryphali, the joints of the antennal club have a rounded 
margin on their superior surface, which is less evident 
below. In this respect they approach Thomson’s sub- 
genus Hrnoporus, which can be characterised by the non- 
emarginate eyes alone. 


HyrorHenemus, Westw. 


I take this genus, as has been done by Leconte and 
others, to be inclusive of Hichhoft’s genus Stephano- 
deres, which it is not possible to separate by means of 
the number of articulations in the antennal funiculus. 
But it is not to be supposed, as Leconte has suggested, 
that the variations of this structure are of no generic 
value in Scolytide. In the majority of genera which 
can be separated by them, there are other structural 
features, or at least a difference of facies, which tend to 
show that they are a reliable guide. In the Hypothenemi, 


84. Mr. Walter F. H. Blandford on the 


however, the smallest insects of the family, it would appear 
that reduction in size leads to a reduction in the number 
of joints, without involving other structural features. 
But we really require to study its structure in more 
species of this difficult and little-known genus. 

I add two more species to the one already described 
from Japan. 


Hypothenemus tristis, Kichh. 


Stephanoderes tristis, Kichh. Scol. Jap., p. 200; Rat. 
Tom., p. 150. No further examples have been taken, 
unless the following insect be a variety of this species. 


Hypothenemus peritus, sp. n. 

Oblongus, subnitidus, fusco-piceus, prothorace in medio 
dilutiore, semi-ovato, antrorsum angustato, margine antico tuber- 
culis 4, duobus mediis majoribus, ornato, anterius tuberculis 
magnis discretis notato, postice rugulose punctato ; elytris ad 
apicem subacuminatim rotundatis, subtiliter lineato-punctatis et 
setosis, interstitiis uniseriatim squamulatis. Long. 1:8 mm. 

One specimen, Nagasaki. 

Oblong, rather shining, deep fuscous, with surface of prothorax 
lighter in middle. Head finely reticulate, front subconvex, punc- 
tured in middle. Antenne sordid testaceous, funiculus 5-jointed, 
suture of club distinctly fringed. Prothorax very slightly 
broader than long, base subsinuate, basal angles obtusely rounded, 
sides slightly dilated before base and rounded, contracted towards 
apex and obscurely sinuate, apical margin with four tubercles in 
the middle, the two centre ones more prominent ; surface obtusely 
gibbous in middle, with thin covering of hairs and scales, anteriorly 
with a roundish patch of large scattered tubercles, becoming closer 
and smaller behind, interstices and base finely reticulate with 
scattered punctures, except over a narrow smooth line from base 
to middle. Elytra rather more than half as long again as 
prothorax, separately rounded at base, humeral angles rounded, 
sides parallel to posterior third, thence obliquely rounded to 
apex ; surface subcylindrical, obliquely declivous and convex 
at apex, with fine lines of punctures bearing minute hairs, the 
lines appearing impressed in certain light, interstices transversely 
rugose, with a single row of cinereous scales along middle and a 
few hairs on either side. Legs infuscate with anterior femora and 
tarsi lighter. 


Quite like H. tristis, Hichh., but differing from the type 


Rhynchophorous Coleoptera of Japan. 85 


in the fewer and stronger tubercles of the prothorax, the 
more scattered punctuation of its base, which presents a 
smooth central line, and the finer elytral striz, which in 
H., tristis are strong and well-marked. It is also rather 
more elongate. 


Hypothenemus expers, sp. 1. 

Oblongus, subnitidus, piceo-niger, prothorace medio dilutiore, 
subrotundato, ante basim dilatato, margine antico bituberculato, 
dorso anterius tuberculis discretis in plaga rotundata compositis 
notato, posterius granulato; elytris lineato-punctatis, pilosis, 
interstitiis setis haud squamatis seriatis ornatis. ong. vix 2 mm. 

Two examples, Kumamoto and Nagasaki. 


Closely allied to H. peritus, but exhibiting the 
following differences :— 

Rather larger and more robust, darker in colour, being black 
with a pitchy tinge, the thorax pitchy-red in middle, and not 
obscurely testaceous. Front of head more convex, very finely 
reticulate and not punctured except over mouth, where it is trans- 
versely impressed. Antenne bright testaceous, basal joint of club 
deeply infuscate, funiculus 5-jointed. Frothorax more strongly 
dilated before base, which is truncate, sides more strongly rounded, 
its tuberculation similar, but marginal tubercles less prominent, the 
two outer ones nearly obsolete, basal half closely reticulate and 
covered with small asperate elevations, closer than the non-elevated 
punctures in H, peritus. Elytra narrower than greatest width of 
prothorax, truncate at base, obliquely rounded at apex, striation 
as in H. peritus ; interstices without evident scales, but with a 
single row of erect hairs, which are stronger towards apex, a few 
finer hairs are found between the rows. Legs clear testaceous. 


The absence of the conspicuous erect hispid scales on 
the elytra, which are replaced by hairs, will at once 
distinguish this species from either of the two former. 


Cosmopgrzs, Hichh. 


This genus was founded by Hichhoff, Rat. Tom., p. 495, 
for OC. monilicollis, % single species from India, which 
differs from Hypothenemus (Stephanoderes, Kichh.) in 
possessing a very short two-articulate antennal funiculus 
and dilated tibiae with their outer borders serrate, the 
serration being absent in Hypothenemus; the typical 
species is also more elongate than a Hypothenemus, and 


86 Mr. Walter F. H. Blandford on the 


differs in colour, and the sculpture of the elytra, which 
have large dilated punctures and subcostate interstices, 
so that they appear reticulate. There is in Mr. Lewis’s 
Ceylon collection a specimen which corresponds in every 
respect to Hichhoff’s description of C. monilicollis, and 
is almost certainly that insect. In the Japanese collec- 
tion is a specimen which agrees with it in generic 
characters, but is much more closely allied in appear- 
ance to Hypothenemus, showing the near relationship 
between the genera, which are, however, sufficiently 
distinct. 


Cosmoderes consobrinus, sp. u. 


Oblongus, subcylindricus, opacus, niger, elytris nigro-piceis, 
antennarum scapo tarsisque testaceis ; prothorace aeque longo quam 
lato versus apicem subangustato, apice rotundato, bituberculato, 
disco gibboso, anterius exasperato, posterius scabrose granulato ; 
elytris prothorace duplo longioribus, seriatim pilosis et striatis, 
striis leniter impressis, vix perspicue punctatis, interstitiis alutaceis 
uniseriatim squamosis. Long. 2 mm. 


One example, without locality. 


Oblong, rather elongate, cylindrical, black, with elytra obscurely 
piceous. Head finely reticulate, front convex, finely and sparingly 
punctured, and very shortly pubescent, eyes oval emarginate ; 
antenne testaceous with club infuscate, short oval, pubescent with 
a smooth patch at base, without evident sutures. Prothorax as 
broad as long, contracted at sides towards apex, base truncate not 
margined, basal angles obtuse, sides slightly rounded behind, sub- 
sinuate and more rounded anteriorly, apical margin strongly 
rounded with two prominent median tubercles ; surface convex 
and gibbous in middle, obliquely impressed on either side behind 
median elevation, in front strongly declivous and rather finely 
asperate, base and interstices dull, covered with fine scabrous pro- 
jections, pubescence very short. Scutellum distinct, triangular. 
Elytra a little wider than base of prothorax and double as long, 
base truncate, humeral angles rounded rectangular, shoulders 
rather prominently elevated, sides subparallel, obliquely rounded 
at apex ; surface subcylindrical, very obliquely declivous towards 
apex, its texture coriaceous, with shallow impressed striz, with 
barely distinguishable punctures and rows of minute hairs, inter- 
stices alutaceous, subconvex, with a single series of erect narrow 


Rhynchophorous Coleoptera of Japan. 87 


scales. Legs fusco-piceous with tarsi lighter, outer border of 
middle tibiz distinctly serrate. 


This insect is very like Hypothenemus tristis and 
peritus, but is more elongate; the base of the thorax 
and the elytra are differently sculptured, and the struc- 
ture of the antennz and legs will at once separate it 
without comparison. 


PityopntuHorvs, Kichh. 


But one species has yet been found in Japan. It is 
likely that there are others as, in addition to the Euro- 


pean species, there are several from both N. and 8. 
America. 


Pityophthorus jucundus, sp. n. 


Elongatus, oblongus, nitidus, fere glaber, niger vel fusco- 
testaceus ; prothorace antice subconstricto, post medium utrinque 
impresso, anterius exasperato, posterius fortiter punctato, linea 
media subelevata laevi; elytris cylindricis, haud acuminatis, 
regulariter punctato-striatis, striis non impressis, interstitiis fere 
planis hic illinc subrugulosis, apice impresso-retuso, striis obsoletis, 
lateribus elevatis et sutura versus apicem tuberculis setigeris raris 
ornatis. Long. 1°6 mm. 

Fem. Fronte media villosa. 


Four specimens, near Nagasaki. 


Head black, finely reticulate, front strongly punctured, with a 
circular patch of villous pubescence in female. Antenne fusco- 
testaceous, their club ovate, tri-articulate with slightly curved 
sutures. Prothorax with base finely margined, truncate, basal 
angles obtuse, sides behind nearly straight, rounded in front and 
sinuate before apex, which is somewhat obtusely rounded, and 
feebly crenate, the apical constriction much slighter than in 
P. lichtensteini, Ratz; surface depressed on either side of a median 
smooth elevated line, reaching to the middle of the prothorax, 
with very short pubescence at sides and apex, its anterior half 
asperate, posterior half with rather strong subrugose punctures 
becoming weaker on sides, Elytra rather narrower at base than 
prothorax, and nearly twice as long, sides straight to middle then 
feebly rounded, apex obtusely rounded, not acuminate but with 
suture slightly prominent ; glabrous except at apex, with regular 
lines of strong punctures, not impressed, interstices impunctate and 
flat, feebly rugose here and there, apex nearly vertically declivous, 


88 Mr. Walter F. H. Blandford on the 


impressed on either side of suture, the impression shining, impunc- 
tate, its outer margins more strongly elevated than suture, with two 
- or three weak setigerous tubercles ; suture very slightly elevated, 
with traces of tuberculation near apex; underside black, thinly 
pubescent. Legs fuscous. 


A slender narrow species, readily distinguished by the 
strong punctures of the thorax, with its elevated median 
smooth line, by the distinct and regular rows of 
punctures on the elytra, which are nearly glabrous and 
not acuminate. 


Eivoryenus, Hichh. 
I add one more species provisionally to this genus. 


Eidophelus imitans, Eichh. 
Eichh., Scol. Jap., p. 200; Rat. Tom., p. 203. 


One example, very small, has occurred since, without 
recorded locality. 


Hidophelus minutus, sp. n. 


Oblongus, nitidus, piceo-niger, fere glaber ; prothorace lati- 
tudine vix longiore, lateribus ad medium subrectis, inde cum apice 
subcirculare rotundatis, supra convexo, antice lineis tuberculorum 
concentricis exasperato, postice punctis discretis fortibus, in medio 
subtilioribus notato; elytris ad apicem oblique declivibus, non 
impressis, lineato-punctatis, punctis post medium obsoletis, inter- 
stitiis subrugosis absque punctis. Long, 1°2 mm. 


One example, Chiuzenji. 


Oblong, piceous-black, with a few hairs on elytra and sides of 
thorax. Head reticulate, front strongly punctured, scantily hairy ; 
eyes oval, scarcely visibly emarginate ; antenne testaceous, funi- 
culus 4-jointed, club orbicular oval with weak curved sutures, 
pubescent at apex. Prothorax little longer than broad, base 
truncate, finely margined, basal angles obtuse but distinct, margined, 
sides straight, slightly narrowed towards middle, then subcircularly 
rounded, apex slightly flatter ; its anterior half with about six 
concentric rows of fused tubercles, forming a series of raised lines 
somewhat as in Cryphalus tilie, the anterior row occupying the 
apical margin when seen from above, posterior half with very 
scattered punctures, strong at sides, smaller and somewhat as- 
perate on middle Scutellum triangular, shining. Elytra rather 


Rhynchophorous Coleoptera of Japan. 89 


narrower than prothorax, and half as long again ; base truncate, 
its angles obtuse, sides weakly rounded to middle and subdilated, 
thence more strongly rounded to apex; surface with rows of 
rather small punctures, separately impressed and not striate, 
becoming weaker from base and nearly obsolete on declivity ; 
interstices slightly rugose, with one or two punctures at base ; apex 
obliquely declivous, convex and not impressed, with traces of one 
or two asymmetrical tubercles near suture. Underside piceous, 
scantily pubescent. Legs infuscate. 

This insect agrees with H. initans in its antennal 
structure, but is separable by the absence of distinct 
pubescence and the sparseness of the punctures on the 
hinder half of the thorax. The entire absence of any 
thoracic constriction and the antennal structure will 
distinguish it from the Pityophthori. 


Tomicus, Latr. (1807). 


The Japanese fauna appears to be poor in this genus ; 
perhaps other species remain to be discovered. 


Tomicus cembrae, Heer. 
Bostrichus cembrae, Heer, Obs. Ent. 1836, p. 28. 
A series taken from larch on Fujisan. 


The specimens show no essential difference from 
European examples. Like the Japanese Myelophilus 
piniperda, they run very large, averaging 0-5 mm. ‘The 
interstitial punctures of the elytra are very weak, but 
traceable. 1’. cembrae is recorded from Siberia and 
Amurland and no doubt occurs over the whole of N. 
Asia; in Europe it is confined to Pinus cembra, which 
is found in Japan, at least, as the variety puwmilus. 
I cannot find any Japanese specimens which correspond 
with Motschulsky’s 7. subelongatus. 


Tomicus angulatus, Hichh. 
Hichh., Scol. Jap., p. 200; Rat. Tom., p. 208. 
Taken commonly in several localities from fir (Pinus 
massomand); Nagasaki, Fujisan, Nikko. 


ACANTHOTOMICUS, nov, gen. 


Antennarum funiculus 5-articulatus, articulis 2—5 latitudine 
crescentibus, clava ovalis, compressa, suturis fere obsoletis fortiter 


90 Mr. Walter I’. H. Blandford on the 


curvatis articulo 1o ovali, ceteris lunatis. Prosternum processu nullo. 
Episterna metathoracica linearia. Tibiee anticee ad apicem dilatate, 
spinose. Corpus cylindricum, elytris ad apicem retusis, ambitu 
retusionis fortiter multi-spinato, stria suturali vix impressa. 


Ciosely allied to Tonicus but differing in the much 
stronger elytral armature, which in Yomicus exhibits a 
uniformly progressive diminution, as the species grow 
smaller, down to the genus or subgenus Pityogenes, 
Bedel. The antennal club is oval, its sutures are quite 
superficial and only marked by a pubescent border, the 
first jot is longitudinally oval and embraced at the sides 
by the succeeding joint, as in \ylocleptes, Ferrari, with 
which the elytral structure has no affinity. The mouth 
parts do not differ from these of Vomicus, the maxilla is 
simply sinuate internally, and not produced into a 
rounded angle, but this angle is wanting in Pityogenes 
and is not a generic character. 


Acanthotomicus spinosus, sp. n. 


Oblongus, cylindricus, nitidus, ferrugineo-testaceus, elytris ad 
apicem infuscatis, pilis longis parce adspersus ; prothorace antice 
constricto, ad medium subgibboso et utrinque impresso, anterius 
exasperato, posterius subtiliter punctulato; elytris prothorace 
dimidio longioribus, punctato-striatis, punctis ad basin subtilibus, 
versus apicem dilatatis, ad apicem abrupte declivibus, utrinque 
8-spinatis, spinula 3a maxima, elongata, 5a—Va minimis, 8a prope 
angulum suturalem magna, intus curvata. Long. 2°5 mm. 


Two specimens taken at Oyayama and Nikko. 


Oblong, cylindrical, reddish-testaceous, with apex of elytra in- 
fuscate. Head with front subconvex, punctured, with a median 
raised line, thinly pubescent ; eyes broad oval, slightly emarginate. 
Prothorax longer than broad, its base truncate, basal angles obtuse, 
sides almost parallel to middle, then constricted and strongly 
rounded to apex, which is somewhat more obtuse, with a crenate 
margin; disc gibbous in middle and impressed on either side, 
asperate anteriorly behind with fine scattered punctures without a 
median impunctate or elevated line. Scutellum small, rounded, 
elytra half as long again as prothorax, their base truncate, 
sides subparallel, with lines of punctures, fine at base, becoming 
stronger, dilated and transversely rugose towards apex, the strix 
appearing impressed towards apex in a certain light only, interstices 


Rhynchophorous Coleoptera of Japan. 91 


slightly convex, very finely and sparingly punctured in rows ; apex 
nearly vertically declivous, margined with 16 spines at the apex of 
the 2nd and succeeding interstices, the 1st two short, 3rd very long, 
curved, 4th and 5th rather shorter, 6th and 7th small, 8th pair 
situated close to apex of suture, curved inwards and as long as 
fourth. Impressed surface rather dull, irregular, strongly punctured, 
with suture elevated. Underside testaceous, glabrous. Antenne 
and legs testaceous. 


The following specimen is probably the female of the 
preceding insect, but I cannot be positive. 


Acanthotomicus spinosus, ? femina. 


Antecedenti differt fronte plana utrinque densissime aureo- 
villosa, prothorace paullo longiore, minus constricto, margine 
antico fortius rotundato, haud crenato, dense villoso; elytris pro 
portione elongatis, prothorace duplo fere longioribus, punctis 
striarum paullo fortioribus et magis confertis, ad apicem singulatim 
breviter 6-spinosis, infra prope angulum suturalem in lobum 
brevem 4-tuberculatum productis ; fundo retusionis punctis mi- 
noribus, confertis, et spinula intra spinam tertiam marginalem 
posita utrinque ornato. Long. 3mm. 


One example taken at Kashiwagi. 


In the dense villosity of the head and prothorax, and 
the reduction in size of the apical spines of the elytra, 
its characters may be considered as simply sexual. But 
the greater length of the elytra and the closer and 
stronger punctuation of the striz, the sutural stria being 
impressed at the apex, together with the presence of a 
spine on the apical declivity internal to the 3rd marginal 
spine, prevent my identifying it positively as the female. 
The apical spines are all short, the 3rd and 5th being 
the most prominent; the lower border is produced on 
either side just outside the suture, into a short transverse 
lobe, which is evidently 4-tuberculate and represents the 
three inferior spines fused. 


Dryocatss, Hichh. 


This genus, in which I include Hichhoft’s Taphrorychus 
apatordes and two allied species, is represented by eight 
species, of which six are new. 


92 Mr. Walter F. H. Blandford on the 


TABLE OF SPECIES. 


1, Prothorax gradually and uniformly rounded from basetoapex 2 
cylindrical at base with apex obliquely declivous 

(Taphrorychus, Hichh.) . . . . eee 6 

2. Apex of elytra retuse, impressed or flattened (size 2. 5m 4mm) 3 

convex (size under 2°5 mm.) 5 

4 


3. Front of head not carinate 7 Py oily Webs 
with a sharp loneriuaole carina . . . afinis. 

4, Strivz of elytrafinely and uniformly punctured, autographus, Ratz. 
coarsely punctured, seein 3 dilated behind 


base: 0.7 <2 kts, ie pum NO RNs ESPUILORDER: 

5. Elytra with strongly punctured eee ser. hl Sy cet ene OBIE 

rows of feeble punctures. . . . . . . mubilus. 

6. Prothorax rounded at apex .. . : gos, wOeRIIE. 
subquadrate, the apex perce: forming an angle 

with sides’. “< 2 5 ue e ek <i 


“J 


. Interstices of elytra not elevated towered apex . dinoderoides, 
sub-costate towards apex, apatoides, Eichh. 


Dryocetes autographus, Ratz. 


Ratz., Forst. Ins., 1., p. 160. 

Hichh., Berl. Ent. Zeit., 1864, p. 39 ; Rat. Tom., p. 281. 

Two examples, Chiuzenji. 

The elytra are almost hairless in these specimens and 
very evidently retuse at apex, with the interstitial 
punctures almost obsolete; but they are quite like 
German specimens in my collection. ‘he species, which 
extends throughout N. Asia and the colder parts of N. 
America, is of variable form. 


Dryocetes pilosus, sp. 0. 

Oblongus, subnitidus, longe pilosus, piceo-niger, antennis 
pedibusque ferrugineis, fronte plana haud carinata; prothorace 
dense granulato exasperato ; elytris prothorace dimidio longioribus, 
transverse rugosis, fortiter punctato-striatis, punctis postice 
subtilioribus, stria suturali subimpressa, interstitiis subtiliter 
uniseriatim punctatis, apice retuso impresso, striis fere obso- 
letis, pilis longis cireumdato. Long. 3 mm. 


One example, Nikko. 


Oblong, pitchy-black, head with front flat, dull, finely punctured, 
and with a faint transverse impression at sides, with long scattered 
hairs, mouth fringed. Prothorax a trifle broader than long, base 


Phynchophorous Coleoptera of Japan. 93 


truncate, humeral angles obtusely rounded, sides and apex rounded 
in a broad ellipse; surface very convex, with uniform scattered 
granulations, their interspaces shining, fringed at margins with 
long scanty hairs. Scutellum small, rounded, convex, shining. 
Elytra narrower than greatest width of thorax, and half as long 
again ; base truncate, humeral angles obtusely rounded, shoulders 
narrowly raised, sides subsinuate, slightly dilated behind and 
strongly rounded at apex ; surface subcylindrical, depressed before 
middle, dilated posteriorly and strongly declivous at apex, with 
rows of strong punctures, the punctures dilated on middle third and 
then becoming finer, sutural stria slightly impressed, interstices sub- 
convex, transversely rugose, with a single series of fine setigerous 
punctures; apical declivity convex, impressed on either side of 
suture, shining, with strive obsolescent ; elytra set with long hairs, 
close at sides and at margins of apical declivity, hairs of declivity 
itself shorter. Underside piceous, punctured ; abdomen with long 
pubescence. Legs ferruginous. 


Readily distinguished by the long pubescence, the 
absence of a frontal carina, the strong punctuation of 
the elytral striz, and the punctured interstices. 


Dryocetes affinis, sp. n. 

Oblongus, niger, subnitidus, pilosus ; fronte carinata ; prothorace 
vix longiore quam latiore, granulate exasperato, postice paullo subti- 
lius ; elytris prothorace minus quam sesqui longioribus, cylindricis, 
apice oblique declivi subretuso, subtiliter lineato-punctatis, stria 
suturali impressa, interstitiis subrugosis seriatim pilosis. Long. 
3mm. 


A single example, Oyayama. 


Black, elytra with a piceous tinge. Head with front impressed 
and longitudinally carinate, coarsely punctured and thinly hairy, 
ciliate over middle of mouth. Prothorax a little longer than 
broad, truncate at base with angles obtuse, sides feebly rounded 
behind and very slightly dilated to posterior third, strongly and 
uniformly rounded in front to apex, the margin of which is 
tuberculaie; surface uniformly convex, asperate, rather more finely 
at base, with scattered hairs. Scutellum small, rounded, shining, 
finely bordered. Elytra rather wider than base of prothorax and 
about one-fourth longer, truncate at base with shoulders narrowly 
elevated, humeral angles rounded rectangular, sides rounded and 
somewhat narrowed at posterior fourth, then rather abruptly 
flexed, and nearly straight at apex, which is not conjointly 


94. Mr. Walter F. H. Blandford on the 


rounded ; surface subcylindrical, obliquely declivous behind, with 
rows of shallow punctures, their interspaces rugose, sutural stria 
alone slightly impressed, interstices rather narrow with series of 
setigerous tubercles ; apex flattened, shining, impressed on either 
side of suture with the stris continued on it; first interstice 
widened. Underside piceous-black, nearly glabrous, first abdo- 
minal segment impunctate in middle, succeeding segments with 
a few very coarse punctures. Legs piceous, anterior tibiz 
rounded, with six or seven outwardly directed spines on outer 
margin, and a curved one at inner angle of apex ; middle tibix 
with a close series of six forwardly directed spines at apex. 


In sculpture this insect is exactly like the one I have 
identified as Coccotrypes graniceps, Kichh. (q.v.), but 
may be distinguished by its cylindrical shape and the 
structure of the tibiz. 


Dryocetes luteus, sp. n. 

Oblongus, subnitidus, flavo-testaceus, parce et breviter pilosus ; 
prothorace oblonge, lateribus postice subparallelis, antice fortiter 
rotundatis, anterius subtiliter exasperato, posterius dense punctato; 
elytris profunde striato-punctatis, stria suturali impressa, inter- 
stitiis subtiliter uniseriatim punctatis in declivitate tuberculatis, 
Long. 2 mm. 


Fem. Fronte densius pilosa. 


Nine specimens, without locality indicated. 


Bright yellow-testaceous, with short scanty pubescence on sides 
of thorax and apex only of elytra. Head strongly punctured, 
with front convex, scantily hairy in male, and ciliate over mouth; 
in female* with a circular tuft of yellow pubescence. Eyes broad 
oval, coarsely granulate, and emarginate anteriorly. Antenne 
testaceous with sutures slightly curved. Prothorax longer than 
broad, with base truncate, its angles obtuse, sides nearly parallel 
to middle, thence strongly rounded to apex; above moderately 
convex, not gibbous, with close punctures, simple at base, and as- 
perate over anterior two-thirds. Scutellum rounded, shining. Elytra 
as wide as prothorax and one-half longer, base truncate, humeral 
angles rectangular, sides parallel to apex, then abruptly flexed, 
their apical margin being almost transverse when seen from above ; 


** Tt would appear that in the J’omicé the possession of a villous 
front is a mark of the female sex, see Eichh., Eur. Bork., p. 203, It 
is not so in the genus Scolytus, and others. 


Rhynchophorous Coleoptera of Japan. 95 


surface cylindrical, nearly vertically declivous at apex, but convex, 
with regular impressed rows of strong punctures dilated after the 
base, sutural stria more deeply impressed and widened behind, 
interstices flat with a single row of fine punctures, replaced by 
small setigerous tubercles on declivity. Underside and legs 
testaceous, the former scantily punctured and pubescent. 


Similar to immature examples of D. villosus, Fabr., 
but twice as small, with the prothorax not asperate 
behind, with scantier pubescence, and the elytral in- 
terstices more distinctly defined. 


Dryocetes nubilus, sp. n. 

Oblongus, subnitidus, parce longius pilosus, piceo-niger, an- 
tennis pedibusque testaceis ; prothorace oblongo-ovali anterius 
rugulis transversis exasperato, posterius subnitido, rugulose punc- 
tuto; elytris ad apicem oblique declivibus, subtiliter lineato- 
punctatis, stria suturali non aut vix impressa, interstitiis post 
medium uniseriatim pilosis. Long, 1-7—2-2 mm. 


Ten specimens, Kiga, Suyama. 

Oblong, lighter, or darker piceous-brown. Front of head 
scantily punctured, weakly impressed at sides, and with a median 
subelevated longitudinal line, its pubescence long and thin, mouth 
ciliate ; antennze testaceous, with sutures of club straight. Pro- 
thorax a little longer than broad, its base truncate, basal angles 
obtusely rounded, sides rounded, slightly behind, more strongly 
towards apex ; surface somewhat depressed, asperate in front with 
concentric lines of transverse tubercles, becoming weaker behind 
and replaced at base by rugose punctures, except over an indistinct 
median smooth line; sides and apex with long scanty hairs. Scutel- 
lum rather large, rounded, shining. Elytra rather wider than 
prothorax at base, and one half longer, truncate at base, with 
shoulders nearly rectangular, sides parallel to near apex, then 
gradually rounded ; apex not obtuse ; surface cylindrical, obliquely 
and convexly declivous for posterior third, with rows of fine 
punctures, the sutural stria alone with a trace of impression in 
some specimens ; interstices subrugose, with a single row of finer 
setigerous punctures, the setee becoming longer and more con- 
spicuous towards apex. Underside testaceous-brown, nearly 
glabrous and impunctate. Legs reddish testaceous. 


Distinguishable from D. alni, Georg., by its smaller 


size, the weaker punctuation of the base of the thorax and 
elytra, and the absence of the lateral impressions and 


96 Mr. Walter F. H. Blandford on the 


elevated suture of the apical declivity. I can see no 
sexual characters, but one example is more elongate, 
with the thorax more distinctly narrowed behind, and the 
median line subelevated. No specimens measure less 
than 2 mm., except one, which is very small (1°7 mm.), 
with the head finely punctured, and without long 
pubescence. It may be distinct, but I can find no 
characters by which it may be separated. 


The three following species differ somewhat in facies 
from Dryocetes proper, but it is not possible to construct 
a new genus for them without dissection, for which more 
material is required. ‘They can be distinguished by the 
shape of the prothorax, which is net uniformly curved 
from base to apex, but is cylindrical at base and 
declivous anteriorly, so as to form an obtuse median 
elevation. Jts anterior half is markedly scabrous, while 
the hinder part is smooth ; but a uniform sculpture of the 
prothorax does not occur in all the species of Dryocetes. 
Kichhoff described one originally as a Dryocetes, but 
afterwards included it, doubtfully, in the genus Taphro- 
rychus. I cannot, however, see adequate grounds for 
this, and prefer to keep it at present in Dryocetes, with 
which the other two species form a connecting link, 


Dryocates mestus, sp. n. 

Oblongus, cylindricus, subnitidus, pilosus, niger, antennis pedi- 
busque testaceis; prothorace aeque lato quam longo, apice rotun- 
dato, tuberculato, supra transverse subgibboso, antice exasperato, 
postice rugose punctato, linea media obsolete elevata; elytris 
lineato-punctatis, stria suturali subimpressa, interstitiis uniseriatim 
subtiliter punctatis et pilosis, ad apicem fortiter declivibus, 
declivitate deplanata, nitida, ad suturam utrinque impressa. Long. 
2°6 mm. 

A single specimen, Nikko. 

Black, cylindrical ; head with front convex, thinly pubescent, 
punctured, with an indistinct elevated longitudinal line ; antenne 
testaceous, club with sutures curved, pilose, basal joint shining, 
equal in length to succeeding joints, apical joint sensitive, pubes- 
cent. Prothorax with length and breadth equal, base truncate, 
basal angles nearly rectangular, sides feebly rounded to near ‘apex, 
then abruptly and broadly rounded, apical margin tuberculate ; 


? 
surface cylindrical at base, declivous at apex, with an indistinct 


Rhynehophorous Coleoptera of Japan. 97 


transverse elevation in middle and an impression behind it on 
either side of a median longitudinal raised line, somewhat shining 
in middle of base only, anteriorly with transverse rugosities, strong 
near apex, and gradually replaced over base by rugose punctures ; 
pubescence short, rather close. Scutellum triangular, shining. 
Elytra as wide as prothorax and less than twice as long, base trun- 
cate, humeral angles rounded, sides subparallel to near apex, which is 
rather abruptly and obtusely rounded ; surface moderately shining, 
with rows of shallow punctures which appear impressed in certain 
lights only, sutural stria rather more strongly impressed, interstices 
somewhat convex, subrugose, with a single row of fine setigerous 
punctures, the setz longer and conspicuous towards apex, which 
is strongly but not abruptly declivous, more shining and flattened, 
with an impression on either side of the suture, its strix nearly 
obliterated, interstices finely tuberculate. Underside black, nearly 
impunctate, with thin pubescence. Legs reddish testaceous, 
anterior tibiz straight, dilated and obliquely rounded at apex, with 
five or six spines, besides a longer spine at inner angle; middle and 
posterior tibiz: with outer border curved and serrate. 


Dryoccetes dinoderoides, sp. n. 


Oblongus, cylindricus, subnitidus, pilosus, niger, antennis pedi- 
busque testaceis ; prothorace quadrato, apice fere truncato, supra 
elevato subgibboso, antice exasperato, postice rugose punctato, 
linea media laevi; elytris lineato-punctatis, stria suturali quam 
minime impresso, interstitiis uniseriatim subtiliter punctatis et 
pilosis, apice convexe declivi, haud impresso. Long. 2:5 mm. 


One specimen, Ichiuchi. 

Very like the last species and probably the male ; but I 
cannot unite them without further evidence. It differs in 
the prothorax, which is very obtusely convex at apex, so 
as to present an evident angle between the sides and 
anterior margin, as in the next species; the anterior 
border is not crenate, the tubercles are less numerous 
and stronger before the median elevation, and the base 
less closely punctured. The elytral striz do not appear 
impressed in any light, with the exception of the first, the 
apex is noi flattened or impressed along the suture, and 
the pubescence is finer though equally dense. The head, 
antenne, and legs are similar. 

These two preceding species are easily distinguished 
from D. apatoides by the interstices not being costate 
towards the apex of the elytra, 


TRANS. ENT. SOC. LOND. 1894.—ParT I. (MARCH.) G 


Ts Mr. Walter I’, Hl. Blandford on the 


Diyoceles apatoides, Michh, 


Dryocwtes (7) apatoides, Wichh., Seol, Jap., p. 201, 
Taphrorychus (¢) apatoides, Wiehh,, Rat, lom., p. 209, 
No further example has occurred, 


The legs of this inseet are piceous-black, not ferru- 
ginous as described by Wichhoff, They are rather more 
slender than in the two last insects, and, therefore, 
more like those of a Vaphrorychus, but there is no real 
difference in their structure, Lf the two last species are 
soxes of the same, the form of the prothorax in this 
insect is probably sexual, 


Coccornyrns, Wiehh, 

ho species of this genus are closely allied to 
Dryocwles, Wichh., and though differing in the maxillary 
armature and in habits, are not always easily separable, 
The prosternal characters seem to me to be of little 
value, and the basal border of the prothorax, which is 
absent in Dryocwtes, is not always readily  distin- 
vuishable in this genus. They can be differentiated 
without dissection by their shape, which (in the 
Japanese species) is shorter and more ovate, and by the 
tibiw, which are distinetly truncate, and not grooved for 
the tarsi; whereas in Dryocates, the anterior tibiae ab 
loast have the outer margin rounded to the inner angle 
of the apex, without abrupt change of direction, and are 
recessed, 

L add two species, 

Coccolrypes graniceps, Wichh, 

Michh,, Rat, ‘l’om,, p. 314. 

Diyocates graniceps, Lewis, Cat, dap. Col, 

One oxample, Nagasaki; alsoat Liogo (von Schonfeldt), 


[ have not seen the type, which is not in Mr. Lewis’s 
collection, but the specimen corresponds absolutely to 
Wichholl’s deseription, except that the whole surface is 
uniform pitchy-brown, Legs ferraginous, the anterior 
tibia with border simple for basal half, then with 
two outwardly directed spines, apex with two forwardly 
directed spines, besides spine at inner angle ; intermediate 
tibie with three forwardly direeted spines at apex. Pune. 
tuation of abdomen finer than in Dryocawles afinis. 


Rhynchopharous Coleoptera of Japan. 0) 


Voveolrypea perditor, ap. 1, 


Ovalia, subnitidus, forruginous, pilosa; prothoraee hand trans. 
vordo, adapicom subanguatato, margine antico tuboreulato, anterius 
tuberculia concentricia vix transversia, postertus longibudinalibus 
ornato 5; elytria lineatoepunetatia, punetin solin adpromsin, inter. 
alitiin dotin longioribus oreelia ornatia, Long, IS mm, 


ive specimens, in the neighbourhood of Nagasalt, 


Oval, convex, lighter or darker forruginousbrown 5 head 


’ 

punctured, with rather long puboseence ab sides and over mouth 5 
front clovated al sides and imprewed ino middle, with w fine 
modian raived ting, Prothorax searcely broader than long, semi« 
olliptical, narrowed towards apex, and not evidently contracted 
it base, which is truneate, with the posterior angles nearly 
roctangular, sides feobly rounded to middle, thence gradually 
more strongly and #ubsinuste to apex, the margin of which is 
cronate, wilh a series of fine tuberclos; surface rather strongly and 
uniformly convex, moderately shining, with seatterod amall 
weporations forming indistinet concontrie lines in front and longi 
tudinal clovations over the base, Seutellam wmall, punetiforn, 
piccous, Elytra a little wider than bawe of prothorax and one- 
fourth longer, truncate ab base with humeral angles rectangular, 
widow subparallol and straight for basal third, then gradually 
rounded to apex ; above convex, and rounded from base to apox 
with rows of shallow setigerous punctures, their soto short, 
intorvuticos narrow and flat, with a single #erion of loss frequent 
fing aciculations bearing long orect eto, Logs forruginous or 
toulaceous, 


Wasily distinguished from C, granicepa by ita much 
amallor size, and the absence of an apical impression on 
the elytra, Allied to CO. pygmens, Wichh., and, therefore, 
to OU, integer, Wichh,, from Siam, which latter [have 
not seen, Shorter and more ovate than CL pygmeua ; 
the thorax rounder at the sides, and more convex, 
more shining, with far fower asperitios, Mlytra less 
than one-sixth longer than broad (in CO. pygqmceus they 
are wm least one-third longer, and appear comparatively 
eylindrical) ; their strim more distinctly and sparsely 
punctured, and wider apart, the interspaces with fower 
and amallor setiverous tuberclos; stria continued on to 
the apex and not obliterated, as in OC, integer, 


100 Mr. Walter F. H. Blandford on the 


Coccotrypes advend, sp. n. 


Oblongo-ovalis, nitidus, pilosus, piceo-ferrugineus, antennis pedi- 
busque dilutioribus ; prothorace antice contracto, angulis posticis 
obtusis, lateribus rotundatis, apice nonnihil deplanato, mutico, 
supra minus conyvexo, punctis haud frequentibus aciculatis notato ; 
elytris fortius lineato-punctatis, punctis haud setigeris, interstitiis 
uniseriatim setosis. Long, 1*6 mm. 


One example near Nagasaki. 


Oblong oval, ferruginous-brown, with long pubescence. Front 
coarsely punctured, flattened and impressed over mouth. Protho- 
rax contracted in front, as long as broad, basal angles obtusely 
rounded and not applied to humeral angles of elytra, sides and 
apex separately and not strongly rounded ; surface only feebly 
convex, shining, with long hairs, arising from scattered aciculate 
punctures, which are weaker on the disc around an indistinet 
median impunctate line. Scutellum rounded, piceous, shining. 
Elytra wider than base of prothorax, and more than one-third 
longer, humeral angles subrectangular, sides parallel to middle, 
thence obliquely rounded to apex ; surface subcylindrical for basal 
third, then dilated and convexly declivous, with rows of punctures, 
strong and dilated at base, weak and shallow, but not obsolete, 
towards apex, without seriate hairs, interstices with a single row 
of fine aciculations bearing erect sete. Legs ferruginous. 


Readily confounded with C. perditor, but more 
elongate; the prothorax much less convex, with 
the apex and sides not conjointly rounded, and the 
posterior angles obtuse and not applied to the elytra, 
not tuberculate, the punctures being scattered and 
only slightly elevated; punctures. of the elytral strize 
much stronger at base and without sete. The shape of 
the prothorax, which is hardly more convex than that 
of most Dryocetes, and its sculpture will distinguish it 
from other species of Coccotrypes. I have not been able 
to dissect it, but the generic characters, as far as observ- 
able, agree with this genus. 


Xyieporvs, Hichh. 


This genus is very well represented in Japan by 29 
species, exclusive of three males, which I have described 


Rhynchophorous Coleoptera of Japan. 101 


separately, as there is nothing to show to what species 
they belong. This is more than one-fourth of the genus 
as at present known, but only a small proportion of the 
species existing in collections have been described. 
Kichhoff in his “ Ratio Tomicinorum ” groups the species 
by the shape of the prothorax, and thereby obtains a 
satisfactory result. I have found his table of species 
easy to work with, but cannot adopt it for the Japanese 
forms, as.in certain species, e.g., X. validus, praevius, 
seriatus, the form of the prothorax is either ambiguous, 
and could be used to place the species in more than one 
of Kichhoff’s sections, or is of a shape that would dis- 
sociate the insect from its allies. Some previous 
familiarity with the forms met with in the genus is 
necessary in order to locate these doubtful species. 
I have, therefore, in drawing up a table, laid less stress 
on this character, and have aimed simply at differentiating 
the species rather than arranging them in a natural 
sequence. ‘The three species of males referred to are not 
included in it ; their characters will be found on page 119. 


TABLE OF SPECIES. 


1. Elytra confusedly punctured, not inevidentrows. . . . 2 


with distinct rows of puhctures. . . .. . <p 00 
2. Form elongate, prothorax longer than broad, pelliculosus, Fichh. 
short, prothorax not longer than broad . . . . . 38 
3. Elytra abruptly truncate at apex, black. . . . . ... 4 


gradually declivous at apex, ferruginous, 
semi-opacus, Kichh. 
4. Prothorax with two tubercles in middle of apical margin, 

mutilatus. 
without tubercles on apical margin . brevis, Eichh. 
5. Prothorax entirely dull, uniformly and closely asperate. . 6 
with posterior half more or less shining, punctured 7 

6. Interstices of elytra with irregular double rows of ar 
eis? 
with single rows. . . rubricollis, Bichh. 

7. Prothorax subglobose, not longer than broad, or subrect- 

angular with sides and apex separately 
rounded; declivity usually carinate below . 8 

oblong, cylindrical, with apex strongly rounded; 

if not longer than broad, declivity of 
elytra not carinate below . ... . .16 


102 Mr. Walter F. H. Blandford on the 


8. 


10, 


11, 


13: 


14, 


ub 


18. 


ING), 


20. 


7a 


Declivity very oblique, beginning abruptly before middle of elytra, 
opaque and squamous. . . . . . . concisus. 
not abrupt, norsquamous . .. . tic ieren 


. Interstices with irregular double rows of HORGhutes . apicalis. 


with a single row of punctures ..... .10 

Basal half of prothorax strongly punctured, covered with 
short erect hairs . atratus, Hichh. 

not strongly punctured, nor uni- 
formly hairy . . . ee 

Base of prothorax with a patch of hair before scutellueny 
species short; smallie i) oa le 
glabrous, species larger, elongate . . . 13 


. Hairs at apex of elytra arising from interstices only, germanus. 


from both striz and inter- 
stices . . compactus, Eichh, 
First interstice of elytra tuberculate at apex. . . . . . 14 
not tuberculate at apex, 
obliquecauda, Motsch. 
Elytral striz not impressed at apex, tuberculation of 
interstices weak . . . . . previus. 
impressed, tuberculation of interstices strong 15 


. Prothorax narrowed towards apex . . . .  validus, Hichh. 


uniformly rounded at sides, not narrowed, aguilus. 


. Apical border of elytra rounded, declivity not excavate . . 17 


truncate or emarginate, declivity 
sulcate or concave; . . . . . 2d 
Declivity of elytra sharply carinate below .... . . 18 
obtusely margined, not carinate . . . 20 

Prothorax scarcely broader than long, elevated in middle 
ofsurface, distinctly punctured at base, festivus, Eichh. 

Prothorax at least one-half broader than long, elevated 
before middle, base feebly punctured. . . . . 19 

Entirely piceous black, apex of elytra abruptly declivous, 
glabratus, Eichh. 

Prothorax ferruginous, elytra infuscate, gradually declivous 
Ab Apex: |. = cw Gio tee ie meee eon OC On aTs 

Body narrowed behind from front of prothorax, 

attenuatus and sobrinus, Hichh. 
eylindrical \,.) (6085 ts we he ae ole OL eee 
All interstices with traces of tuberculation on apical declivity 22 

First and third interstices tuberculate on declivity, second 
unarmed,’ 4) .Vonvee vs ue aes ea ine os es er 


5) 


Rhynchophorous Coleoptera of Japan. 108 


All interstices tuberculate above declivity, which is im- 
pressed and unarmed (length less than 2 mm.), minutus. 

22. Prothorax as broad as long, elytra with alternate series of 
longer and shorter hairs . . . . . . seriatus. 
longer than broad, hairs of elytra uniform, muticus. 


23, Piceous with thorax sometimes lighter. . . . . adumbratus. 
Ferruginous or testaceous . . . alee ce « 24 

24, Elytra rather strongly striato- famataie Be oN fade Eichh. 
finely punctured in rows. . . . .  vicarius, Kichh, 


25. Declivity with a narrow suleate impression along suture, 
each lateral margin with four or five small 


tubercles’... fe: - . « schaufusst. 

widely impressed, eis eed margin with two 
strong spines . . . See . 26 

26. Ferruginous brown, apical deprension mearl Serra feably 
emarginate below .. . tite Pele) AfOnsUs: 

Black, apical depression very oblique, us ongly emarginate 
GLOW fy a! var al oe. Mt Ce emote ct eee al, eh, NEES Cs 


Xyleborus mutilatus, sp. n. 


Fem. Curta, subnitida, fusco-pilosa, nigra, antennis pedibusque 
ferrugineis ; prothorace magno globoso, lateribus leniter, apice 
fortiter rotundato et tuberculis duobus ornato, supra in medio 
transverse elevato et postice utrinque impresso, anterius exasperato, 
posterius dense punctato, supra scutellum hirto ; elytris prothorace 
brevioribus, a basi fere oblique et abrupte declivibus, supra irregu- 
lariter punctatis, declivitate striata, interstitiis granulatis, lateribus 
et apice infra marginatis. Long. 3°5 mm. 


One example, without locality. 


Black, with the base of the elytra alone shining, covered with 
fine erect fuscous hairs. Head large, prominent, finely reticulate, 
front convex, punctured, and hairy; mouth ciliate with yellow 
pubescence ; eyes small, flat, finely emarginate ; antenne ferruginous, 
club round, basal joint large, shining, reaching nearly to apex of 
club. Prothorax a little longer than broad, its base bisinuate, 
produced behind, basal angles nearly rectangular, sides straight 
and subparallel behind, becoming more rounded in front with apex 
broadly convex, apical margin bisinuate, slightly produced in 
middle and armed with two prominent tubercles ; surface very 
convex, cylindrical at base, with an obtuse transverse élevation in 
middle, in front of which it is declivous to apex, anteriorly 
separate and pilose, posteriorly densely punctured except over 


104 Mr. Walter F; H. Blandford on the 


two shining lateral impressions, with thin pubescence at sides and 
a dense patch before scutellum, which is large, shining, and 
rounded, Elytra as wide but not as long as prothorax, basal 
borders feebly convex, shoulders narrowly elevated; at first 
cylindrical, then obliquely declivous from basal fourth to apex, 
shining and irregularly punctured, the punctures rugose and con- 
fluent along basal margin ; apical declivity rounded oval with a 
sharp raised margin to sides and lower border, its surface subconvex, 
pilose, with impunctate striz, interstices closely granulate ; lateral 
border of elytra declivous, parallel throughout to margin of apical 
declivity. Underside black, thinly pubescent, anterior coxe 
separated by a narrow prosternal process. Legs ferruginous, 
tibie strongly dilated and obsoletely spined; tarsi short, their first 
three joints compressed, and pilose beneath. 


The most extreme type of the truncate Xylebori I have 
seen, and a remarkable instance of the diversity of form 
which the genus presents. 


AXyleborus brevis, Eichh. 


Hichh., Rat. Tom., p. 319. 
Four examples, Nikko; originally taken at Hagi by 
Hiller. Identified by the description. 


Xyleborus lewisi, sp. n. 


Fem. Oblonga, cylindrico-convexa, pilosa, rubra elytris infus- 
catis ; prothorace subgloboso, apice tamen deplanato, supra sub- 
aequaliter convexo, opaco, antice fortiter postice paullo subtilius 
asperato ; elytris subnitidis, striato-punctatis, interstitiis subtiliter 
biseriatim punctatis in declivitate tuberculatis, apice oblique declivi 
ad suturam impresso. Long. 4°5 mm. 


Nikko, Hakone, Miyanoshita, nine specimens. 


Oblong, convex, and cylindrical, pilose with long hairs, red with 
elytra infuscate. Head coarsely and rugosely punctured, with an 
indistinct elevated median line, thinly hairy and ciliate over 
mouth. Prothorax transverse, nearly globose, but with sides and 
apex separately rounded and antero-lateral angles more strongly 
rounded, base subsinuate with obtusely rounded angles ; above gib- 
bous and convex, but without median elevation, dull and entirely 
scabrous, the asperities a little weaker posteriorly. Scutellum 
rounded, shining, infuscate, anteriorly impressed. Elytra as wide as 
base of prothorax and more than one-half longer, truncate at base, 


Rhynchophorous Coleoptera of Japan. 105 


with humeral angles rounded-rectangular, sides straight but sub- 
divergent to apex, which is rather abruptly and broadly rounded 
and inconspicuously carinate below; surface very convex with 
scarcely impressed irregular stria of large shallow punctures and 
long coarse pubescence, especially at apex, interstices finely 
punctured in irregular double rows, declivity very convex, with 
first interstice widened, impressed and finely tuberculate, the second 
very strongly, the rest less strongly tuberculate. 


‘The largest Japanese species of the genus; very like 
X. rubricollis, Hichh., in colour and in the completely 
asperate prothorax, but very much larger, and with the 
elytral interstices punctured in double instead of single 
rows. ‘the prothorax is flattened in front, a feature not 
found in other species with a short subglobose prothorax. 


Xyleborus vubricollis, Kichh. 
Hichh., Scol. Jap., p. 202; Rat. Tom., p. 330. 
No further examples taken. 


Xyleborus apicalis, sp. n. 

Fem. Oblonga, subnitida, longius pilosa, piceo-nigra, antennis 
pedibusque ferrugineo-testaceis ; prothorace orbiculato, transverso, 
margine antico tuberculato, disco transverse elevato, postice sub- 
tiliter punctato, margine basali hirto; elytris prothorace, sesqui 
longioribus, lineato-punctatis, interstitiis irregulariter punctatis, 
setosis, ad apicem convexe declivibus, praeter suturam impressis, 
interstitio 20 in summa declivitate obtuse spinato, 30 elevato, 
tuberculato. Long. 3 mm. 


One specimen, without locality. 


Oblong, pitchy black with antenne and legs reddish testaceous. 
Head finely reticulate, front with scattered strong punctures, thinly 
hairy, mouth ciliate. Prothorax transverse, truncate at base, basal 
angles obtusely rounded, sides and apex rounded, the latter more 
strongly and tuberculate ; surface with a median transverse 
elevation, anteriorly exasperate, posteriorly with fine scattered 
aciculate punctures, with long pubescence scattered at sides and 
apex, denser along basal margin. Scutellum triangular, piceous. 
Elytra rather narrower at base than greatest width of prothorax, 
and one-half longer, humeral angles obtuse, sides rounded from 
middle to apex, which is somewhat obtuse ; surface cylindrical at 


base, with rows of punctures, very fine at base becoming strong 


106 Mr. Walter F. H. Blandford on the 


about middle, interstices with an irregular series, usually double, 
of finer setigerous punctures, apical declivity convex, strong but 
not abrupt, not acutely carinate below, impressed on either side of 
suture and margined by the elevated and finely tuberculate 3rd 
interstice, 2nd interstice with a short obtuse spine at upper angle 
of declivity, hairs long, especially towards apex. 


Not unlike X. atratus, Hichh., but distinguished by 
the prothorax being finely punctured behind, the elytral 
interstices being biseriately punctured, and the tubercle 
on the third. Separable by the structure of the pro- 
thorax from Dryocetes pilosus and affinis. 


Xyleborus atratus, Kichh. 
Hichh., Scol. Jap., p. 201; Rat. Tom., p. 324. 
A few more specimens taken at Kiga and Nagasaki. 


AXyleborus germanus, sp. n. 

Fem. Breviter cylindrica, nitida, picea, antennis pedibusque 
testaceis ; prothorace subgloboso, convexo, indistincte transverse 
elevato, anterius exasperato, posterius subtiliter punctulato, margine 
basali medio piloso ; elytris prothorace sesqui fere longioribus, ad 
apicem oblique declivibus et infra carinatis, subtiliter striato- 
punctatis, interstitiis rarius uniseriatim punctatis in declivitate 
seriatim pilosis. Long. 2—2'3 mm. 


Apparently common; sixteen specimens altogether 
have been taken at various places, two or three at most 
from each. 


Oyayama, Nikko, Subashiri, Kiga, Miyanoshita. 


Short, cylindrical, piceous, shining. Head finely reticulate, with 
front convex, scantily punctured, mouth ciliate ; eyes flat, oblong, 
deeply emarginate. Prothorax as long as broad, base truncate, 
basal angles rounded, sides scarcely rounded behind, broadly in 
front; surface convex with an obtuse median transverse elevation, 
anteriorly with concentric rows of exasperations, strong over apex, 
posteriorly shining with scattered fine punctures, pubescence short 
and sparse, except at apex and in middle of basal margin. 
Scutellum large, subtriangular, shining. Elytra as wide as thorax, 
and nearly twice as long ; humeral angles obtusely rounded, sides 
subparallel to apex, which is abruptly and broadly rounded and 
carinate below; surface convex, somewhat rounded from base to 
middle, then obliquely but not abruptly declivous, piceous-brown, 


Rhynchophorous Coleoptera of Japan. 107 


with fine rows of punctures which appear impressed in certain 
lights, interstices rather wide, flattish, each with a row of very fine 
punctures, and behind middle with erect sete arising from slight 
tubercles. Underside brown, punctured, pubescent. Legs tes- 
taceous. 


Closely allied to X. compactus, Hichh., but twice as 
large, with distinct though feebly impressed elytral striae 
and regular rows of hairs on the apical declivity, which 
are confined to the interstices. 


Xyleborus compactus, Hichh. 
Eichh., Scol. Jap., p. 201 ; Rat. Tom., p. 328. 


No further examples taken. The hairs are shorter 
and more depressed at the apex of the elytra than in the 
last species, and arise from the striz as weil as from the 
interstices. 

Xyleborus semi-opacus, Hichh. 

Hichh., Rat. Tom., p. 380. 

Five specimens, Konose, Kioto, Chiuzenji; also at 
Hong-Kong in China (J. J. Walker). 

The type is not in Mr. Lewis’s collection, but the 
Species is unmistakable. In certain lights rows of 
punctures can be seen on the elytra, but they are not at 


all clear, and cannot be confounded with the distinct 
lines occurring in most of the genus. 


Xyleborus concisus, sp. Nn. 

Oblongus, nitidus, ferrugineus, prothorace et elytris ad apices 
infuscatis, breviter ac parce pilosus ; prothorace transverso lateribus 
leniter, apice fortius rotundato et crenato, supra in medio obtuse 
transverse elevato, postice discrete punctato ; elytris latitudine a 
basi crescentibus, lateribus rectis, apice rotundato, striato-punctatis, 
interstitiis subtilius uniseriatim punctatis, declivitate valde obliqua, 
ante medium incipiente, subconvexa, squamis obtecta, striata. 
Long. 2°4 mm. 


One example, without locality. 
Oblong, ferruginous, with apex of prothorax and elytra slightly 
infuscate. Head piceous, finely reticulate, front subconvex, sparsely 


and strongly punctured towards sides and scantily pubescent, witha 
transverse impression over mouth, above which is a somewhat 


108 Mr. Walter F. H. Blandford on the 


tuberculate elevation. Antenne testaceous. Prothorax transverse, 
not narrowed towards apex; base truncate, posterior angles rounded, 
sides and apex rounded, the former very feebly, the latter more 
strongly, with margin crenate ; surface with an obtuse transverse 
elevation in middle, rather finely asperate anteriorly, posteriorly 
with distinct scattered punctuation. Scutellum triangular, shining. 
Elytra rather less than twice as long as prothorax, and as wide at 
base, which is truncate, with the humeral angles nearly rectangular, 
sides straight but divergent to apex, which is circularly rounded 
and narrowly emarginate at suture ; surface at base cylindrical and 
longitudinally convex, striato-punctate, interstices flat, with a single 
row of very fine punctures, apical declivity sharply marked, very 
oblique, beginning before middle of elytra, dull and clothed with 
cinereous scales, subconvex, impressed along inferior margin which 
is acute, striate, the interstices flat. Underside testaceous, thinly 
pubescent. Legs testaceous, the anterior tibie obliquely truncate 
and uncinate at apex, middle and posterior tibiz rounded. 


This species corresponds closely to Hichhoff’s de- 
scription of X. sordicauda, Motsch., from which it differs 
in no important respect except in size. X. sordicauda is 
said to be twice as large as X. semi-opacus, Hichh., 
whereas this specimen is rather smaller. 

Minor points of difference from the description of 
X. sordicauda are found in the prothorax, which is not 
contracted anteriorly, and in the elytra being dilated 
posteriorly, with the apical margin more strongly rounded. 
I conclude that it is distinct, and that we have an instance 
of what is common in the genus, a well-marked form 
represented by more than one species. It may easily be 
separated from X. semi-opacus by the punctured striz 
of the elytra. 


Xyleborus validus, Hichh. 
Kichh., Scol. Jap., p. 202; Rat. Tom., p. 358. 


Taken in greater numbers than any other species; 
extending from Sapporo and Junsai in Yezo, to 
Nagasaki and Oyayama in Kiushiu, also at Nikko, 
Miyanoshita, etc. As there is a closely allied and hardly 
distinguishable species from Ceylon, it represents an 
Oriental rather than a Palearctic; type. It is found 
in firs. 


Rhynchophorous Coleoptera of Japan. 109 


Xyleborus obliquecauda, Motsch. 


Phleotrogus obliquecauda, Motsch., Bull. Mosc., 1863, 
is, p. Obs. 

Xyleborus carinipennis, Hichh., Berl. Zeit. 1868, p. 152. 

AX. obliquecauda, Eichh., Rat. Tom., p. 351. 


One specimen, near Yokohama, Oct. 7th, 1881. It 
appears to me identical with examples taken by Mr. 
Lewis in Ceylon, and corresponding to Hichhoff’s descrip- 
tion of X. obliquecauda. The prothorax is a trifle more 
depressed, and the elytra] strie more regular and not 
at all impressed ; but there is no more variation than might 
be expected between specimens from distant localities. 


Xyleborus aquilus, sp. n. 

Fem. Oblonga elongata, nitida, breviter pilosa, ferrugineo- 
picea, antennis pedibusque ferrugineis ; prothorace subqvadrato, 
lateribus et apice singulatim leniter rotundatis, in medio transverse 
gibboso, postice subtiliter disperse punctato; elytris ad apicem 
oblique declivibus, infra subtiliter carinatis, leniter  striato- 
punctatis, striis in declivitate impressis, interstitiis planis uni- 
seriatim setosis, in declivitate convexis, tuberculatis. Long. 3:°2— 
oo mm, 

Six examples taken at Oyayama and Hitoyoshi in 
Kiushiu, one near Kashiwagi. 

Somewhat similar to Y. eurygraphus, Ratz, but narrower and less 
convex, with the prothorax shorter and less evidently quadrate, 
the elytral striz much finer, and the interstices convex at apex, 
more finely and evenly tuberculate. Ferruginous- brown or 
piceous ; head dull, front punctured, thinly pubescent, with an 
obsolete median raised line. Prothorax a little longer than broad, 
truncate at base, with posterior angles obtusely rounded, sides and 
apex separately slightly rounded, antero-lateral angles more 
strongly ; surface gibbous in middle, the elevation transverse but 
short, its anterior half rather finely asperate, the posterior half 
shining, scantily and finely punctured. Scutellum small, triangular, 
shining. Elytra slightly narrower than prothorax at its widest 
part and about one-half longer, humeral angles rounded rect- 
angular, sides parallel to apex then subcircularly rounded and 
margined ; surface slightly convex from base to declivity, which 
is oblique, with fine punctured striz, little or not at all impressed 
before apex, where they are dilated, with a slight outward curve, 
interstices with a single row of setz, flat anteriorly, convex at 


110 Mr. Walter F. H. Blandford on the 


apex and finely tuberculate, the first having traces of tubercles 
from the middle. Underside ferruginous, thinly hairy. 


There are two forms of this species, one slightly 
shorter with the elytral striae not impressed at all on the 
declivity, which is impressed transversely, somewhat 
dull, and sub-acuminate at apex; the tubercles are finer 
than in the other form, in which the striz are impressed, 
and the apex is shining and subconvex. One specimen, 
however, appears to be intermediate, and therefore I 
do not separate them, though at first sight they appear 
specifically distinct. 


Xyleborus praevius, sp. n. 

Fem. Elongata, subcylindrica, nitida, parce pilosa, picea, 
antennis pedibusque testaceis ; prothorace paullo longiore quam 
latiore, lateribus leniter, apice fortiter rotundato et crenato, disco 
umbonato, postice sparsim distincte punctato ; elytris prothorace 
plus quam sesqui longioribus, ad apicem oblique declivibus et 
infra carinatis, declivitate utrinque subimpressa, supra subtiliter 
lineato-punctatis, interstitiis rarius uniseriatim punctatis, postice 
obsolete tuberculatis. Long. 3 mm. 


One example, without locality. 


Similar to X. aquilus, but smaller, more slender and 
with the interstices flat on the apical declivity. 


Head dull with front flattened, coarsely punctured at sides, 
with a smooth elevated area in middle, pubescence short except 
over mouth, eyes wide, not deeply emarginate, antennz testaceous. 
Prothorax a little longer than broad, its base truncate, basal 
angles obtuse, sides scarcely rounded behind, becoming gradually 
more strongly rounded to apex, which is crenate, surface elevated 
in middle, the elevation not evidently transverse, anterior half 
with rather scattered transverse asperities, becoming very fine 
over median elevation, posterior half shining, with irregular 
scattered but distinct punctures. Scutellum small, obtuse trian- 
gular, shining. Elytra as wide as base of thorax, and more than 
half as long again, humeral angles obtusely rounded, sides not 
quite parallel, slightly dilated in middle, subcireularly rounded 
and carinate at apex, surface slightly convex from base to apex, 
thence somewhat obliquely declivous, with rows of shallow 
punctures, which are not impressed, interspaces between punctures 
transversely rugose, interstices with a single row of sete, arising 
from fine tubercles on the whole of the first two, and the apice 


Rhynchophorous Coleoptera of Japan. 111 


of the remaining interstices. Apical declivity with a shallow 
impression on either side of the suture. Underside ferruginous, 
with abdomen darker, very scantily punctured and pubescent. 
Legs reddish testaceous. 


Xyleborus seriatus, sp. n. 

Fem. Oblonga, cylindrica, subnitida, pilosa, fusco-picea elytris 
dilutioribus, antennis pedibusque rufescentibus ; prothorace longi- 
tudine et latitudine aequali, apice fortiter, lateribus paullo rotundatis, 
supra transverse subelevato, postice subtiliter reticulato et fortiter 
punctato, linea media laevi; elytris ad apicem oblique declivibus, 
declivitate subimpressa subtus haud acute carinata, setis alterne 
longioribus et brevioribus  seriatis, lineato-pr-nctatis, interstitiis 
uniseriatim punctatis versus apicem tuberculatis. Long. 2:5 mm. 


Two examples taken at Nikko and Miyanoshita. 

Oblong, cylindrical, dull piceous with elytra lighter. Head 
finely reticulate, front nearly flat, scantily punctured and pube- 
scent, with an indistinct median longitudinal elevation, mouth 
ciliate, eyes deeply emarginate, antennz testaceous. Prothorax 
as broad as long, truncate at base with posterior angles obtuse, 
sides slightly rounded, in front. strongly rounded to apex ; above 
with an- obtuse transverse elevation in middle, anterior half 
asperate, posterior half finely reticulate and strongly punctured, 
the punctures becoming asperate at sides, with a smooth median 
line, pubescence short and scanty. Scutellum small, rounded, 
piceous. Elytra as wide as prothorax, and more than half as 
long again, truncate at base with humeral angles rounded rect- 
angular, sides parallel to middle, then obliquely and gradually 
rounded to apex ; surface cylindrical to middle, obliquely declivous 
behind, the declivity impressed weakly at sides of suture, with 
inferior margin inflexed but not carinate, with scarcely impressed 
rows of strong close setigerous punctures, the sete very short and 
fine, interstices with a single row of punctures, a little finer and 
less numerous, bearing longer setze, and very finely tuberculate 
for apical half. Legs ferruginous. 


This species resembles X. dryographus, Ratz., but 
the prothorax is shorter and more strongly punctured 
behind. The elytral punctuation is much stronger, and 
the rows of shorter and longer setz are quite distinctive. 
It is allied to Hichhoff’s section + + +, in which the 
prothorax is defined as being cylindrical and oblong; 
but in this species the prothorax is so short, that it will 
not serve to separate it from the insects of section +. 


ET Mr. Walter I’. H. Blandford on the 


Xyleborus pelliculosus, Kichh, 
Hichh., Rat. Tom., p. 336. 


Two examples, Kiga. 


I have not seen the type of this insect, and am less 
certain, than with other unseen species, that I have here 
the true X. pelliculosus. But the description fits it, and 
my doubt is rather owing to the fact that there exist 
other at present undescribed species of similar facies 
in the Oriental region. It is quite easily distinguished by 
the rather dense and almost downy pubescence, and the 
confusedly and finely punctured elytra, which bear only 
the feeblest traces of striz. One example is black, the 
other sordid testaceous. ‘The shape of the prothorax 
would lead me to put it in Hichhoff’s group + + + rather 
than +, but it is a distinct form, and not very closely 
related to any other described species. I have placed 
it in the neighbourhood of X. muticus, which is of 
similar build, without evident elevation of the prothorax, 
but that species has the elytra plainly lineato-punctate. 


Xyleborus muticus, sp. n. 

Fem. Elongata, cylindrica, subnitida, pilosa, rufo-picea, 
antennis pedibusque ferrugineis; prothorace oblongo, antice 
fortiter rotundato, supra parum gibboso, post medium punctis 
subaciculatis subtilibus notato; elytris lineato-punctatis, inter- 
stitiis subtilissime uniseriatim punctatis et pilosis, apice fortiter 
declivi, ad suturam subimpresso, striis impressis, interstitiis vix 
perspicue tuberculatis. Long. 3 mm. 

Two examples, near Kashiwagi. 


Ferruginous-pitchy, with rather long soft pubescence. Head 
ferruginous, dull, front convex, punctured at sides with a median 
impunctate slightly raised line, pubescence scanty and short except 
over mouth; antenne ferruginous. Prothorax oblong, base 
truncate, basal angles obtusely rounded, sides straight, slightly 
divergent from base towards apex, which is strongly rounded ; 
surface with only the slightest trace of a transverse elevation, 
thinly pubescent especially at sides and apex, its anterior third with 
rather close transverse asperities which become weaker, but are 
continued back to posterior third, which is very finely reticulate, 
dull and subaciculately punctuate, the punctures rather close at 
the sides. Seutellum rounded, piceous, shining. Elytra slightly 


Rhynchophorous Coleoptera of Japan. 113 


wider than prothorax at base and about half as long again, 
humeral angles subrectangular, shoulders narrowly callose, sides 
parallel to behind middle, thence rounded to apex, which is not 
earinate below ; surface cylindrical with slightly impressed rows of 
oblong punctures, interstices narrow with a single row of very fine 
setigerous punctures, about as numerous as those of striz, the 
hairs stronger towards apex, which is strongly but obliquely 
declivous, slightly flattened, and impressed along the suture with 
the punctures of the striz larger, rounded, and shallow ; interstices 
flat with microscopic traces of tuberculation, rather more evident 
on the third interstice. Underside and legs ferruginous, the former 
shining, punctured, and pubescent. 


In the feeble development of any apical armature to 
the elytra, this species differs from any described by 
Kichhoff. Other examples of the same type occur in the 
Oriental and Polynesian regions, e.g., X. obliqwus, Sharp, 
from Hawai. 


Xyleborus festivus, Eichh, 
Kichh., Scol. Jap., p. 2/2; Rat. Tom., p. 366. 
No further examples have been taken. 


Xyleborus glabratus, Hichh. 
Hichh., Rat. Tom., p. 381. 
Yokohama, Higo; a few specimens. 


Xyleborus bicolor, sp. n. 

Fem. Elongata, cylindrica, nitida, subglabra, capite et pro- 
thorace ferrugineo-testaceis, elytris fusco-piceis, antennis pedi- 
busque testaceis : prothorace oblongo, antice fortiter rotundato, in 
medio subgibboso, postice sparse punctato, linea media obsolete 
elevata laevi; elytris ad apicem oblique declivibus et infra 
carinatis, subtiliter lineato-punctatis, interstitiis lo et 30 ad apicem 
tuberculatis. Long. vix 2 mm. 


Four examples, Nagasaki, Feb. 21st, 1881; one at 
Inasa on Kashinoki (Ilex sp.). 

Reddish testaceous with the elytra piceous brown. Head finely 
reticulate, front flattened, impressed on either side with a group 
of punctures, pubescence very scanty, mouth ciliate. Eyes oval 
with a deep angulate emargination. Antenne testaceous. Pro- 
thorax oblong, base truncate, basal angles obtusely rounded, sides 


TRANS. ENT. SOC. LOND. 1894.—ParTI. (MARCH.) H 


114 Mr. Walter F. H. Blandford on the 


nearly straight, slightly divergent from base to near apex, which 
is strongly rounded, surface somewhat depressed, declivous for 
anterior third, its junction with the posterior cylindrical part 
marked by a fine elevation at the apex of an indistinct raised line 
reaching nearly to base ; finely asperate in concentric lines before 
elevation, with sparse short hairs, basal half shining, feebly 
punctured, with a row of more distinct punctures on either side 
of central line. Scutellum rounded, piceous, shining. Elytra as 
wide as base of prothorax and half as long again, basal angles 
rounded rectangular, humeral callosities distinct, sides subparallel, 
apex circularly rounded and acutely margined below by the 7th 
interstice ; surface subdepressed, very obliquely declivous at apex, 
punctured in rows, interstices flat witha single row of very fine 
sparse punctures ; declivity with 1st and 3rd interstices elevated, 
finely tuberculate and setose. Underside and legs testaceous, the 
former punctured at sides and at apex of abdomen. 


Readily distinguished by its small size, narrow form, 
fine sculpture and colour. 


Xyleborus attenuatus, sp. n. 


Fem. Elongata cylindrica, a prothorace medio posterius sub- 
angustata, subnitida, pilis erectis brevibus pubescens, picea ; 
prothorace oblongo, in medio transverse elevato, postice sublaevi, 
punctis minutis notato ; elytris ad amussim punctato-striatis, iuter- 
stitiis uniseriatim punctatis, apice subopaco, oblique rotundato, 
sutura elevata, interstitiis lo, 30, et 40 tuberculatis, 20 inermi., 
Long. 2.6. mm. 


One example, Nikko. 


Very closely allied to X. saxeseni, Ratz., like the 
following species, but in my opinion distinct from either. 
From X. saweseni it differs as foilows: 


The body is more elongate and evidently narrowed, especially 
towards the apex of the elytra, and is furnished throughout, 
including the posterior half of the prothorax, with short upstanding 
hairs ; thoracic tubercle more elevated, the surface behind less 
shining, more evidently and closely punctured ; elytral punctures 
stronger and rather closer, 2nd interstice more deeply impressed at 
apex, and the tuberculation stronger; third stria also somewhat 
impressed. From. sobrinus it can be distinguished by its larger 
size, the distinct and wegular rows of punctures on the elytra, 


~ 


Rhynchophorous Coleoptera of Japan. 115 


and the wider space between the two inner rows of tubercles on the 
apex, which is distinctly punctured in the line of the two first 
strie. 


Xyleborus sobrinus, Kichh,. 


Hichh., Scol. Jap., p. 202; Rat. Tom., p. 363. 
Four specimens added from Chiuzenji. 


Somewhat variable in the sculpture of the elytra and 
the development of the elytral tubercles, which are usually 
much stronger than in any specimens of X. saveseni. 
The tubercles of the Ist interstice begin about the 
middle of the elytra, whereas in the single example of 
X. attenwatus they are confined to the declivity. 

A specimen from Hiogo in Colonel von Schoénfeldt’s 
collection is not separable from saweseni by any cha- 
racters. As it is quite possible that there may be two 
or three species allied to saweseni in Japan, I allow this 
and the preceding species to stand as a help for further 
investigations. 


Xyleborus adumbratus, sp. n. 

Fem. Oblonga, cylindrica, nitida, pilis erectis adspersa, nigra vel 
picea, prothorace nonnunquam rufescente, antennis pedibusque 
testaceis, his infuscatis; prothorace oblongiusculo, apice fortiter 
rotundato, in medio transverse elevato, postice sparse subtiliter 
punctulato ; elytris punctato-striatis, interstiis subrugosis, uniseri- 
atim subtiliter punctatis et pilosis, duobus primis a basi tuberculatis, 
apice declivi deplanato, sutura ad apicem solum callose elevata, 
interstitiis lo et 30 tuberculis acutis ornatis. Long. 3 mm, 


Hight examples taken at Nagasaki, Hitoyoshi, Oyama, 
and Subashiri. 


Closely allied to X. pfeili, Ratz., of which it is perhaps 
a mere geographical variety. 


Tt varies in colour from black to piceous brown, with the thorax 
sometimes lighier than the elytra. Head as in X. pfeili, with 
the front strongly punctured, with a more or less evident longi- 
tudinal carina. Prothorax as in X. pfeili, but with the hairs 
longer, and the central elevation more prominent and trans- 
verse ; its basal half very shining, diffusely punctured except 
behind elevation, where the punctures are closer. Elytra one half 
longer than prothorax, a little shorter and more conyex than in 


116 Mr. Walter F. H. Blandford on the 


X, pfeili; sides parallel at base, slightly narrowed and rounded 
behind middle, apex feebly produced in middle, sinuate on 
either side ; their sculpture similar to that of X. pfeili, but with 
the punctuation a little stronger and the points of insertion of 
the hairs on the two first interstices more or less evidently elevated 
and tuberculate. Apical declivity with suture not elevated except 
at apex, where it is callose, the tubercles more prominent, and 
surface less wrinkled and more clearly punctured. Underside 
piceous, or testaceous, with abdomen darker. 


Xyleborus badius, Hichh. 


Hichh., Berl. Ent. Zeit., 1868, p. 280; Rat. Tom., 
p. 379. 


Hiogo, one example (von Schdnfeldt). 


I can see no difference whatever between this insect 
and a typical example from Madagascar. 


NXyleborus vicarius, Kichh. 
Hichh., Scol. Jap., p. 203; Rat. Tom., p. 376. 
No further examples taken. 


The apical declivity of the typical specimens is 
decidedly dull, as in X. affinis, Hichh., and not shining as 
described by Hichhoff. 


Nyleborus minutus, sp. n. 

Fem. Oblonga, nitida, pilis brevibus erectis ornata, ferrugineo- 
testacea ; prothorace latitudine paullo longiore, lateribus leniter, 
apice fortiter rotundato, supra in medio elevato-nodoso, postice 
discrete subtiliter punctulato ; elytris lineato-punctatis, interstitiis 
vix perspicue uniseriatim punctatis, ad apicem oblique declivibus, 
declivitate deplanata, subimpressa, interstitiis omnibus subtilissime 
post medium tuberculatis, versus apicem muticis. Long. 1:5— 
1-7 mm. 


Three examples, Inasa. 


At present the smallest species in the genus, shorter 
though more robust than X. bicolor. 


Oblong, subcylindrical, testaceous-brown, a little darker at the 
extremities. Head finely reticulate, testaceous with epistoma 
darker, front subconvex, punctured at sides and shortly pubescent, 
with a median longitudinal elevation towards vertex, carinate in 
one example, mouth shortly ciliate, eyes oval, flat, emarginate, 


Rhynchophorous Coleoptera of Japan. 17 


antennz testaceous. Prothorax a little longer than broad, slightly 
rounded at base, basal angles obtuse, sides gently rounded to near 
apex, which is strongly rounded; surface with a slight nodose 
elevation in middle, scantily pubescent at sides and apex, its 
anterior half with concentric asperations, the posterior half shining, 
finely punctured with a groundwork of very fine parallel scratches. 
Scutellum rounded, shining, infuscate. Elytra as wide as base of 
prothorax and rather less than half as long again, base truncate, 
humeral angles subrectangular rounded, sides very feebly rounded, 
nearly parallel, apex strongly rounded, with the lateral margin 
inflexed below for a very short distance, but scarcely carinate ; 
surface shining, convex from base to posterior third, thence 
obliquely declivous, the declivity flattened and more or less 
impressed, with rows of punctures which appear large, rounded, and 
dark when strongly illuminated, interstices somewhat irregular, 
with single rows of very fine sparse punctuation and short erect 
hairs, behind middle with fine tuberculation, obliterated on the 
impressed apical area on which the strie are slightly impressed, and 
curved inwards at the tip. Underside and legs testaceous. 


Xyleborus schaufussi, sp. n. 

Elongatus, cylindricus, nitidus, ferrugineo-brunneus vel piceus, 
pilosus ; prothorace oblongo, apice rotundato supra vix gibboso, 
postice subtiliter punctulato; elytra subtiliter lineato-punctatis, 
striis haud impressis, interstitiis discretius uniseriatim punctatis, 
apice retuso, laevi, ad suturam profunde impresso, lateraliter 
utrinque elevato, tuberculis 4 aut 5 ornato. 

Mas. Minor, angustior. 

Long. Mas.—2‘7 mm.; FemM.—3 mm. 


Several examples taken at Lake Junsai and Nikko. 


Varying in colour from ferruginous-brown to piceous. Head 
dull, front convex strongly punctured at sides, smooth in middle 
with a slight longitudinal elevation over mouth ; pubescence grey, 
rather long, mouth ciliate with yellow hairs, eyes not deeply 
emarginate, antenne testaceous-brown. Prothorax oblong, nar- 
rower in male than in female, base truncate, basal angles rounded, 
sides parallel, apex strongly rounded; surface with only a trace 
of acentral elevation, with very short scanty pubescence, longer 
at sides and apex, its anterior third with rather fine imbricate 
asperities, basal part shining, finely punctured with an indistinct 
smooth central line. Scutellum very small, triangular, depressed, 
Elytra barely as wide as prothorax and rather more than half as 


118 Mr. Walter F. H. Blandford on the 


long again, separately, but slightly rounded at base, humeral 
angles rectangular, shoulders scarcely prominent, sides parallel 
to behind middle, thence gently rounded, apices nearly truncate, 
or separately rounded with slight median emargination ; surface 
shining, finely punctured in rows, interstices with a single row of 
punctures, as strong but not so frequent as those of stria, and 
with a series of fine hairs which are longer posteriorly ; behind 
convex, strongly and nearly vertically declivous, deeply impressed 
along suture for posterior third of elytra, the depression smooth, 
shining, with elevated callose sides, bearing four or five tubercles 
on the third, and finer ones on the succeeding interstices, first and 
second interstices with two or three fine tubercles before apical 
impression. Underside testaceous-brown, punctured at sides and 
very scantily pubescent. Legs testaceous with knees infuscate. 
Male similar in sculpture to female, but smaller and more slender. 


The impression of the elytral apex is much deeper 
than in X. confusus, Hichh., and resembles that of 
certain Pityophthort. 


Xyleborus defensus, sp. n. 


Fem. Elongata, cylindrica, ferrugineo-brunnea, breviter pilosa; 
prothorace oblongo, apice rotundato, supra leniter gibboso, postice 
distincte sparsim punctato; elytris lineato-punctatis, interstitiis 
uniseriatim punctatis, apice subcirculatim excavato retuso, infra 
emarginato, in retusionis margine dentibus duobus acutis utrinque 
armato. Long. 3 mm. 


One specimen, Sapporo. 


Similar to the last species, but with the following 
distinctive features : 


Body stouter and more convex ; prothorax slightly narrowed 
towards the apex and not regularly rounded, median elevation 
evident, though small, punctuation of the basal half stronger 
and rather irregular. Punctures of the elytra stronger and less 
numerous, interstices somewhat rugose ; apex widely impressed, 
with the elevated callose margin subcircular, but not reaching 
the suture below, so that the lower border has a shallow 
emargination, impressed surface shining and impunctate; there 
are, on each side, two minute tubercles before it close to the 
suture, and two spines just within the lateral margin, one near 
the upper extremity, the other about the middle. Antenne, legs 
and underside ferruginous. 


Rhynchophorous Coleoptera of Japan. 119 


I have not seen X. fallax, Eichh., to which this 
species is allied in the structure of the elytra; but the 
former differs according to the description in several 
points. Itis more elongate, with the posterior half of 
the prothorax very finely punctured, the apical emargi- 
nation of the elytra deep, and the impressed surface 
rugosely punctured. 


Xyleborus exesus, sp. 0. 

Fem. Elongata, cylindrica, nitida, pares pubescens, piceo- 
nigra, antennis pedibusque ferrugineis ; prothorace oblongo, parum 
gibboso, postice distincte ac discrete punctato, linea media laevi ; 
elytris lineato-punctatis, interstitiis subtilius uniseriatim punctatis 
subrugulosis, apice oblique declivi, fortiter excavato-retuso, 
margine elevato utrinque bispinato, infra subcirculatim emargi- 
nato. Long. 3°6 mm. 


Two examples, Miyanoshita. 


Cylindrical, pitchy-black. Head dull, convex, with long thin 
pubescence, mouth ciliate, front punctured at sides, with an 
elevated smooth space widened behind and impressed in middle, 
eyes broad oval, anteriorly emarginate, antennz ferruginous. 
Prothorax about one-half longer than broad, base slightly rounded, 
basal angles obtusely rounded, sides subparallel behind becoming 
rounded gradually and more strongly towards apex; surface 
cylindrical behind, declivous in front, but without median elevation, 
its pubescence thin, rather long at apex and sides, absent over 
basal area, anterior half with fine imbricate asperities, posterior 
half shining, distinctly punctured, with a smooth central line, the 
punctures stronger and more scattered towards the base. Scutel- 
lum small, rounded, not depressed. Elytra as wide as prothorax, 
and rather more than half as long again, base truncate, shoulders 
subrectangular rounded, sides parallel to behind middle, thence 
gently rounded to apical border, which is strongly and almost 
abruptly rounded, with the median third subcircularly emarginate ; 
surface shining, gently convex from base to apex, with a few 
hairs, distinctly punctured in rows, first stria with punctures 
stronger towards base, but not impressed ; interstices rugose, 
with a row of fine scattered punctures, the first widened postericrly 
with two or three fine tubercles before declivity, which is very 
oblique, beginning at the base of the posterior third of the elytra, 
and excavate; the excavation smooth, impunctate, and shining, 
with the suture narrowly ‘elevated, its lateral margins sharply 


120 Mr. Walter F. H. Blandford on the 


raised, crenate, fringed with a few long hairs, and armed with two 
spines, one small, near the suture, at the apex of the 2nd 
interstice, the other about the middle, at the apex of the 5th 
interstice, longer and directed backwards. Underside deep ferru- 
ginous. Legs ferruginous with knees infuscate. 


Readily distinguished from X. defensus by its larger 
size, colour, and the obliquity of the apex, the impressed 
surface of which is not circular but elongate; the apical 
emargination is much deeper and more abrupt. It is 
wlied to X. emarginatus, Eichh., but as the apical 
impression in that species is described as being sub- 
rugose and somewhat closely punctured, it is obviously 
distinct. This type of Xyleborus, with an impressed, 
emarginate, and spined apex to the elytra, appears to be 
rather common in the Oriental region. I have other 
undescribed species in my collection, which can easily 
be separated by comparison, though they run very close 
in structural features. ‘They are, in spite of their shape, 
true Xylebori, and show no generic differences upon 
dissection. 


It is a peculiarity of this genus that the descriptions 
and differentiations of the species it contains are based 
almost entirely on the characters of the females. The 
males are so rarely taken, that but very few have ever 
been described among exotic species. As they are sub- 
apterous, ard incapable of flight, they are not to be 
obtained except by a special search in the burrows they 
inhabit, a task usually too tedious to be attempted by a 
collector who is devoting his attention to one or more 
Orders in a foreign country. A further acquaintance 
with them would be of material assistance towards 
grouping the species of the genus, as, though small and 
ill-developed in comparison with the females, they 
present some well-marked differences of structure. In 
form they are of two types, one, short and subglobose, 
as X. dispar ¢, the other, cylindrical and similar to the 
female, but shorter, often more convex and less robust, 
as X. saxesent ¢, and XY. schaufussi ¢. For the insects 
whose males are of the former type, Ferrari has pro- 
posed the genus or subgenus Anisandrus, which is at 
present of no use, as one cannot yet say with certainty 
what characters of the female are connoted with that 
particular type of male, which probably merges into the 


Rhynchophorous Coleoptera of Japan. 121 


other by intermediate forms. Other differences, besides 
those of form, are found in the prothorax, which is 
frequently impressed in front, and furnished with a 
structure on its apical margin, varying from a minute 
tubercle, as in X. dryographus ¢, to a spine; in some 
species, as in .Y. coronatus, Hichh., from Venezuela and 
Brazil, and others from Madagascar, this median spine is 
strongly developed and bifid, and there are two lateral 
processes. In the present collection the males of but 
two species are associated with their females, Y. vicarius, 
Hichh., and X. schaufussi, m.; there are, moreover, the 
males of three species which I cannot assign with 
certainty to any female forms, and am compelled to 
describe under separate provisional names. ‘The dates 
and localities of capture have proved of no assistance 
towards identifying them. In one species, Y. cucullatus, 
the anterior margin of the prothorax presents a new 
modification, being produced obliquely forwards and 
downwards into a flat plate, which completely conceals 
the head, in a second, Y. orbatus, there is no process of 
the prothorax, and the general type is that of X. dispar 
¢, while the remaining species, X. galeatus, has a deep 
anterior prothoracic impression with a strong apical 
spinous process which conceals the head, and is of the 
type of Y. eurygraphus g, Ratz. 


Xyleborus cucullatus, sp. n. 


Mas. Breviter oblongus, subnitidus, pilosus, piceo-niger, 
antennis pedibusque ferrugineis, prothorace longitudine et lati- 
tudine subaequali, a basi fere oblique declivi, deplanato, et in 
lobuia transversum marginatum supra caput producto, dense 
punctato, punctis versus apicem exasperatis; elytris lineato- 
punctatis, interstitiis multipunctatis, ad apicem fortiter declivibus, 
subtruncatis, declivitate swbconvexa, immarginata, striata, granu- 
lata. Long. 2—2°6 mm, 


Four examples, Kurigahara, and Konose in Higo. 


Short, oblong, slightly shining, lighter or darker piceous, with 
rather long pubescence. Head completely hidden from above and 
in front, ferruginous, smooth and indistinctly punctured, pube- 
scence almost absent, eyes very small, quite flat with few facets 
antenne ferruginous, of the usual structure, the sensitive surface 


192 Mr. Walter F. H. Blandford on the 


of club very oblique. Prothorax rather broader than long (its 
apparent length varying according to the angle at which it is 
viewed), base truncate, basal angles obtusely rounded, sides 
rounded, above obliquely declivous and flattened anteriorly, the 
declivity prolonged backwards in the middle line to basal third, 
anterior border produced in a line with the declivous surface into a 
transverse lobe, covering head, its margin rounded and elevated ; 
surface obtusely elevated behind apical lobe and at sides of declivity, 
closely punctured, and with a more or less evident smooth median 
line from base to middle, the punctures, finely asperate anteriorly 
for a variable distance, interspaces finely reticulate. Scutellum 
small, rounded triangular. Elytra narrower than prothorax, an 
about one third longer, base truncate, basal angles rounded, humeral 
elevations scarcely traceable, sides parallel to behind middle, thence 
rounded to apex ; surface convex, subcylindrical at basal half, with 
indistinct rows of punctures, interstices multipunctate, apex 
very strongly but obliquely declivous, the declivity subconvex, 
circular in outline, not acutely margined, granulately punctured 
and indistinctly striate. Legs rather long, apices of anterior tibi» 
subtruncate, outer margin of middle and posterior tibie broadly 
rounded serrate. 


The specimens vary in colour, gloss, and the closeness 
and degree of asperity of the thoracic sculpture. One 
example from Higo is at first sight very different. It is 
smaller, darker, more truncate, with the prothorax 
shorter and the anterior lobe more declivous. It may 
quite possibly be the male of a different species, but 
presents no essential differences of structure or sculp- 
ture. ’. cucullatus is perhaps the male of X. brevis, 
Eicbh., and is certainly allied to it; but the Higo ex- 
ample alone appears small enough to stand as the male 
of that species, and in all the rows of punctures on the 
elytra are more evident than in _X. brevis. The specimen 
from Kurigahara has been placed, by Mr. Lewis, on the 
same card with the next described species, as the 
corresponding sex. But I do not see my way to accept- 
ing this. If it be so, XY. cucullatus must be the female, 
and in all characteristics it isa male; the generative 
organs, as far as I have been able to examine them, agree 
with those of Y. dispar g, the elytra appear to be 
soldered, and the wings are either completely absent or 
very minute. 


Rhynchophorous Coleoptera of Japan. 123 


Xyleborus orbatus, sp. n. 


Mas. Brevis, subglobosus, subnitidus, ferrugineus, longius 
pilosus, prothorace transyerso, aeque rotundato convexo, anterius 
exasperato, posterius rugoso; elytris a basi usque ad apicem 
rotundatis, lineato-punctatis, interstitiis subrugosis uniseriatim 
punctatis. Long. 1:5 mm. 


One example taken with the last species at Kurigahara. 


Of the type of Xyleborus dispar, g. Very short and subglobose, 
somewhat depressed, ferruginous with very long scattered pu- 
bescence. Head with front nearly flat, reticulated, sparingly 
punctured and pubescent, eyes small, antenne testaceous. Pro- 
thorax one-third broader than long, base truncate, basal angles very 
broadly rounded, sides and apex strongly rounded, surface 
uniformly but not strongly convex from base to apex, which is not 
produced in front, asperate anteriorly, the asperities becoming more 
scattered and punctiform towards base, interstices finely reticulate 
except at extreme base. Scutellum triangular. Elytra as wide as 
prothorax and half as long again, with humeral angles rounded, 
sides rounded from base to apex, rather more strongly behind 
middle, surface uniformly convex with indistinct rows of shallow 
punctures, interstices with a single series of punctures bearing long 
hairs. Legs long, slender, testaceous; tibia feebly dilated and 
spined. 


This insect may be the male of X. germanus, M., 
or semi-opacus, Hichh. I have described it for the 
sake of completeness, and not because I think that any 
useful purpose is served by describing these isolated 
males unless they exhibit some salient features. 


Xyleborus galeatus, sp, n. 

Mas. Oblongus, cylindricus, subnitidus, parce longius pilosus, 
ferrugineo-piceus ; prothorace oblongo., versus apicem fortiter 
excavato suleato, margine apicali in corniculum validum producto, 
supra anterius exasperato, posterius disperse punctato ; elytris ad 
apicem oblique et convexe declivibus, declivitate infra marginata, 
supra lineato-punctatis, interstitiis rarius uniseriatim punctatis, lo 
et 5o in declivitate tuberculatis. Long. 3-5 mm. 


One specimen near Nagasaki. 


Oblong, cylindrical, shining, deep ferruginous-brown, pubescence 
thin and long. Head completely hidden in front and below by 


124, Mr. Walter F. H. Blandford on the 


prothorax, antenne ferruginous. Prothorax longer than broad, 
subtruncate at base with posterior angles rounded, sides slightly 
rounded, apical angles strongly rounded, apex transverse but 
produced in middle into a strong triangular spine directed forwares 
and upwards, anterior opening of protkorax horizontal, slightly 
produced downwards below spine ; surface convex, with a median 
obtusely pointed elevation, in front of which is a deep triangular 
impression reaching apical spine, its anterior half very finely 
asperate, its posterior half with distinct scattered punctuation. 
Scutellum very small, triangular. Elytra narrower than prothorax 
and one-third longer, separately convex at base with margin 
elevated, humeral prominences obsolete, sides subparallel to apex, 
which is broadly rounded; surface nearly cylindrical, declivous 
and convex at apex, the declivity finely carinate below for a short 
distance, with rows of fine punctures ; interstices flat, subrugose, 
with a single row of remote piliferous punctures, on declivity more 
distinctly and transversely rugose, Ist and 3rd with four or five 
fine tubercles, 2nd with traces of two tubercles on summit only. 
Legs ferruginous, tibize strongly dilated and serrate. 

This insect is the male of a species in Hichhoff’s section 
+ + to which Xyleborus obliquecauda, Mots., X. aquilus, 
M., and X. validus, Hichh., belong. It is most like the 
last species, but does not resemble it very closely, and 
may be the male of a species of which Mr. Lewis has 
not taken the female. 


TRYPODENDRON, Steph. 


‘There are at present two Japanese species, one of which 
is merely a geographical variety of the European Trypo- 
dendron quercus, Kichh. 


Trypodendron quercus, Hichh., var. niponicum. 


Majus, prothorace pro maxima parte nigro, plaga basali testacea, 
elytrorum lateribus ad versuram apicalem angulatis, subproductis, 
apice utrinque distincte sinuato. Long. 36—4 mm. 


A dozen examples taken at Miyanoshita, and one at 
Oyayama. 

The black patches on the thorax and the elytral vittz 
are of the same shape as in the type, but better marked, 
the thorax being suffused as a rule with black except at 
the base, and the elytral vittee reaching the base in all 
mature specimens. ‘he most distinct character is the 
prominence of the angle of junction between the sides of 


Rhynchophorous Coleoptera of Japan. 125 


the elytra, and the apical margin which is thereby sinuate 
at the sides. This feature, though not usual in Huropean 
examples, is distinctly traceable in a less degree in one 
specimen of my series from the Ardennes, and is therefore 
not specific. I do not think that the identification of this 
species with Fabricius’ Apate signata is proved. 


Trypodendron pubipenne, sp. n. 

Oblongum, subnitidum, pilis longis adspersum, fusco-nigrum, 
elytris testaceis, sutura et apice infuscatis, antennis pedibusque 
testaceis ; clava antennarum ovali, haud acuminata; prothorace 
obscuro, anterius exasperato, posterius in medio subtilissime 
aciculato-punctato, lateribus muticis ; elytris apice obtuso, vix 
perspicue lineato-punctatis, interstitiis subrugulosis, multipunctatis, 

Mas. Capite angusto, fronte depressa, supra inter partes 
oculoram superiores carinula transversa nitida ornata; prothorace 
anterius fortiter angustato. 


Fem. Capite lato, fronte subconvexa; prothorace minus 
angustato, Long. 3 mm. 


Four specimens taken at Sapporo, Kiga, Miyanoshita, 
and Ichiuchi. 


Oblong, fuscous-black with elytra dull testaceous, darker along 
suture and at apex, moderately shining, with long downy 
pubescence. Head in male narrow with front flattened, longi- 
tudinally impressed, dull, sparsely punctured and pubescent, upper 
divisions of eyes with a narrow raised glossy margin continued as a 
transverse grooved ridge across vertex ; in female broader, sub- 
convex, transversely rugose, scantily punctured and hairy. Antenne 
with club oval, not acuminate, completely pubescent. Prothorax 
transverse, strongly constricted in front in male, less so in female, 
its sides rounded, apical margin bituberculate, acuminate in male, 
the whole surface dull with very fine granulations, gibbous in 
female, depressed in male, with scattered hairs, absent over sides of 
base, in front with asperate tubercles, becoming finer and produced 
in middle nearly tu base, which is without aciculations at sides. 
Scutellum piceous, triangular, pubescent. Elytra less than twice 
as long as prothorax and narrower than its greatest width, sides 
subparallel, apex obtusely rounded, surface with long pubescence 
chiefly at sides and apex, finely punctured in rows, interstices 
multi-punctate, subrugose, the punctures as in the allied species 
not readily distinguishable from those of the striew. Underside 
fuscous-black, almost impunctate, pubescent, 


126 Mr. Walter F. H. Blandford on the 


In the weakness of the elytral stria and the length 
of the pubescence this species is allied to T'rypodendron 
politum, Say (unicolor, Hichh.), from N. America, which 
I have not seen. It has no trace of a suture on 
the antennal club, nor is it especially narrow, both 
points being characteristic of J’. politum (Lec. Rhyne. 
N. Am., pp. 357, 358). It differs from Hichhoff’s 
description of 7’, wiicolor, in being not elongate, in the 
absence of a transverse elevated line on the prothorax, 
which is constricted in front, and in the elytra being not 
unicolorous. 


SCOLYTO-PLATYPINI. 


ScoLyto-PLaAtypus, Schauf. 


Scolyto-platypus tycon, m. 
daimio, m. 


3) 3 
os »  slomio, m. 
> » | mkado, m. 


I have nothing to add to my account of these insects 
(Trans. Ent. Soc. Lond., 1893, pp. 425—442). 


The following species is new, and should be placed 
after S. tycon. 


Scolyto-platypus shogun, sp. n. 
Oblongo-cylindricus, fere glaber, niger vel piceus, antennis pedi- 
busque ferrugineo-piceis ; prothorace transverso, subopaco, parce 
et subtiliter punctulato; elytris ante declivitatem haud striatis 
irregulariter punctatis, declivitate striata, interstitiis convexis 
seriatim tuberculatis. Long. 3°5 mm. 


Mas. Fronte excavata, opaca, pilis fulvis ciliatis circumdata, 
antennarum scapo fortiter clavato, funiculo brevi, flexili, clava 
elongata acuminata, ad apicem ciliata. 


Several examples, probably taken at Higo. 


Intermediate between S. tycon and daimio, and closely 
resembling dark specimens of the former in shape and 
size, but the elytra show no trace of striz, and are not 
pubescent at the apex. Male with head deeply excavate, 
dull with a thick marginal fringe of coarse hairs, antennze 


Rhynchophorous Coleoptera of Japan. 127 


constructed as in S. daimio 3, but with the ciub elongate 
and pointed. Female with antennal club a little more 
elongate than in S. tycon 2. Hlytra with the sutural 
stria alone faintly expressed in the male, declivity with 
Ist and 38rd interstices tuberculate throughout, the 
remainder more finely at the upper angle. Prothoracic 
foveee of male distinct. Anterior tarsal joints trigonate. 


The male has the prosternum prominent in the middle, 
the prominence corresponding to a wide deep anterior 
“nocket”; the anterior margin of the prosternum 
is constructed similarly to that of S. mikado $, but the 
two hooks are replaced by a transverse chitinous plate, 
the anterior angles ot which are acute and prominent. 


PLATDY PINE: 


One of the most admirable features of Chapuis’ “Mono- 
graphie des Platypides,’ upon which our knowledge 
of this sub-family is almost entirely based, is the ability, 
almost to be called intuition, with which he has grouped 
forms, often widely different in appearance, as_ the 
respective sexes of the species which he described ; and 
an examination of his own collection, or’ of any other 
containing species named by him, affords proof that his 
judgment was in the main correct, which is remarkable, 
if it be remembered that he had to reducé to order a 
vast and heterogeneous mass of material from all parts 
of the world. For he raised the number of species from 
15 (excluding a few unrecognized forms) to 202; and 
entomologists have been so far content to accept his 
work as final, that since the publication of his ‘‘ Mono- 
eruphie ” they have added but six species to those therein 
described. But, identification of two forms as the 
respective sexes of a single species is obviously quite 
compatible with error as regards the reference of them 
to their proper sexes, and it is a matter of common 
opinion among entomologists that he has, throughout the 
work, reversed the sexes and called the male the female 
and vice-versa. This was first suggested to me by the 
late Mr. Janson; it has not been, to my knowledge, 
explicitly stated in print, but Hichhoff has indicated a 


128 Mr. Walter F. H. Blandford on the 


doubt as to the correct interpretation of the sexes by 
Chapuis (“ Die eur. Borkenk.,” p. 306, note). M. Bedel 
(“« Coléop. du bassin de la Seine,” vi., p. 40+) inverts the 
sexes of P. cylindrus, as given by Chapuis, though with- 
out comment; he has kindly informed me that he was 
led by Hichhoff’s expression of doubt to dissect dried 
examples of P. cylindrus, which led him to the conclusion 
that Chapuis was wrong. Now Chapuis was probably 
guided by the generalisation of Perris, since disproved 
for the Tomicint by Lindeman and Hichhoff, that in the 
Scolytide a greater development of the elytral armature 
was a female characteristic, and by the sexual features 
of certain species of Crossotarsus, in which the antennee 
have the scape remarkably developed in the sex that he 
indicates as the male. ‘his antennal development, 
together with the deeply excavate and fringed front in 
the same sex of some species, and the more developed 
legs of the opposite sex (in Crossofarsus), which agree 
with the sexual features of the Scolyto-platypini, - are 
not easily reconciled with the usual sexual characters, if 
the sexes are reversed; but the opposite may be said of 
the constantly stronger elytral armature of the females 
(Chap.), and their occasional possession of abdominal 
armature (Crossotarsi genuini, Platypus blanchardi) or of 
a concave and short abdomen (Crossotarst). It is 
obvious that external sexual characters are not constant 
among the Scolytide, and that little dependence can be 
placed on them. The question can only be decided by 
direct examination of the generative organs. Like 
M. Bedel, I have examined them in dried specimens of 
P. cylindrus, and also of Crossotarsus wallacet. ‘This 
examination has given results which leave me little doubt 
in the matter; but it so difficult, in these cases, to 
correctly interpret the structures exhibited in dried 
specimens, that I do not feel justified at present in 
publishing my conclusions. I can, however, say that 
the sexes of those two species, as given by Chapuis, 
correspond, and there is no evidence of his having 
indicated them rightly in some genera and erroneously 
in others. He is entirely right or entirely wrong. For 
the present, therefore, | prefer to describe the sexes in 
accordance with Chapuis, as a matter of convenience 
solely, and without implying acquiescence in his views. 


Rhynchophorous Coleoptera of Japan. 129 


This will cause no confusion, and the same cannot be 
said of a readjustment of the sexual characters which 
should eventually be proved to be itself erroneous. 

The question can be settled beyond dispute by the 
dissection of fresh examples; and if J can obtain them 
J hope to solve it in that way. 


Crossorarsus, Chap. 
This Oriental genus is represented by three species. 


Crossotarsus chapuisi, sp. n. 

Fem, Sub-elongata, picea, nitidissima ; capite dense ac rugose 
punctato; prothorace irregulariter punctato, punctis ante 
suleum et versus basim crebrioribus; elytris striato-punctatis, 
interstitio 30 ad basim dilatato, laevi, ad apicem subangustatis, 
leniter declivibus, apice pilis aureis ciliato, supra depressionem 
posticam angustam sublinearem producto, angulo externo vix 
perspicue elongato ; abdominis segmento apicali fortiter excavato- 
impresso ; tibiis posticis dilatatis ad apicem in lobos productis, 
Long. 8 mm, 


One specimen, Higo. 


More elongate than the majority of the genus, shining piceous, 
smooth and regularly cylindrical. Head with front subquadrate, 
very closely and rugosely punctured, the punctures longitudinal, 
with an indistinct median depression, vertex with three shining 
longitudinal vittze separated by punctured spaces, eyes rather large 
and prominent, subcircular, antenne with scape enlarged towards 
the base or inner side, but not produced, funiculus inserted at 
apex of scape. Prothorax oblong, not strongly emarginate, 
posterior angles of emargination rather prominent, median sulcus 
fine, surrounded by a smooth cordate area; punctures longi- 
tudinally oval, sparse over apical half, closer at base and over a 
patch in front of sulcus, lateral margin with two or three deep 
asymmetrical pore-like punctures behind middle. Scutellum 
indistinguishable. Elytra double as long as prothorax, with sides 
narrowed and slightly constricted at apex ; striate, the striz with 
an irregular single row of punctures grouped in pairs, 3rd and 4th 
conjoined at base, interstices smooth, flat, with very fine irregular 
punctures, 3rd dilated at base with a few stronger punctures; 
gently depressed towards apex, with the striz at first deeper and 
then obliterated before posterior margin, interstitial punctuation 

TRANS. ENT. SOC. LOND. 1894.—parRT 1. (MARCH.) I 


130 Mr. Walter F. H. Blandford on the 


stronger, suture longitudinally impressed before apex ; posterior 
margin gently rounded and fringed with aureous pubescence 
which conceals the very shortly produced apical angles ; 
posterior impression narrow, sublinear, concave, hidden by 
apical margin. Underside ferruginous, with scanty pubescence, 
abdomen concave, not strongly punctured, sides of 2nd and 4th 
segments scarcely produced, last segment with a large deep 
transversely oval impression, bordered in front by a fringe of 
aureous hairs, and reaching the prominent apical margin. Anterior 
tibix with five carine, and indistinct traces of others at base ; 
intermediate tibiz with a single carina on anterior surface ; 
posterior tibiz dilated, with outer border convex, and apex 
produced into two serrate lobes on either side of tarsal articu- 
lation, densely ciliate, with yellowish hairs. Posterior coxe with 
a prominent free margin to inner and apical borders, which with 
the posterior margin of the trochanter is sharply serrate. 


I cannot refer this fine and distinct species to any 
of Chapuis’ groups. I therefore propose for it a distinct 
group, Crossotarst mutici, characterized in the female 
by the gently declivous elytra, without elevated inter- 
stices or conspicuously produced apical angles, and with 
a narrow deep posterior impression, in which it 
approaches C. minav, Walk. The structure of the 
posterior tibia, and the abdomen, together with the 
absence of a distinct scutellum, are also characteristic 
features. The specimen is undoubtedly a female—in 
the conventional sense. 


Crossotarsus niponicus, sp. 1. 


Ferrugineus, elytris in medio paullo dilutioribus, postice 
infuscatis ; prothorace subquadrato, irregulariter punctato, punctis 
ad basim haud profundioribus; elytris lineato-punctatis, stria 
suturali per totum, ceteris modo ad basim apicemque impressis, 
apice convexe declivi, angulis externis productis, depressione 
angusta ; abdominis segmento primo spinula obliqua armato. 


Mas. Interstitiis elytrorum in declivitate non elevatis, ad 
basim granulatis, angulis externis brevius productis, abdominis 
spinula brevi. Long. 6—6°5 mm. 

Frm. Interstitiis in declivitate subelevatis, striis subsulcatis, 


spinula abdominis longa, segmento 50 subconcayo. Long. 5°7— 
6-2 mm, 


Rhynchophorous Coleoptera of Japan, 131 


Generally distributed; numerous specimens taken at 
Sapporo and Hakodate in Yezo, also at Miyanoshita, 
Yuyama, etc., and in Kiushiu. 


Lighter or darker ferruginous-brown, with head and apical 
third of elytra infuscate ; the elytra sometimes evidently lighter 
in the middle, particularly in the males, Head with front 
flattened, dull, with longitudinal strigose punctures which are 
stronger in the female, and with a longitudinal impressed line, 
nearly obsolete in the male, vertex with three shining vitte ; 
antennz with scape simple, slightly dilated internally at base. 
Prothorax subquadrate, its central furrow shallow ; in the female 
with the sides of the furrow slightly elevated, with irregular 
punctures, the punctures closer and rather deeper towards sides 
behind emargination, internal to which there is a shallow depres- 
sion; in the female without sides of sulcus elevated, with fine 
irregular punctures, usually rather closer on two slight impressions 
on either side of anterior extremity of sulcus, one or two 
impressions along lateral margin are usually present but not 
constant. Elytra longer in the male than in the female, in both 
sexes with rows of fine punctures, first stria impressed throughout, 
more strongly at base, the rest impressed at base and on declivity, 
3rd and 4th conjoined at base ; interstices with a very fine ground- 
work of punctures, and one or two larger punctures, 3rd finely 
granulate at base in male; apex gently declivous with external 
angles produced more strongly in female, striz of male impressed but 
not dilated, interstices flat, pubescent, strize dilated in female, sub- 
suleate, interstices pubescent, convex ; marginal impression narrow, 
shining, tuberculate at extremities of 3rd and 4th interstices. 
Underside ferruginous, 1st abdominal segment with an oblique 
spine, long in female, tuberculiform in male, apical segment 
strongly punctured in female and concave. Legs ferruginous. 


This species belongs to the Crossotarsi subdepressi, 
and is closely allied to C. fairmairei, Chap., from 
which it can be separated by the absence of the close 
variolose punctures at the base of the prothorax, and 
by the second interstice of the elytra not being im- 
pressed in the middle. The produced angles of the 
elytra are longer, and extend more obliquely backwards 
than in C. fairmairet. 


Crossotarsus contaminatus, sp. 0. 


Fem. Picea, fronte subconcava, opaca, disperse punctata ; 
prothorace oblongiusculo, in medio ntrinque impresso, irregulariter 


132 Mr. Walter F. H. Blandford on the 


punctato, ante suleum congerie punctorum confluentium notato ; 
elytris subtiliter striato-punctitis, striis ad basim impressis, inter- 
stitiis laevibus, rarius subtiliter punctulatis, ad apicem subde- 
clivibus, angulis externis subtus productis, interstitiis variolose 
punctatis, pilosis, depressione postice angusta, lunata ; abdominis 
segmentis inermibus, 50 subconcavo, Long. 5:3 mm. 


One example, Higo; three without locality. 


Piceous ; head with front subconcave, shortly pubescent, dull 
with scattered punctuation, more strongly over mouth, the 
punctures elongate towards vertex, and with a median impressed 
line, vertex with a median shining line and two indistinct lateral 
lines, antennz with scape simple, linear. Prothorax longer than 
broad, its lateral emargination very feeble, slightly impressed on 
either side at middle of surface, sulcus very fine, not reaching 
base, and terminated in front by an irregular patch of coalesced 
shallow punctures, punctuation of rest of surface scattered, 
irregular, stronger in front and on either side of a median smooth 
line running from central patch to apex. Elytra shining, finely 
striato-punctate, the strise wider and deeper at base, 3rd and 4th 
not conjoined, sutural stria impressed throughout, interstices 
scarcely convex, with a few fine scattered punctures, rather closer 
at base, without reticulate or punctured ground; apical extremity 
rounded and declivous, with interstices coarsely and rugosely 
punctured and pilose, not carinate, the punctuation of the Ist 
not extending farther forwards than that of succeeding interstices, 
external angles produced downwards in the plane of the terminal 
depression, which is narrow, lunate and subconcave, forming a 
marked angle with the surface of the elytra. Underside deep 
ferruginous, with metasternum and abdomen piceous, middle of 
former and latter strongly punctured; abdominal segments unarmed, 
2nd and 4th narrowly produced at sides, 5th concave. Legs 
piceous. Posterior coxee with a spine at internal angle ; posterior 
femora not crenate below. 


This species must, I think, be placed with the 
Orossotarsi subdepresst, though the interstices are not 
carinate at the apex of the elytra, and the marginal 
impression is more sharply separated from the surface 
than is usual in that group. It can be distinguished 
from C. niponicus by the absence of a spine on the first 
abdominal segment, and by the drd and 4th elytral 
strie not meeting at base, so as to shorten the 4th 
interstice. 


Rhynchophorous Coleoptera of Japan. 133 


Pratypus, Hbst. 


The five species in this collection are all new; the 
males of two alone are represented. 


TABLE. 
1. Elytra with impressed striz 
with rows of fine punctures REAR Morice re ir 
2. Apex of elytra with a small terminal impression, unarmed . 3 
declivous and convex, with two small tu- 
bercles ondeclivity . . . modestus, 9 
produced at externalangles. . . . tata oe 
3. Prothorax quadrate, with a narrow patch of close eacennee 
round sulcus, the anterior two or three large 
and porelike, elytra pilose from base to apex 
modestus, 3 
oblong, with a broad cordate patch of uniform 
punctures rovnd sulcus, elytra glabrous before 
apex a: HK i Gay “Wewisd, ig 
4. Interstices spined at summit aE apie decuenty of elytra ; 4th 
abdominal segment with two spines . .  lewisi, Q 
Interstices not spined, 4th abdominal segment unarmed 
severini, 
5. Terminal impression oval, with an inferior emargination not 
reaching middle . . . . calamus, 
lunate, emarginate to middle . hamatus, 


+0 


+0 +0 


Platypus modestus, sp. n. 


Ferrugineus, prothorace subquadrato, sparsim piloso ; elytris 
sulcatis, sulcis postice latioribus, interstitiis subconvexis, nitidis, 
rarius pilosis ; apice convexe declivi, interstitiis opacis, granulatis, 
pube densiore vestitis. Long. 5 mm. 

Mas. Fronte valde concava, opaca, fundo bifoveolato ; protho- 
racis sulco congerie punctorum circumdato, punctis inaequalibus, 
depressione elytrorum postica parva, subtriangulari, granulata. 

Fem. Fronte quam minime concava, opaca ; prothorace medio 
utrinque impresso, subaequaliter punctato, punctis in utroque sulci 
latere paullo crebrioribus; declivitate ad apicem interstitii 31 
spinula ornata. 


Four specimens, Nikko and Shimidzu Toge. 


Rather robust and not elongate; ferruginous, with bead and 
apex of elytra darker, the latter nearly testaceous at base, 


134 Mr. Walter F. H. Blandford on the 


MALgE, with front of head rather deeply concave, its surface 
closely granulate, slightly shining in parts, with a fine central 
impressed line and a circular fovea on either side ; vertex convex, 
with a shining central and two indistinct lateral vitte. Prothorax. 
subquadrate, median sulcus with a narrow patch of punctures on 
either side, the front half of each patch formed of the usual close 
small punctures, except for the two anterior punctures, which are 
larger and almost porelike, and those of the posterior half, which 
are larger, shallow, and less numerous ; remainder of surface rather 
regularly punctured and thinly pubescent. Elytra sulcate, the 
sulci punctured throughout, becoming a little wider towards apex, 
interstices convex, shining, very finely punctured and pubescent, 
drd and 5th closely granulate at base, 2nd and 4th abbreviated 
at base with a few small granules ; apical extremity convex, the 
sulci shallower, interstices granulate, more thickly pilose, posterior 
impression small and not well marked, subtriangular, irregularly 
granulate. 

FEMALE, with front very slightly concave, quite dull, very 
finely punctured above, with short median impressed line, vertex 
with median vitta alone shining. Prothorax subquadrate with an 
impression on either side about middle, nearly uniformly 
punctured, the punctures rather closer at sides of sulcus. Elytra 
with surface sculptured as in male, but with the sulci wider 
behind middle and their punctures coalesced ; all interstices granu- 
late for a short distance at base ; apical extremity convex, the 
striz ceasing before posterior margin, interstices dull, granulate 
and pilose, apex of 3rd marked by a small pointed tubercle. 

Underside and legs testaceous or ferruginous, abdomen more 
convex in the male. 


One pair is rather darker, and the surface of the 
thorax and elytra are hairless before the apex. ‘This is, 
however, due to the specimens being more mature and 
having been rubbed. ‘The species belongs to the 
Platypi sulcati, and is not unlike P. jansoni, Chap., but 
the prothorax is shorter and the elytral interstices are 
less elevated, and are not spined at the margin of the 
apical declivity. 

Platypus lewisi, sp. n. 

Ferrugineo-piceus, prothorace oblongo, elytris sulcatis, interstitiis 

convexis, glabris, nitidis. Long. 5°5 mm. 


Mas. Fronte plana, opaca, prothoracis sulco congerie punctorum 
magna, cordiformi, circumdato, elytrorum interstitiis subsimilibus, 


Rhynchophorous Coleoptera of Japan. 135 


ad apicem, 30 et 50 etiam ad basim granulatis, depressione postica 
parva, subtriangulari, granulata. 


Fem. Fronte antice subconcava, opaca, prothoracis sulco con- 
gerie punctorum minore ovali circumdato, interstitiis lo et 20 in 
summa declivitate in spinam communem magnam, 3o, 50, To in 
spinulas parvas productis, declivitate utrinque tuberculata, angulis 
externis in lobos subquadratos productis, abdominis segmento 
40 bispinato. 


Five specimens, Miyanoshita, Kiga, and Yuyama. 


Elongate, ferruginous, or inclining to pitchy. 

Mate with front of head flat, dull with very fine cross reticula- 
tion, punctures rather fine, longitudinally strigose towards mouth. 
Prothorax oblong, diffusely punctured, rather closer at extreme sides, 
with a porelike puncture within anterior angle of emargination 
and a broad cordate group of small uniform punctures round 
median sulcus. Elytra sulcate, the sulci with irregular confluent 
punctures, not widened behind and obliterated before apical 
impression ; interstices convex, shining, with fine scattered oblong 
punctures, lst narrow throughout, bases of 2nd and 4th abbreviated, 
impressed and more strongly punctured, the latter with two or 
three longitudinal asperities, bases of 3rd and 5th elevated with 
transverse granulations ; apical extremity with interstices dull, 
granulate, and pilose, terminal impression subtriangular, closely 
granulate, with erect hairs, suture shining. Underside reddish 
testaceous, abdomen unarmed. 


FEMALE with head similar to male in sculpture, front impressed 
over mouth and subconvex behind. Prothorax oblong, slightly 
impressed in middle of either side, sulcus surrounded with an oval 
patch of punctuation narrower than in male, rest of punctures 
scattered and rather fine, except along lateral border. Elytra sulcate, 
the sulci wider behind with punctures confluent, interstices convex, 
shining, with fine scattered oblong punctures, the ist narrow 
throughout, base of 3rd enlarged, more closely punctured, bases of 
2nd and 4th impressed, punctured, and asperate as in male, two first 
with a large common spine, 3rd, 5th, and 7th with small spines 
at summit of apical declivity, the rest unarmed ; declivity convex 
with the sulci continued on it, and the interstices finely asperate, 
the third terminating at a stout tubercle, external angles produced 
backwards into a vertical sabquadrate lobe, with its posterior edge 
concave, so as to form two blunt teeth atangles. Fourth abdominal 
segment armed with two spines, the fifth flattened. 


136 Mr. Walter F. H. Blandford on the 


This species is allied in its main features to the Platypi 
sulcati, with which I must associate it. It differs from 
the other described species in the strongly produced 
external apical angles of the elytra, which are not 
triquetrous as in the P. trispinati. The female 
shares with P. quadridentatus, Ol. (blanchardi, Chap.), 
the peculiarity of possessing two spines on the fourth 
abdominal segment, but is much larger and easily 
distinguished by the group of punctures on the thorax. 
The male is separable from that of P. modestus by its 
larger size, flat forehead, and oblong thorax; the elytra 
are quite glabrous before the apical extremity. 


Platypus severiivi, sp. n. 

Fem. Piceo-ferruginea, elytris postice infuscatis; fronte sub- 
concava, rugosa ; prothorace paullo longiore quam latiore, utrinque 
ad medium vitta obliqua subelevata, antice rarius irregulariter, 
postice crebrius punctato ; elytris ad apicem subdeclivibus et in 
processus divaricatos attenuatis, striato-punctatis, striis impressis, 
interstitiis subconvexis ad apicem pilosis, duobus primis per totum, 
ceteris ad basim subtiliter punctulatis, processibus desuper aspectis 
bidentatis. Long. 5—5:3 mm. 


Numerous examples of one sex only, taken from beech 
at Nikko, Chiuzenji, and Hakodate. 


Deep ferruginous brown, with elytra darker towards apex. 
Head with front subconcave, covered with close shallow rugose 
punctuation, rather smoother towards mouth, vertex rather abruptly 
separate from front, with three smooth vitte, interspaces coarsely 
punctured. Prothorax a little longer than broad, median sulcus 
fine but sharply marked, surface with very fine reticulation, and an 
indistinct oblique elevation at either side of anterior extremity 
of sulcus, punctures of anterior half scattered, and absent over 
middle line, of posterior half closer, shallow, and longitudinally 
oval. Elytra declivous towards apex and produced into two 
divergent processes, with subsulcate striz, which are wider and 
shallower towards apex, with the punctures fused ; interstices convex, 
finely reticulate and punctured at base, the first and second with a 
single row of punctures along inner border, all interstices flatter 
towards apex, inconspicuously tuberculate, and with serrate hairs ; 
apical processes declivous in the plane of the posterior termination, 
produced outwards to form an obtuse oval emargination at apex, 


Rhynchophorous Coleoptera of Japan. 137 


their upper border curved and continuous with second interstice, 
extremity two-spined when seen from above, inner spine longer and 
truncate. Last abdominal segment subconvex, rugosely punctured. 


The species belongs to the Platypi oxyuri, and may be 
distinguished from the Pyrenean P. orywrus, Dup., by 
the thoracic sculpture, and by the processes of the elytra 
being two-spined when seen from above instead of from 
the side, as in the latter species. From P. solidus, 
Walk., the divergence of these processes will at once 
separate it, as will its larger size, and the convex 
interstices which, except the first two, are impunctate 
after the base. 


Platypus calamus, sp. n. 


Fem. Elongata, angusta, ferrugineo-testacea, elytris apice 
infuscatis ; fronte concava, linea media impressa; prothorace 
sesqui longiore quam latiore, in utroque sulci latere plaga punc- 
torum angusta ; elytris lineato-punctatis postice haud declivibus, 
interstitiis planis, vix perspicue punctatis, apice oblique excavato- 
impresso, ovali, inferne emarginato, angulis externis oblique 
productis, margine externo elevato sinuato, ad suturam supra 
valde obliquo, depressione nitida, ante angulos extremos impressa. 
Long. 3°7 mm. 


Sixteen examples of one sex only, taken at various 
places, Miyanoshita, Oshima, Kiushiu (Higo, Yuyama, 
etc.). 

Slender, reddish testaceous, elytra with apex darker. Head with 
front concave, anteriorly dull, with a deep longitudinal impressed 
line in middle, above it, with rather strong scattered punctures, 
upper part forming an obtuse angle with vertex, which has only 
the median elevated line present, narrow and infuscate. Prothorax 
one half longer than broad, the sulcus fine, with a narrow group 
of punctures on either side, closer anteriorly, remainder of 
surface irregularly punctured, the punctures more evident towards 
the base, anterior and posterior borders with fine transverse 
reticulation. Elytra finely punctured in lines, sutural stria 
impressed throughout, remainder at base only ; interstices with a 
few fine punctures, 1st, 3rd, and 5th elevated at base, the former 
impressed with a fine line beginning behind base, 2nd and 4th with 
base evidently punctured. Apex of elytra with an oblique terminal 
impressed surface, its margin elevated and slightly everted, curving 


1388 Mr. Walter F. H. Blandford on the 


away from suture so as to form an angle with its fellow above the 
terminal impression, and then sinuate at sides; external angles 
produced in a curve downwards and backwards, their outer border 
serrate ; impressed surface very shining, elevated at suture, and 
concave above external angles, its outline an incomplete oval, 
emarginate below, the emargination deeper than wide, but shorter 
than the terminal surface measured along the suture, its sides 
nearly straight, not toothed, and apex rounded. Last abdominal 
segment concave. 


This species is one of the Platypi cupulati, and can 
be separated from all described species except the 
following, by the fact that in them the suture is notched 
or narrowly emarginate above the terminal impression, 
and the svtural border forms a sharp and distinct angle 
with the upper margin of the impression, whereas in these 
species the sutural margin is gradually rounded into that 
ef the impression without trace of any angle, so as to 
form a wide emargination. It can also be separated from 
most by the absence of any trace of a tooth on the lower 
apical emargination. 


Platypus hamatus, sp. n. 


Fem. Elongata, ferrugineo - testacea, elytris apice paullo 
obscurioribus ; fronte impressa, in medio foveolata ; prothorace 
oblongo, punctis magnis et parvis irregulariter notato, parte antica 
sulci utrinque punctis paucis circumdata; elytris lineato- 
punctatis, interstitiis planis, parcissime punctatis, apice excavato- 
impresso, lunato, inferne late emarginato, angulis extremis longe 
productis, margine supra depressionem intus ad suturam curvato, 
convexo. Long. 4°2 mm 


Two examples at Yuyama and Miyanoshita. 


Elongate, reddish testaceous, with elytra darker at apex. Head 
with front subconcave, foveolate in middle, finely reticulate, and 
rather coarsely punctate above, vertex strongly punctured with 
a fine median elevated infuscate line. Prothorax more elongate 
than in P. calamus, sulcus deeper at extremities than in middle, 
with about a dozen punctures on either side of anterior extremity, 
remainder of surface with irregular punctures of two sizes, a 
slight impression on either side, about middle more closely 
punctured, anterior and posterior borders finely reticulate. Elytra 
with sculpture of surface similar to that of P. calamus, termi- 
nating in a vertical impressed surface, which is lunate and widely 


Rhynchophorous Coleoptera of Japan. 139 


emarginate below, lateral borders meeting obliquely at suture 
above impression, and curved out as in P. calamus, and then 
sinuate, external angles produced downwards and_ backwards, 
longer and more curved than in the latter species; terminal 
impression emarginate almost to middle, the emargination broader 
than long, with inferior sutural angles very slightly produced, its 
surface shining, foveolate on either side, with suture elevated and 
tuberculate. Last abdominal segment concave. 


This species, also one of the Platypi cupulati, is best 
separated from P. calamus by the structure of the apical 
impression, the length of which measured along the suture 
is about equal to the depth of the emargination, whereas 
in P. calamus it is about double its depth. The tuber- 
cles along the suture, the minutely produced sutural 
angles at the fundus of the interior emargination, and a 
fine serration of the superior border, where it curves 
away from the suture, are peculiar to this species. P. 
hamatus is evidently closely allied to P. forficula, Chap., 
of which I have been unable to see a typical specimen. 
I have, however, an unnamed Platypus, taken by Mr. 
Wallace in Gilolo, which agrees with the description of 
P. forficula, and is probably to be referred to that species, 
as Chapuis appears to have seen all Wallace’s Platypini. 
It most closely resembles P. hamatus, but the sutural 
border of the elytra makes a sharp angle with the upper 
margin of the terminal impression, which is much 
narrower than in the latter species, being not half as 
deep as the inferior emargination. 


Diapus, Chap, 


An Oriental genus, represented in Japan by one 
species. 


Dapus aculeatus, sp. n. 


Frm. Picea, margine prothoracis postico et elytris ad basim 
dilutioribus ; fronte deplanata, in medio breviter carinata ; protho- 
racis disco nitidissimo, basi fortiter punctata ; elytrorum interstitiis 
lo, 20, 30 elevatis, 30, 50 ad apicem in spinulas validas productis ; 
7o breviter spinoso, extus cum 80 et 90 serratis conjuncto, angulo 
externo in spinulam obliquam angustissimam producto, depressione 
postica breviter bispinata. Long. 2°8 mm. 


Two examples, Higo. 


140 Mr. Walter F. H. Blandford on the 


Rather slender and very shining, head and prothorax deep 
piceous, the latter with basal border narrowly fusco-testaceous, 
elytra fusco-testaceous at base, becoming pitchy towards apex. 
Head with front nearly flat, impressed slightly over mouth, dull, 
with rather distinct and strigose punctuation below, the punctures 
less apparent towards vertex, in the middle with a short median 
elevated keel, which is not perceptible towards vertex ; elevated 
lines of the latter very evident and shining. Prothorax very 
shining, rather deeply emarginate at sides, apex with a row of 
piliferous punctures, rest of surface quite impunctate, except at 
base which is closely and strongly punctured. Elytra with first 
two strize and marginal stria impressed throughout, third and 
fourth at base only, outer striz indicated by lines of irregular 
punctures, which coalesce towards apex, interspaces almost im- 
punctate ; 1st interstice with a minute spine at apex, 3rd and 5th 
with two long spines, 7th with a shorter spine, the outer border 
of which is conjoined with two serrations formed by the ex- 
tremities of the succeeding interstices, external angle with a very 
slender spine directed obliquely downwards, apex fringed with a 
few long ciliate hairs, apical impression oblique, subconvex, with 
two spines shorter than the three long spines of the interstices. 
Underside testaceous, metasternum and abdomen piceous, last seg- 
ment of the latter nearly flat, dull, fringed with long curled cilia. 


This species is closely allied to D. quinque-spinatus, 
Chap., of which it may be the Japanese form. It 
presents the following distinctive features: the colour is 
deeper and the surface more shining, the frontal carina 
is shorter, nearer the mouth, and not traceable towards 
the vertex as in D. quinque-spinatus, in which it appears 
to be longer and interrupted; the prothorax is quite 
impunctate except at the apex and base, and the basal 
punctures are more numerous; the interstices of the 
elytra are scarcely punctured, the spine of the 7th 
interstice is shorter and that of the 9th only represented 
by a small tooth, the spine at the external apical angle 
is pecuhar to this species, as is the terminal fringe of 
hairs; the spines of the posterior depression are also 
shorter. 


Rhynchophorous Coleoptera of Japan. 141 


(Genus incerte sedis.) 


Grnyocerus, Motsch. 
Ktud. ent. vii., p. 68. 
G. adustipennis, Motsch., l.c. ix., p. 19. 


Whatever this insect may be, it cannot be a Scolyto- 
platypus, as it is described in these terms: Figura 
Platypi cylindri, sed glabrior, etc. 


Par 
ens, Mi) 
aA 


iY ve 
RAN ee 


X a Ms 
Sa ak ta 
i) wt 


) Aa SRN say a ee ; AAR 0 Alaa ve 
, i hs \ ip : at ee : er" i i ils 
Th Chante i ¢ 


3 
=~ 
™ 


( 143 ) 


V. Description of the female of Hypochrysops scintillans, 
Butl. By Hamitron H. Drucz, F.Z.S8. 


[Read Feb. 7th, 1894. ] 


We have lately procured a fine pair of this species ; 
the ¢ only of which has been described, and as the ¢ 
presents considerable differences I have written the fol- 
lowing deseription. ‘They were received from Mioko, 
New Ireland, the ¢ agreeing with the type in the 
National Museum from New Britain. 


Hypochrysops sceimtillans. 


Miletus scintillans, Butl., Ann, and Mag. Nat. Hist. 
(5), x; p. 150: (1882). 

Hypochrysops scintillans, H. H. Druce, Trans. Ent. 
Soc. Lond. (1890), p. 191. 


@. Upperside brown with a creamy-white central patch on 
forewing ; no trace of biue. Forewing rich dark brown, with a 
disc of creamy-white arranged much as the blue in H. cratevas ¢?, 
and extending from the base to beyond the cell, lightest in colour 
towards the centre of the wing, and becoming gradually sub- 
merged in the brown borders, the inner margin being lighter 
brown than the rest of the wing. Hindwing brown, considerably 
paler than the forewing, darker along the subcostal nervure to 
the base of the subcostal nervule, where it becomes somewhat 
suffused ; the median nervure from about its middle, and the two 
median nervules from their bases, distinctly and evenly yellow. 
Costal margin from base nearly to apex pale creamy-white. 
Upper half of anal fold greyish, lower concolorous with wings. 
Cilia of both wings brown, that of hindwing being rather darker 
than ground colour. Underside much as in ¢, but the median 
space of forewing (bordered by the lower wall of the cell and the 
subapical red band) creamy-white in place of greyish-brown ; the 
red bands of the hindwing being somewhat wider, and their 
metallic borders scarcely so brilliant. Thorax and abdomen 
brown ; head dark brown, with a whitish frontal line, and a yellow 
inner border to the eye ; legs and palpi yellow. Antenne black, 
white spotted below, yellow tipped. Expanse, 9 1,3, in., ¢ 1} in. 


144 Mr. H. H. Druce on Hypochrysops scintillans. 


Hab. Mioko, New Ireland (Mus. Druce). 


In December 1890, I published, in the ‘Transactions’ 
of this Society, a short monograph of the genus, 
recognizing thirty-two species of these gorgeous little 
butterflies, and to that number must be added a new 
species, since described by Mr. Miskin in the “ Anns. 
Queensl. Mus.,” No. 1 (Supp.), p. 85 (1891), under the 
name H. apollo, from Herbert River, W. Queensland, 
and two other Australian species described by the same 
author in the “P. Z. 8. N.S.W.” (ser. 2) iii, pp. 1517 & 
1518 (1888), viz.: H. ewelides and H. olliffi; the former 
allied to H. chrysanthis, Feld., the two latter to H. 
ignita, Leach, 


VI. A list of the Lepidoptera of the Khasia Hills. Pt. II. 
By Colonel Cuartes Swinnog, M.A., F.L.S. 


[Read Oct. 18th, 1893.] . 
Prats II, 


HETEROCERA. 
i. 


In arranging the order in which the Heterocera should 
stand I have been working on the lines set forth in 
Mr. Hampson’s excellent classification in Faun. Brit. 
Ind., but I cannot altogether follow the order in which 
he has, for the sake of convenience, put the different 
families, and prefer dividing them into three groups. 


Ist. Fore wing with vein 5 (lower radial) from 
centre of discocellulars, or nearer 6 than 4. 


2nd. Fore wing with vein 5 from lower angle of 
cell, or nearer 4 than 6. 


3rd. Fore wing with veins 4 to 11 arising from the 
cell at almost even distances apart. 


| Grovp I. 


— 


Sub Group 1. No. 1. Sphingide  - Frenulum present. 
” 2, ,, Ll. Saturniide - 


5 2 Brahmeide -\|q. 

es eect ta of Frenulum absent. 
3 od. ,, Ll. Hupterotide - 

4, 1. Notodontide - 

3) ” ay 

nos et at ag, Seam eee 
Fs 5. ,, 1. Epicopiide -)Frenulum  rudi- 

» 2 Uraniide | mentary or ab- 

» 0 Micronude -) sent. 

» 4, Hpiplemide - Frenulum present. 
eA 6. ,, 1. Geometers - Frenulum present. 

Group II. 


Sub Group 1. No. 1. Callidulidx “Le preset sometimes 


: resent sometimes 
Ge Tye ) AG 
5, 2 Drepanulide acai. 


TRANS. ENT. SOC. LOND. 1894,.—ParTI, (MARCH.) Kk 


146 
Sub Group 1. No. a Limacodidx 


. Lymantriide 
. Hypside 

. Arctiide 

. Agaristide 


. Castniidx 


i) 


. Psychide 

. Cosside. - 
. Arbelide 

. Hepialidexe 
. Syntomidee 
. Zygaenide 
. Noctuidee 

. Thyridide 
. Pyralide 


. 
1 O> 
° . 

~~ 


po Eno = DOIG eno ag or oe 


Group III. 


Sub Group 1. No. 1. Sesvide - 
ue 


1. Tineide - 


3) 33 


Group I. 


Sub Group I. 


. Lasiocampidx fan absent. 


Colonel Charles Swinhoe on the 


~ +Frenulum present. 


. Pterothysanidx 


Frenulum present. 


Frenulum absent. 


Frenulum present. 
- Frenulum present. 


. Laietulam present. 


*y 


a t Frenulum present. 


Family SPHINGIDA. 
Sub Family ACHERONTIIN A. 


Genus AcHrrontiA, Ochsen. 


1. A. styz, Westw., Cab. Or. Ent., p. 88; pl. 42, fig. 3. 


2. A. lachesis (Fabr.), Ent. 
27 (1798). 


A. morta, Hiibn., 


Syst. 


Suppl., 434, 26, 


Verz., p. 140 (1816). 


Sub Fam. SMERINTHIN A. 
Genus LANGLIA, sees 


8. DL. zeuzeroides, Moore, P. Z. 
L. khasiana, Moore, L.c., p. a 


, 1872, p.. 560. 


Recorded from the Khasias; not received by me, 


Lepidoptera of the Khusia Hills, 147 


Sub Fam. AMBULYCINA. 
Genus Catymnta, Walker. 
4. C. panopus (Cram.), Pap. Hxot., iii., p. 50, pl. 224, 
figs. A, B (1779). 
Recorded from the Khasias, 


Genus Amsunyx, Westw. 
5. A. substrigilis, Westw., Cab. Or. Hnt., p. 6) pl. 30: 
fig. 2 (1848), 
Shillong and Cherra Punji. 
6. A. semifervens (Walker), xxxi, p. 38 (1864), 
A. turbata, Butler, P. 7. Sy, L87a,0.252., 
Cherra Punji. 
7. A. deucalion (Walker), viii., p. 236 (1856). 
Recorded from the Khasias, 


8.. A. gigantea, Rothschild, Nov. Zool., i, p. 96 (1894), 
Recorded from the Khasias. 


Sub Fam. CHAROCAMPINA, 
Genus Acosmeryx, Boisd. 


9. A. sericeus (Walker), viii., p- 181 (1856). 
Many examples ; Shillong. 


10. A. cinerea, Butler, P. 7. Si Soy pe 245, 
Several examples; Shillong and Cherra Punji. 


11, A. naga, Moore, Cat. Lep. E. I. C., 1.,p. 271 (1857). 

Shillong and Cherra Punji. Out of the numerous 
specimens of this genus received there has not been a 
single typical anceus, Cram. Hampson has put these 
four very distinct species together ; they appear to me 
to be very distinct, easily distinguishable and without 
intermediate forms. 


148 Colonel Charles Swinhoe on the 


Genus Camrocampa, Dup. 


12. C. macromera, Butler, P. Z. 8., 1875, p. 7. 
Shillong. 


13. OC. alecto (Linn.), Syst. Nat., i., p. 803 (1767). 


Common. 


14. C. theylia (Linn.), Lc. 
C. rafflesii, Butler, Trans. Zool. Soc., 1876, p, 556. 
Examples of both forms of this species. 


15. ©. oldenlandie (Fabr.), Sp. Ins., ii., p. 148 (1781). 


Common. 


16. C. vagans (Butler), Ill. Typ. Lep. Het. B. M., v., 
p. 4, pl. 78, fig. 7 (1881). 


One example; Shillong. 


17. OC. mydon (Walker), viii., p. 155 (1856). 
Three examples ; Shillong. 


18. C. olivacea (Moore), P. Z.S., 1872, p. 567. 


Common. 


19. C. velata (Walker), xxxv., p. 1853 (1866). 
Pergesa aegrota, Butler, P. Z.8., 1875, p. 246. 


Metopsilus albomarginatus, Rothschild, Nov. Zool., i., 
p- 78 (1894). 


Shillong and Cherra Punji; in great numbers. Hamp- 
son has included aurifera, Butier, and acuta, Walker, with 
this species ; they appear to me to be distinct ; they have 
broad pale yellow prominent bands on each side of the 
thorax, a character wanting in velata. The form albo- 
marginatus is not uncommon in the Khasias ; I cannot see 
how it can be separated from velata: in this variable 
species there are scarcely any two examples identical. 


20. OC. castor (Walker), viii., 153 (1856). 


Many examples, a common species in the Khasias ; 
omitted by Hampson. I have it from both Shillong and 
Cherra Punji, it seems to be a link between velata and 


Lepidoptera of the Khasia Hills. 149 


olivacea, having the colouration of the former with the 
white marginal suffusion on the fore wings above, of the 
latter. 


21. C. busiris (Walker), viii., p. 158 (1856). 
Panacra hamiltoni, Rothschild, Nov. Zool., 1., p. 82. 
Cherra Punji. 


The type specimen is much faded, but I have a fine 
series and feel certain of my identification. 


22. C. helops (Walker), viii, p. 180 (1856). 
Philampelus orientalis, Felder, Reise Nov. Lep., pl. 77, 
fig. ] (1874). 
One example; Shillong. The type of helops is recorded 
from Port Natal, which must be a mistake. 


23., C. clotho (Drury), Ill. Exot. Ins. u., p. 48, pl. 28, 
fie. 1 (1773). 
Sphina butus, Cram., Pap. Exot., u., p. 88, pl. 152 
Ghats). 
Deilephila cyrene, Westw., Cab. Or. Ent., p. 138, pl. 6, 
fig. 1 (1848). 
Many examples. The type of cyrene is in O, M.; it 
is identical with clotho. 


24. CO. punctivenata, Butler, P. Z. S., 1875, p. 248. 

Many examples. I have both sexes from Cherra Punji. 
It should not be confounded with elotho. Gonograpta 
from Southern India also appears to be a good species. I 
have a long series in my Museum. 


25. C. lineosa, Walker, viil., p. 144 (1856). 
26. C. major, Butler, P. Z.8., 1875, p. 249. 
27. C. minor, Butler, l.c. 


Shillong and Cherra Punji; in great numbers. At 
first I put them together, following Hampson, but with 
the numbers now before me I cannot do'so. They are 
undoubtedly somewhat alike, as are all the species of 
this group, but each has its own distinctive wing 
markings, and major has a pale yellowish broad longitu- 
dinal shade, central on forewings, discal on hindwings, 
which is very characteristic. 


150 Colonel Charles Swinhoe on the 


28. C. pallicosta, Walker, viii., p. 145 (1856). 
Three examples; Shillong. 


Genus Darunis, Hubn. 
29. D. nerit (Linn.), Syst. Nat., i., 2, p. 798 (1767). 
Recorded from Khasias. 
30. D. bhaga, Moore, P. Z. 8., 1865, p. 794. 
Numerous specimens from Shillong and Cherra Punji. 


Genus THrererra, Hubn. 


31, T. nessus (Drury), Ill. Ins. Exot., n., p. 46, pl. 27, 
fig. 1 (1778). 
Shillong and Cherra Punji. 


82, T. actea. (Cram.), Pap. Exot., ui., p. 938, pl. 248; 
fig, as(1779). 
Common. 


Genus Exipia, Walker. 


38. H. dolichus (Westw.), Cab. Or. Ent., p. 61, pl. 30, 
fig. 1 (1848). 
Shillong and Cherra Punji. 


Sub Fam. SPHINGINE. 
Genus Protoparcr, Burmeister. 


34. P. orientalis, Butler, Trans. Zool. Soc. 1876, p. 609, 
pl. 91, figs. 16, 17 (larva). 
Common. 
Genus Pseuposrumx, Burmeister. 
30. P. inexacta (Walker), viii., p. 208 (1856). 
Many examples. 
36. P. fo (Walker), viii., p. 195 (1856), 
Shillong and Cherra Punji. 
37. P. discistriga (Walker), vili., p. 209 (1856). 
Common. 


Lepidoptera of the Khasia Hills. 151 


38. P.khasianwm (Rothschild), Nov. Zool.,i., p. 90 (1894). 
Recorded from the Khasias; must be closely allied to 
if distinct from the preceding which is a very variable 
species. 
Genus Nupuete, Hiibner. 
39. N. hespera (Fabr.), Syst. Ent., p. 546 (1775). 
Recorded from the Khasias. 
Genus Aprocatypsis, Butler. 
40. A. velov, Butler, Trans. Zool. Soc., 1876, p. 641. 


One example; Shillong. 


Genus PsrvpopoLpina, Rothschild. 
41. P, velowina, Rothschild, Nov. Zool.,i., p. 91, pl. 6, 
fig, 18 (1894). 
Recorded from the Khasias. 


Sub Fam, MACROGLOSSIN 
Genus Gurutca, Kirby. 
42. G, hyas (Walker), viii., p. 107, 1856. 
Shillong. 
43. G. masuriensis (Butler), P. Z. §., 1875, p. 244, 
pl. 3663. 
Numerous specimens, the common form apparently. 
I have received it from both Shillong and Cherra Punji. 
Genus RuoratorsycHeE, Butler. 
44, R. nycteris (Kollar), Hiig. Kasch., iv., 2, p. 408, 
pl. 19, fig. 5 (1844). 
Cherra Panji. 


Genus Macrogtossa, Scop. 
45, M. belis (Cram.), Pap. Exot., i, p. 147, pl. 94, fig. c 
(L276). 
Common. 
46. M. gilia, Herr Schiiff., Samml. Ausser. Eur. Schm., 
pp. 59, 79, pl. 23, fig. 107 (1850). 


Several examples. 


152 Colonel Charles Swinhoe on the 


47, M. proxima, Butler, P. Z. 8., 1875, p. 4, pl. 1, fig. 1. 
Three examples ; Shillong. 


48, M. imperator, Butler, P. Z. §., 1875, p. 243, pl. 37, 
fig. 4. 
Cherra Punji. 
Genus CepHnopzs, Hiibn. 
49. C. hylas (Linn.), Mant., i., p. 539 (1771). 
Recorded from the Khasias. 


Genus Saraspes, Moore. 


50. S. infernalis (Westw.), Cab. Or. Ent., p. 61, pl. 30, 
fig. 3 (1848). 


Common. 


Sub Group IT. 
Fam. SATURNIIDA. 
Genus Acrtras, Leach. 


51. <A. selene (Hiibn.), Samml. Exot. Schm.,i., pl. 172, 
fig. 3 (1806-19). 


Common. 


52. A. menas, Double., Ann. Mag. N. H., xix., p. 95, 
pl. 7, fig. 1 (1847). 


Common. 
Genus Arracus, Linn. 
53. A. atlas, Linn., Mus. Lud.-Ulr., p. 366 (1764). 
Common. 


54. A. edwardsi, White, P. Z. 8., 1859, p. 115, pl. 57. 
Shillong and Cherra Punji. 

00. A. cynthia, Drury, Exot. Ins., u1., pl. 6, fig. 2 (1773). 
Very plentiful. 

56. A. ricini, Boisd., Ann. Soc. Ent, Fr. (8), i, p. 790 


(1854). 
Many examples. 


Oo 


Lepidoptera of the Khasia Hiils. W5e 


Genus AnTHEeRmA, Hiibn. 


57. A. roylei, Moore, P.Z.S., 1859, p. 256, pl. 64, fig. 1. 
Cherra Punji. 

58. A. paphia (Linn.), Syst. Nat., i, p. 809 (1767). 
Cherra Punji. 

59. A. helferi, Moore, P.Z. S., 1859, p. 257, pl. 64, fig. 2. 
Cherra Punji. 

60. A. knyvetti, Hampson, Faun. Brit. Ind. Moths, 1, 

p. 19 (1892). 

One female ; Shillong. 


This species has been in my collection for some years 
under Moore’s M.S. name of tumida. 


Genus Saturnia, Schrank. 


61. S. extensa (Butler), Ill. Typ. Lep. Het. B. M.,, v., 
p. 61, pl. 94, fig. 2 (1881). 
Two examples ; Shillong. 


Genus Lozpa, Moore. 


62. L. katinka (Westw.), Cab. Or. Ent., p. 25, pl. 12, 
fig. 2 (1848). 


Numerous examples. 


Genus Satassa, Moore. 

63. Salassa megastica, sp. n. 

dé @. Of the same brick-red colour as in S. lola, but very 
much larger, the antemedial line of forewings much closer to the 
base, white outwardly edged with brown, very much outwardly 
curved and runs close into the base on the hinder margin ; the 
hyaline spot is large, ringed with a black and a white line deeply 
excavated on the outer side, the excavation filled up by a deep 
black patch ; on the hindwing the difference is greater, the hyaline 
spot is larger than on the forewings, but of a similar nature, the 
rings round it are black, white, and again black, then a broad light 
red space, which on the outer side is very broad and extends well 
beyond the discal black and white spotted band, which is thin in 
the disc and curves broadly round the ocellus, the space between 
it and the red portion being filled in with dark grey; halfway 


154 Colonel Charles Swinhoe on the 


between this discal band and the outer margin is a duplex brown 
band, the inner portion straight, the outer dentate. The female, 
instead of being brown, as in S. lola, merely differs from the male 
in having the brown bands darker, and the hyaline spots very large 
on both wings, being nearly half an inch in diameter, and of 


about equal size. Expanse of wings g 9, 6,%, 6,3; inch. 
Three males and one female, from Cherra Punji. 
Genus Cricuna, Walker. 
64. C. trifenestrata (Helfer), Journ. As. Soc. Beng., v1., 
p. 45 (1837). 
Common. 


Fam. BRAHM IDA. 
Genus Braumama, Walker. 
65. B. wallichii, Gray, Zool. Misc., p. 39 (1852). 
Many examples; Shillong and Cherra Punji. 


66. B. hearsayi, White, Proc. Ent. Soc., 1861, p. 26. 
Eleven examples; Shillong. 


Fam. BOMBYCIDA. 
Genus Ocrnara, Walker. 


67. O. signifera (Walker), Journ. Linn. Soc., vi., p. 130 
(1862). 
Shillong and Cherra Punji. 
68, O. cyproba, sp.n. (Pl, ih, fe.) 


36 9. Antenne reddish grey, body and wings pure white, 
wings thinly clothed, almost semi-hyaline, with many grey bands ; 
first ante-medial, indicated on hindwing by cell spot and a short 
band at abdominal margin, second discal third sub-marginal, both 
complete, these bands on the forewings are crossed by longitudinal 
similar bands, which fill up the second, third, and fifth interspaces, 
almost from the base to the centre margin, and are crossed by the 
white veins on both wings. Expanse of wings ¢ 1,35, 9 1,4 inch. 

Many males and two females, from Cherra Punji and 
Shillong. 

Far more thinly clothed than O. signifera, and with- 
out any spots. 


Lepidoptera of the Khasia Hills. 155 


69. O. apicalis, Walker, Journ. Linn. Soc., vi, p: 180 
(1862). 


Shillong and Cherra Punji. 


Sub Group III. 
Fam. EUPTEROTIDA. 


Genus GancariDes, Moore. 
70. G. roseus (Walker), xxxil., p. 513 (1865). 
Two males and one female, Cherra Punji. 
Genus Drzuata, Walker. 


71. D. hades, Walker, iv., p. 908 (1855). 

Shillong and Cherra Punji. Many males and females. 
The females only differ from the males in their larger 
size and brighter colour, having sometimes a reddish 
tinge. 

Genus Patirisa, Moore. 
72. P. lineosa (Walker), iv., p. 912 (1855). 

Shillong and Cherra Punji. One male and several 
females. 

73. P.cervina (Moore), P. Z. 8., 1865, p. 807. 

Shillong, one pair. 

Genus Tacora, Walker. 
74. T. patula, Walker, v., p. 1189 (1855). 

T. undulosa, Walker, v., p. 1196. 

Cherra Punji. 

75. J. khasiana (Moore), Descr. Ind. Lep. Atk., i, 
Dati, L809: 

Shillong and Cherra Punji. 

76. T. pallida (Walker), iv., p. 912, 1855. 

Sphingognatha asclepiades, Felder, Reise Nov. Lep., 

iv., pl. 94, fig. 1 (1868). 

Cherra Punji. 

I have received numerous specimens of patula and 
khasiana, and one male and two females of pallida. 


156 Colonel Charles Swinhoe on the 


Hampson puts the first two together, but there are un- 
doubtedly three species and not two. There is a small 
pale fawn-coloured species, d 3x5, ? 4 inches in expanse 
of wings, similarly coloured in both sexes, identical with 
Moore’s type of khasiana, very uniform in size; it has a 
black head and pale tuft of hairs at basal joint of antennz. 
A medium sized form, ¢ 4:4, 2 5 inches, uniform in size, 
very variable in colour, from pale fawn colour to dark grey 
and dark reddish fawn colour in the male, always of the 
latter colour in the female, the females of which fairly 
well correspond to Walker’s types of patula and wn- 
dulosa. This species has also in the male a black head 
and pale tuft of hairs. Finally, there is a large species 
of which I have received only one male and two females, 
measuring, d 5, 6:5 inches. The male is coloured 
pale fawn colour like the males of pallida from Java and 
Malayana in my collection. It has an ochreous-red head 
and tuft of hairs a little darker than in the other species. 
The females are dark reddish-fawn coloured like the 
females of patula, and also identical in colour with 
Malayan and Javan females of pallida. 


Genus Pszuposana, Hampson. 
77, PB. incandescens (Walker), iv., p. 910 (1855). 
Cherra Punji; two females. 
Genus Ganisa, Walker. 
78. G. postica, Walker, v., p. 1190 (1855). 
Shillong. 
79. G. pandya (Moore), P. Z. S., 1865, p. 807. 
Shillong. 
Genus ApHA, Walker. 
80. A. subdives, Walker, v., p. 1180 (1855). 
Shillong. 
Genus Huprerore, Hiibn. 
81. H. lineosa (Walker), vi., p. 1440 (1855). 


Murlida fraterna, Moore, Trans. Ent. Soc., 1884, 
pearls 
Shillong. 


Lepidoptera of the Khasia Hills. 157 


82. H. calandra, sp..n. 

¢ @. Antennz (of the male) with the plumes grey; palpi, 
head, and fore part of thorax ochreous ; shaft of antennz, body, 
and wings, above and below, bright citron-yellow, absolutely with- 
out any markings ; thorax beneath and legs ochreous. Expanse of 
wings, ¢ 24, 9 3:4 inch. 

Cherra Punji and Shillong. Numerous examples of 
both sexes. Antenne of the male weak, with the pec- 
tinations short as in #. geminata, Walker, but the wings 
are broader, with margin of fore wings less oblique. 
The absence of all markings makes it very distinctive. 


83. H. assimilis, Moore, Trans. Ent. Soc., 1884, p. 363. 
Fi. dissimilis, Moore, l.c., p. 368. 
Cherra Punji. 

84. H. testacea (Walker), iv., p. 906 (1855). 
Shillong ; common. 


85. H. avesta,sp.n. (Pl. IL, fig. 2.) 

3. Plumes of antenne grey ; palpi and head red-brown ; body 
and wings above of a uniform sandy-yellow colour ; wings crossed 
by several brown bands ; first antemedial outwardly curved, indis- 
tinct, obsolete in hind wings ; second medial, macular curved in on 
to the costa of fore wings, followed by three equidistant discal 
bands, outwardly curved, and composed of small spots. The 
last on hind wings is linear, a sub-marginal darker band of double 
lunules, which on hind wings is single and sinuous, cilia brown, 
underside darker, with the inner band obsolete on both wings ; all 
the other bands present and prominent ; legs pink-brown, covered 
with yellow hair ; 2 pale reddish-brown, with the inner lines more 
or less obsolete. Expanse of wings, 2,4; inch. 

Shillong and Cherra Punji; several examples. 

Allied to Hi. testacea, but the bands on the latter are 
linear and mostly straight. 


Genus Nisaga, Walker. 
86. N. simplex, Walker, iv., p. 885 (1855). ; 
Shillong and Cherra Punji. 


The female of this species, of which I have received 
two, has simple antenne, the body is dark, the wings 
pale brickdust-colour, slightly tinged with pink. 


158 Colonel Charles Swinhoe on the 


Sub Group IV. 
Fam. NOTODONTIDA. 
Genus Tarsouepis, Butler. 
87. T. fulgurifera (Walker), xiv., p. 1347 (1857). 
Cherra Punji; one example. 


88. T. remicauda, Butler, Ann. Mag. N. H. (4), x, 
p. 125, pl. 8 (1872). 


Shillong; one example. 


Genus Dupvusa, Walker. 


89. D. nobilis, Walker, xxxu., p. 447 (1865). 
Shillong ; one female. 


This is the first record of this species within Indian 
limits ; the type came from North China. I have it also 
from Celebes and Singapore. 


Genus Barapesa, Moore. 


90. B. lithosioides, Moore, P. Z. 8., 1883, p. 17, pl. 5, 
fig. 2. 
Shillong and Cherra Punji; numerous specimens. 


Genus Patera, Moore. 


91. P. parivala, Moore, Cat. Lep. EH. I. C., ii., p. 434 
(1859). 
Shillong and Cherra Punji. 
92. P. raya, Moore, l.c., p. 434. 
Shillong and Cherra Punji. 


. Genus Garcerta, Walker. 
93. G. costigera, Walker, xxxu., p. 455 (1865). 
Shillong and Cherra Punji; common. 


94. G. curvaria, Hampson, Faun. Brit. Ind. Moths, 
1; poo Gleg2}: 


Shillong. 


Lepidoptera of the Khasia Hills. 159 


Genus Turnaca, Walker. 


95. T. delineivena, sp. n. 

2 Palpi dark brown with pale tips to the hairs, antenne 
reddish-brown, forewings and thorax grey, forewings with the 
median and outer veins dark brown and with brown bands on each 
‘of these veins, more broadly and better defined in the median vein 
along its whole length, hinder portion of the wing also suffused 
with brown, hindwings brown with pale costal margin. Expanse 
of wings, 2.°; inch. 

Cherra Punji; two examples, 
Genus Twacona, Walker. 
Porsica, Walker. 
96. T. ingens (Walker), xxxv., p. 1823 (1866), 
Shillong ; two examples. 


Genus Pypna, Walker. 

97. P. testacea, Walker, vii., 1754 (1856). 

Common, 

98, P. kamadena (Moore), P. Z. 8., 1865, p. 812. 
Shillong. I have both sexes of this species, it is 

quite distinct from the preceding, differiug in size, 

markings, and colouration. 

99, P. decurrens (Moore), Desc. Ind. Lep. Atk., i., 
p. 66 (1879), ; 
The type came from Cherra Punji, not received by 

me. 

100. P. notata, Swinh., Trans. Ent. Soc., 1891, p. 479, 

ple LO; tie, 16: 

Shillong; one example. 

Hampson makes this a synonym of P. pallida, Butler, 
from Japan, but it cannot even belong to the same 
section of the genus, having simple antenne in both 
SeXeS. 

101. P. longivitta (Walker), vii., p. 1754 (1856). 
Shillong ; two males, one female. 

102. P. bela, sp. n. 


Antenne grey ; upper side of palpi dark brown; underside 


160 Colonel Charles Swinhoe on the 


yellowish-grey ; head, thorax, and forewings dark reddish-ochreous ; 
abdomen and hindwings white, tinged with ochreous ; forewings 
irrorated with brown atoms; these, clustered together, form a 
longitudinal fascia in the centre up to the apex and indications of 
another fascia below it; some indistinct brown spots in an out- 
wardly curved line from centre of hinder margin towards apex, 
only apparent in one example ; hind wings with the outer border 
tinged with red. Expanse of wings, 1,4 inch. 
Cherra Punji; three examples. 
Genus Crira, Walker. 


103. C. aurata (Moore), Descr. Ind. ep. Atk., 1, 
p. 64 (1879). 
Recorded from the Khasia Hills, by Moore. 
104. C. fasciata (Moore), l.c., p. 66. 
Shillong ; two specimens. 
Genus Somera, Walker. 
105. S. viridifusca, Walker, iv., p. 882 (1855). 
Shillong and Cherra Punji; many examples. 
Genus Crerura, Schrank. 
106. C. liturata, Walker, v., p. 988 (1855). 
Shillong. 
Genus Hyrermscura, Butler. 


107. H. pallida, Butler, Ann. Mag. N. H. (5) vi. 
p. 65 (1880). 


Shillong; numerous examples. 
108. H. basalis (Moore), P. Z. S., 1865, p. 813. 
Cherra Punji. 
Genus AntHsua, Walker. 
109." A. servula (Drury), Ins. Mxot, a p.20, plidie 
fio, 11 (1773). 
Cherra Punji. 
Genus Anticyra, Walker. 
110. A. combusta, Walker, p. 1092 (1855), 
Cherra Punji. 


Lepidoptera of the Khasia Hills. 161 


Genus Fantonta, Butler. 
111. F. argentifera (Moore), P. Z. 8., 1865, p. 813. 
Cherra Punji. 
Genus Nutri, Walker. 
112. N. viridescens (Walker), vi., p. 1504 (1855). 
Cherra Punji. 
| Genus Spatanta, Hiibner, 


113. S. argentifera (Walker), Journ. Linn. Soe., vi., 
p» 140 (1862). 
Shillong. 
114. S. awritracta (Moore), P. Z. S., 1865, p. 811. 
Shillong. 


115. S. plusioides (Moore), Descr. Ind: Lep. Atk., i., 
p. 62 (1879). 
Cherra Punji. 
Genus Lornoprrmryx, Stephens. 


116. L. ferruginosa, Moore, Descr. Ind. Lep. Atk., i., 
B07 (S79): 


Cherra Punji. 
Genus Brsara, Walker. 


117. B. rubiginea, Walker, xxxii., p. 459 (1865). 
Recorded from the Khasias. 


Genus MerascHais, Hampson. 


118. M. disrupta (Moore), Descr. Ind. Lep. Atk., i., 
p. 62 (1879). - 


Cherra Punji. 
Genus Icutuyura, Hiibner. 
119. TF. fulgurita, Walker, XXX, p. 433 (1865), 
Shillong. 
120. I. pallida (Walker), v., p. 1077 (1855). 


Shillong, one example. 
TRANS. ENT, SOC, LOND. 1894,.—ParRT I. (MARCH.) Vi 


162 Colonel Charles Swinhoe on the 


Fam, CYMATOPHORIDAL, 
Genus Haprosyne, Hiibner. 
121. H. indica (Moore), P. Z. S., 1867, p. 44. 
Shillong. 


122. H. armata, Moore, Descr. Ind. Lep. Atk., ii, 
p. 90 (1882). 


Recorded from the Khasia Hills. 
Genus T'nyatira, Hiibner. 
1238. T. batis (Linn.), Syst. Nat., i., p. 836 (1735). 
Shillong and Cherra Punji. 
Genus Gaurena, Walker. 
124. G. florescens, Walker, xxxii., p. 620 (1865). 
Recorded from the Khasias. 
Genus Potypioca, Hiibner. 


125. P. albicosta (Moore), P. Z. 8., 1867, p. 45. 


Cherra Punji. 


Sub Group V. 
Fam. EPICOPIID. 


Genus Epica1a, Westw. 


126. H. polydora, Westw., Arcana Ent., p. 19, pl. 5, 
fig. 1 (1843). 


Shillong and Cherra Punji. 
127. H. varunea, Moore, P. Z. 8., 1865, p. 799. 


Shillong and Cherra Punji. 


Fam. URANIIDA. 


Genus Nycratemon, Dalman. 


128. N. zampa, Butler, Ent. Mo. Mag., v., p. 273 
(1868). 


Shillong and Cherra Punji. 


Lepidoptera of the Khasia Hills, 168 


Fam. MICRONIIDA. 


Genus Uraprerorpes, Moore. 
129. U. astheniata (Guén.), Phal., ii., p. 24 (1857). 
Cherra Punji. 
Genus StropuipiA, Hiibner. 


130. S. fasciata (Cram.), Pap. Exot., i1., p. 12, pl. 104, 
fig. p (1779). 
Shillong. 


Genus Micronipia, Moore. 


131. M. simpliciata (Moore), P. Z. 8., 1867, p. 646. 
Cherra Punji. 


Genus Micronia, Guén. 
132. M. aculeata, Guén., Phal., ii., p. 12, pl. 18, 
fig. 8, 9 (1857). 


M. gannata, l.c., p. 26, g 
M. sparsaria, Walker, xxiil., p. 818 (1861). . 


Shillong and Cherra Punji; common. 


Genus Psrupomicronia, Moore. 
133. P. celata, Moore, Descr. Ind. Lep. Atk., i, 
p. 257 (1887). 
Shillong ; one example. 


Genus Acroptsris, Hiibner. 
134, A. striataria (Clerck.), Icon., pl. 55, fig. 4 (1759). 
Shillong and Cherra Punji. 
135. A. iphiata (Guén.), Phal., ii., p. 29 (1857). 
Micronia pontiata, Guén., l.c. 
M. convewaria, Walker, xxiii., p. 819 (1861). 
Common. 


The type of convexaria came from Landoor, it was 
omitted from Cotes and Sunbury’s catalogue by over- 
sight. 


164 Colonel Charles Swinhoe on the 


136. A. ciniferaria (Walker), xxxv., p. 1642 (1866). 
Micronia obliquaria, Moore, P. Z.8., 1877, p. 622, 
pl. 60, fig. 17. 
Shillong and Cherra Punji. 
The type of ciniferaria from Siam is in the O.M., 
the type of obliquaria is from the Andaman Islands ; 
they are not separable. 


137. A. vagata (Moore), Pia. 8.; I877; p. G22; pleas 
ne, 18; 
Shillong. 
Genus AvuzEA, Walker. 

138. A. rufifrontata, Walker, xxvi., p. 1487 (1862). 

Cherra Punji. 

Genus Caatamia, Moore. 

139. CO. flavescens (Walker), ii., p. 406 (1854). 

Shillong. 


Fam. EPIPLEMIDAS. 
Genus Amana, Walker. 
140. A. angulifera, Walker, ii., p. 662 (1855). 
Shillong and Cherra Punji. 
Genus Decretia, Walker. 
141. D. mimicusaria, Walker, xx., p. 95 (1860). 
Shillong and Cherra Punji; several examples. 


Genus Parapecertta, Noy. 


With the veination of Decetia, but with very different 
antenne. In Decetia the antennee are pectinated in both 
sexes. In this genus the antenne of the male are 
dentated, the teeth with flat ends and very close 
together; in the female the rudiments of the dentations 
only are present. 


142. P. albistellaria (Walker), xxvi., p. 1522 (1862). 
Shillong and Cherra Punji. 


IT have it also from the Shan States (Manders). The 
type is in B.M., without locality. 


Lepidoptera of the Khasia Hills. 165 


Genus HrteLema, Herr Schiff. 
143. H. reticulata (Moore), Descr. Ind. Lep. Atk., iu., 
: p. 259 (1887). 
Shillong. 


144. H. himala (Butler), Ann. Mag. N. H. (5), vi., 
p- 221 (1880). 
Shillong. 
145. H. insolita (Walker), xxxv., p. 1648 (1866). 
Divades conchiferata, Moore, Lep. Ceylon, iii., p. 403, 
pl: 186, fig. 1h (1887). 
Cherra Punji. 


146. H. fulvilinea, Hampson, Ill. Typ. Lep. Het. B. M., 
vilil., p. 103, pl. 150, fig. 19. (1891). 
Shillong. 


I have this also from the Nilgiri BUTS and from F'ort 
Stedman, Shan States. 
147, H. quadricaudata (Walker), xxiii., 847 (1861), 

Hrosia varisaria, Walker, xxili., 848, and xxvi., 1797 

(1862). 

Cherra Punji. 

148. H. moza (Butler), Ann. Mag. N.H. (5), 1, p. 402 
(1878). 

Cherra Punji. 

The type of this species came from Japan, the Cherra 
Punji are almost identical with it. 
149. H. ocusta, sp. n. 

¢. Pale chocolate-brown, irrorated with black and brown 
atoms, both wings crossed by two pale lines, ante and post medial, 
arising from costa of forewings, from which they are first extended 
outwards, then abruptly curving inwards run direct to the hinder- 
margin, are extended across the hindwings, both being deeply 
elbowed outwardly in their centres, the inner line edged outwardly 
and the outer line inwardly with black, the black edgings being 
broad in parts, the outer portion of both wings are suffused with 
black and brown, two indistinct black spots near hinder angle of 
forewings and an ochreous patch near outer margin above the 
middle on the hindwings ; on the forewings there is a sub-marginal 
pale line outwardly edged with black, from the apex, stopping at 


166 Colonel Charles Swinhoe on the 


one-third before the hinder angle, at which part of the margin 
the wing is produced into a small angle; marginal line white, 
cilia dark brown edged with ochreous grey. Expanse of wings, ;°> 
inch. 


Cherra Punji; three examples. 


150. H. sreapa, sp. n. 

gd. Blackish brown, nearly uniform in colour, striations black, 
two black transverse lines across both wings, .ante and post 
medial most distinct on costa of forewings from which they run 
outward, and then become sinuous, directly descending to the 
hinder margin, the outer one having white points; on the hindwings 
they are not sinuous, the inner one is almost invisible, the outer 
one is double, an ochreous line between them, it is bent acutely 
outwards in its centre where it has two small teeth, and some _ 
white points in its lower half, a black sub-marginal streak on 
forewings below the apex, marginal line ochreous, fringe blackish 
brown. Expanse of wings, 1,2, inch. 


Cherra Punji; one example. 
Genus Drrapzs, Walker. 
151. D. bicaudata (Moore), P. Z.8., 1867, p. 645. 
Cherra Punji. 
Genus Orupiza, Walker. 
152. O. protheclaria, Walker, xxiii., p. 858 (1861). 
Common. 
Sub Group VI. 
GEOMETERS. 


Fam. MENOCTENIIDA. 
Genus Sarcinopes, Guén. 
153. S. carnearia, Guén., Phal., 1, p. 188 (1857), 
Very common, 
154. 8. restitutaria (Walker), xxvi., p. 1527 (1862). 
Auxima sumatraria, Walker, xxxv., p. 1577 (1866). 


Sarcinodes xgrota, Butler, Il. Typ. Lep. Het. B. M., 
vi., p. 60, pl. 115, figs, 3, 4 (1886). 


Very common. 


Lepidoptera of the Khasia Hills. 167 
155. S. equilinearia (Walker), xxi., p. 292 (1860). 
Shillong ; several examples. 


156. S. lilacina, Moore, Descr. Ind. Lep. Atk., in., 
p. 234 (1887). 


Cherra Punji and Shillong; common. 

157. S. debitaria (Walker), xxvi., p. 1527 (1862). 
Cherra Punji and Shillong; several examples. 

158. S. susana, Swinh., Trans. Ent. Soc., 1891, p. 488. 
Shillong and Cherra Punji; many examples. 


Genus Norra, Walker. 
159. N. ajaia (Walker), Journ. Linn. Soc., i, p. 195 
(1859). 
Shillong. 


Genus Eomeri, Duncan. 


160. H. rosalia (Cram.), Pap. Exot., iv., p. 152, pl. 368, 
fig. r. (1782). 
Common. 
161. H. flavata, Moore, Lep. Ceylon, iii., p. 440, 
pl. 198, fig. 3 ¢ (1887). 
Shillong; one example. 


162. HE. aureliata, Guén., Phal. i., p. 394, pl. 22, fig. 6 
(1857). 


Shillong. 


Fam. ORTHOSTYXID Ai. 


Genus Ozota, Walker, xxiv., p. 1080 (1861). 
Carima, Walker, xxvi., p. 1630 (1862). 
Zarmigethusa, Walker, l.c., p. 1637. 

163. O. microniaria, Walker, xxiv., p. 1080 (1862). 
Cherra Punji. 

164. O. extersaria (Walker), xxiii., p. 926 (1861), 
Shillong and Cherra Punji. 


168 Colonel Charles Swinhoe on the 


165. O. biangulifera (Moore), Descr. Ind. Lep. Atk., in., 
p. 263 (1887). 


Shillong and Cherra Punji; common. 


The inner band of forewings is acutely bent in on to 
the costa, but this species can more easily be distin- 
guished from ewxtersaria by having the discal band 
waved up to costa near apex, whereas in evtersaria it is 
acutely angled inwards on to the corta at one-third from 
apex. 

Genus Rampara, Moore. 
166, &. efila, sp. n. 

é. White, both wings with a brown ringlet at the end of the 
cells, almost lunular in shape, a brown mark in centre of hinder 
margin of forewings, and a short brown streak before centre of 
abdominal margin of hindwings, giving with the cell marks the 
appearance of a broken medial band across both wings; an 
indistinct discal sinuous brown line on hindwings with slight traces 
of one on the forewings, a large black sub-apical spot on fore- 
wings, and black dots on the outer margin of both wings between 
the veins, Expanse of wings, 1,4; inch. 


Cherra Punji; two examples of this fine Rambara. 


Genus Naxa, Walker. 
167. N. textilis, Walker, vii., p. 1743 (1856), 

Shillong; two examples. 

Orthostixis hiigeli, Felder, Reise Nov. Lep., pl. 130, 
fig. 19 g (1874), is generally made a synonym to the 
above, but this wrong: there are two distinct species, 
although the pattern is very much the same; /hégeli has 
antenne: very much more deeply pectinated than in 
tewtilis. 

Genus CeLteruna, Walker. 
168. C. divisa, Walker, Trans. Ent. Soc. 1862, p. 72. 

Shillong and Cherra Punji. 


Genus, IorapHora, Nov. 


Palpi short, covered ~with short stiff hairs, last joint depressed ; 
antenne of male bipectinate, with short stiff branches to the tips, 
the pectinations ciliated. Female with simple antenne ; hind- 


Lepidoptera of the Khasia Hills. 169 


wings with vein 2 one fourth from lower end of cell, 3 just 

before the end, 5 above the centre of disco-cellulars, rather close 

to the upper end of cell, 6 and 7 from end of cell ; forewings with 

veins 2 and 3 as in hindwings, 5 from centre of disco-cellulars ; 

hind tibiee with two pairs of short spurs. 

169. I. iridicolor (Butler), Ann. Mag. N. H. (5), vi, 
p. 227 (1880). 

Cherra Punji; one pair. 


Genus Tosaura, Swinh. 
170. T. falcipennis (Moore), Deser. Ind. Lep. Atk., iu., 
p. 266, pl. 8, fig. 29 (1887). 
Shillong and Cherra Punji; common. 
Vis Pomecaria, Swinh., Trans. Ent. Soc., 13892;' p. 16; 
pli, fie? 1G: 
Shillong ; many examples. 


Fam. GEOMETRIDAS. 
Genus HuscuEema, Hiib. 
Hazis, Boisd. 
Heleona, Swainson. 
172. HE. militaris (Linn.), Mus. Lud. Ulr., p. 375 (1764). 
Shillong. 
173. H. excubitor, Moore, P. Z. 8., 1878, p. 846. 
Shillong. 


Genus Pacayopes, Guén. 
174. P. hemataria (Herr Schiiff.), Exot. Schm., figs. 2065, 
206 (1850-69). 
P. ornataria, Moore, Descr. Ind. Lep. Atk., in., 
p. 249 (1887). 
Common. 
175. P. vigens (Butler), Ann. Mag. N. H. (5), vi., 
p. 127 (1880). 
Common. 
176. P. apicalis (Moore), Descr. Ind. Lep. Atk., iii., 
p. 247 (1887). . 
Common. 


170 Colonel Charles Swinhoe on the 


177. P. costistrigaria (Moore), P. Z. 8., 1867, p. 633. 
Shillong and Cherra Punji. 
178. P. similis (Moore), Descr. Ind. Lep. Atk., ii. 
p. 248 (1887). 
P. ruficosta, Hampson, Ill. Typ. Lep. Het. B. M., viu., 
p. 108, pl. 150, fig. 16 (1891). 
Shillong. 
179. P. erionoma, Swinh., Ann. Mag. N. H. (6), xu., 
p- 219 (1893). 
Shillong; numerous examples. 


Genus ABSALA, Swinh. 
180. A. doreada, Swinh., Ann. Mag. N. H. (6), xii, 
p. 149. (Pl: Tie...) 
Shillong; three males, one female. 


Genus Psrupotrerpna, Hiibn. 
Hypochroma, Guén. 
Dindica, Moore. 
Pingasa, Moore. 
181. P. polyphenaria (Guén.), Phal., i., p. 280 (1857). 
Shillong and Cherra Punji; common. 
182. P. para (Swinh.), Trans. Ent. Soc. 1891, p. 490. 
Shillong; many examples. 
183. P.crenaria (Guén.), Phal., i., p. 278 (1857). 
Hypochroma distenta, Walker, xxi., p. 434 (1860). 
H. sublimbata, Butler, Ann. Mag. N. H. (5), x., p. 232 
(1882). 
Shillong and Cherra Punji. 
184. P.chlora (Cram.), Pap. Exot., iv., p. 233, pl. 398, 
fig. c (1782). 
Hypochroma lariaria, Walker, xxi., p. 433 (1860). 
H. irrorataria, Moore, P. Z. S., 1867, p. 632. | 
Shillong and Cherra Punji; very common. 
185. P. rufofasciata (Moore), Descr. Ind. Lep. Atk., i., 
p. 247 (1887). 
Shillong and Cherra Punji. 


Lepidoptera of the Khasia Hills. 171 


186. P. ruginaria (Guén.), Phal., i., p. 278 (1857). 
Hypochroma perfectaria, Walker, xxi., p. 454 (1860). 
H. nyctemerata, Walker, l.c., p. AA, 

Shillong and Cherra Punji; very common. 
187. P. alba (Swinh.), Trans. Ent. Soc. 1891, p. 491, 
ple LO tices; 
Shillong ; several males. 


Genus CuLoroponroperA, Warren. 
188. C. discospilota (Moore), P. Z. 8., 1867, p. 621. 
Shillong. 
189. C. chalybeata (Moore), P. Z. 8., 1872, p. 580, pl. 34, 
fig. 4. 
Shillong and Cherra Punji. 


Genus Hprpristis, Meyrick. 
Terpnidia, Warren. 


190. E. minimaria (Guén.), Phal., i., p. 279 (1857). 
Hypochroma parvula, Walker, xxi., p. 435 (1860). 
Acidalia truncataria, Walker, xxiii., p. 774 (1861). 
Shillong and Cherra Punji. 


Genus Hrrocaroma, Swinh. 
Actenochroma, Warren. 
191. H. baba, Swinh., Ann. Mag. N. H. (6), xu, p. 148 
(1893). 

Shillong; many specimens. 

Mr. Warren’s paper on Indian Geometers in Coll. 
Elwes was published in the same month as mine in the 
Annals, but Part ii. of P. Z. 8S. was not issued from the 
printers until three weeks later than the issue of the 
August number of the Annals and Magazine of Nat. 
Hist. in which my paper appeared. 


192. H. subtepens (Walker), xxi., p. 438 (1860). 
Shillong; numerous examples. 


The type ijn the O.M. came from Sarawak, it is, how- 
ever, a common species in the Khasia Hills, and the 
Khasia Hill examples are identical with the type. 


— 
“TI 
Lo 


Colonel Charles Swinhoe on the 


198. H. viridaria (Moore), P. Z.8., 1867, p. 632. 
Shillong and Cherra Punji ; common. 


194, H. muscicoloraria (Walker), xxvi., p. 1543 (1862). 
Shillong and Cherra Punji. 


Genus Maxartrs, Moore. 
195. M. celataria (Walker), xxii., p. 552 (1861). 
Shillong. 


196. M. macariata (Walker), xxvi., p. 1562 (1862). 
Shillong. 


Genus Lissonica, Nov. 


Differs from Maxates in the simple antenne in both sexes; in 
the genus Mazates the antenne in the male is bipectinate for two- 
thirds its length, in the veination also vein 2 of the hindwings 
arises much nearer the end of the cell. 

197. L. polygrapharia (Walker), xxi., p. 435 (1860). 

Macaria vagata, Walker, xxii., p.927 (1861). 

Shillong ; common. 


Genus EprsoTHatma, Swinhoe. 

198. H. sisunaga (Walker), xxii., p. 550 (1861). 
Thalassodes macruraria, Walker, xxvi., p. 1561 (1862). 
Shillong; nine specimens. 

199. H. ocellata, Swinh., Ann. Mag. N. H. (6), xii., p..218 

(1893). (PIN, Ger os) 
Shillong and Cherra Punji; two examples. 


Genus Cacamopa, Nov. 


d. Antenne simple, hindwing produced into a tail ; veins 6 and 
7 stalked at end of cell, 5 emitted very near 6, forewing with vein 
3 emitted before end of cell, 6 and 7 stalked before end of cell. 


200. C. viridata (Moore), P. Z. 8., 1867, p. 625. 
Shillong. : 


Lepidoptera of the Khasia Hills. 173 


“Genus AGATHIA, Guén. 
201. A. lycenaria (Kollar), Hiig. Kasch., iv., p. 486 
(1848). . 
Common. 
202. A. prasina, Swinh., Ann. Mag. N. Ele) xe, 
p. 219 (1893). 
Shillong; three males. 
203. A. letata (Fabr.), Ent. Syst., iii., 2, 164 (1793). 
Shillong; four specimens. 
204. A. quinaria, Moore, P. Z. 8., 1867, p. 639. 
Cherra Punji and Shillong ; many examples. 


Put with hilarata, Guén., in the B.M. collection, but 
is certainly distinct; hilarata only differs from lcetata in 
the medial transverse band of forewings: lxtata, hilarata, 
and quinaria are closely allied, but the three forms 
appear constant without intermediates. I have many 
examples of each from different parts of India. 


205. A. gemma, Swinh., Trans. Ent. Soc., 1892, p. 8. 
Shillong ; five examples. 

206. A. hemithearia, Guén., Phal, i., p. 381 (1857). 
Shillong. 


207.: A. beata, Butler, Ann. Mag. N. H. (5), vi., p. 218 


(1880). 
Shillong. 


208. A. arcuata, Moore, P. Z. 8., 1867, p. 64. 
Shillong and Cherra Punji; very common. 
209. A. codina, Swinh., Trans. Ent. Soc., 1892, p. 7, 
pled igs: 
Shillong ; four males of this fine species. 
Genus Tanaoruinus, Butler. 
210. T. viridiluteata (Walker), xxii., p. 515 (1861). 


Shillong and Cherra Punji; apparently a common 
species in the Khasias. 


174 Colonel Charles Swinhoe on the 


211. T. reciprocata (Walker), xxii, p. 515 (1861). 
Geometra dimissa, Walker, l.c., p. 516. 
Shillong; two examples. 
212. T. kina, Swinh., Ann. Mag. N. H. (6), xu, p. 150 
(1893), (PL Tl Mie 1.) 
Shillong; four males. 
Genus AporanpriA, Noy. 


Antenne of male deeply pectinated for two-thirds its length, the 
pectinations ciliated ; palpi upturned, thickly covered with hair, 
last joint short, tip square ; hindwings with veins 3 and 4 and 
6 and 7 forked one-third their distance beyond the cell, vein 2 
emitted less than half the distance from the base to the fork of 
veins 3 and 4 ; forewing with vein 3 emitted a short distance before 
cell, vein 2 from the middle, vein 6 one-fifth beyond cell ; hind 
tibiz with two short terminal spurs. 


213. A. specularia (Guén.), Phal., 1., p. 342 (1857). 
Shillong. 


Genus Loxocurta, Butler. 
214. ZL. variegata, Butler, Ill. Typ. Lep. Het. B. M., 


vii., p. 104, pl. 136, fig. 3 (1889). 
Shillong. 
Genus Mrcatocutora, Meyrick. 
215. MM. vittata (Moore), P. Z.S., 1867, p. 636. 
Shillong. 
216. M. avicularia (Guén.), Phal., i., p. 342 (1857). 
Shillong. 
Genus Eucroris, Hiibn. 
Thalassodes, Guén. 
217. H. quadraria (Guén.), Phal., i., p. 8360 (1857). 
Thalassodes inconclusaria, Walker, xxii., p. 556 (1861). 
Shillone. 


218. EH. lunifera (Moore), Descr. Ind. Lep. Atk., i., 
p. 270 (1887). 
Type Cherra Punji in Coll. Staudinger: I have not 
seen this species. 


Lepidoptera of the Khasia Hills. 175 


219, H, opalina (Butler), Ann. Mag. N. H. (5), vi., p. 214 
(1880). 
Shillong and Cherra Punji. 


Very closely allied to the preceding, but uniformly 
larger. 
220, H. liliana (Swinh.), Trans. Ent. Soc., 1892, p. 7, 
pl. 1, fig. 2! 
Shillong. 
221. H. acte, Swinh., l.c., p. 6, pl. 1, fig. 13. 
Shillong. 
222. H. thalassica (Moore), Lep. Ceylon, ni, p. 430, 
pl. 194, fig. 6 (1387). 
Cherra Punji. 
223. H. devexata (¢) (Walker), xxii., p. 573 (1861). 
Agathia scintiligera (¢), Butler, Ann. Mag. N.H. (5), 
vi., p. 216 (1880). 
Shillong. 


Genus THALERouRA, Nov. 


Cell of hindwings larger than in Euchloris, vein 7 emitted 
nearer its upper end, antennz bipectinate for three-fourths from 
base, the pectinations evenly ciliated. 


224, T. goniaria (Felder), Reise Nov. Lep., pl. 128, 
fig. 3 2 (1874), 

Cherra Punji. 

225. T. urapteraria (Walker), xxxv., p. 1608 (1266), 

Shillong; two examples. 


Genus Getasma, Warren. 
226. G. thetydaria (Guén.), Phal., 1., p. 8358 (1857), 
Thalassodes bifasciata, Walker, xxvi., p. 1562 (1862), 
Shillong. 


176 Colonel Charles Swinhoe on the 


227. G, griseoviridis, Warren, P. Z, S., 1898, p. 353, 
pl. 31, fig. 6. 
Cherra Punji. 


Genus CGinospina, Swinhoe. 
228. Ci. lyra, Swinh., Trans. Ent. Soc., 1892, p. 5. 
Shillong and Cherra Punji; many specimens. 
229. GH. strix (Butler), Ill, Typ. Lep. Het. B. M., vii, 
p. 105, pl. 136, fig. 8 (1889), 
Cherra Punji. 


Genus Nemortia, Hiibn. 
Hemithea, Dup. 


930. N.insularia (Guén.), Phal.,i., p. 315 (1857). 
Thalera insularia, Hampson, Ill. Typ. Lep. Het. B. M., 
Will., 20, 771 (L891). 
Shillong and Cherra Punji. 


The Cherra Punji examples are much larger than 
those from Shillong, but there is no specific difference 
between them. 


Genus T'satera, Hiibn. 
231. T. disjuncta, Walker, xxii., p. 595 (1861). 
Shillong. 
232. T. graminea, Hampson, Ill. Typ. Lep. Het. B. M., 
viul., p, 109; pl. lol; fig, 1 (1891), 
Cherra Punji. 


Genus Eucrosrzs, Hiibn. 
Comibena, Hubn. 
Uliocnemis, Warren. 


233. H. sanquilineata (Moore), P. Z.8., 1867, p. 638. 
Cherra Punji. 

234, H. partita (4) (Walker), xxi, p. 573 (1861). 
Comibena felicitata, (9), Walker, /.c., p. 579. 
Thalera concisiplaga (¢), Walker, l.c., p, 598, 
Cherra Punji. 


Lepidoptera of the Khasia Hills. 177 


235, LH. integranota (Hampson), Ill. Typ. Lep. Het. B. M., 
ix., p/ 146, pl £70; fe. 13. 


Cherra Punji. 
236. I. kafebera, sp. n. 


d White, forewings with a green transverse sub-basal band, a 
broader discal band, and between these a large green spot separated 
from the bands by a fine white line ; this discal band is continued 
on the hindwings as a medial band, bent outwards in the middle ; 
both wings with a submarginal macular green band, the spots 
formed by the white veins and a white outer festoon; these bands 
have between them white bands, through the centre of which runs 
a fine sinuous green line, outer margin and cilia green, the veins white 
and white streaks in the cilia opposite the veins ; thorax green, 
abdomen with green segmental band ; underside white. Expanse 
of wings, 12; inch. 


Cherra Punji; two examples. 


I feel uncertain as to the genus of this insect, and 
as Mr. Hampson is now working out the genera of the 
Geomeiridz, I prefer leaving it to him to determine. 


Genus Berta, Walker. 
237. B. chrysolineata, Walker, xxvi., p. 1621 (1862), 
Shillong. 


Genus rmanpra, Dup. 
238. T. aventiara (Guen.), Phal. ii, p. 3 (1857). 
Shillong. 


239. T. convectaria (Walker), xxiu., p. 800 (1861). 
Timandra comptaria, Walker, xxvi., p. 1615 (1862). 
Shillong and Cherra Punji. 

240. T'. responsaria (Moore), Descr. Ind. Lep. Atk., iii., 

p- 255 (1887). 
Shillong and Cherra Punji. 
TRANS. ENT.S0C. LOND. 1894.—PaRT I, (MARCH.) Mt 


Wie! Colonel Charles Swinhoe on the 


Fam. ACIDALIDA. 
Genus Pertxera, Meyrick. 
241. P. obrinaria (Guén.), Phal., i., p. 414 (1857). 
Shillong. 
242, P. absconditaria (Walker), xxvi., p. 1580 (1861). 
Common. 


Genus Musorropnr, Hampson. 
243. M. expunctaria (Walker), Journ. Linn. Soc., ii., 
p. 195 (1859). 
A, ovisignata, Moore, Lep. Ceylon, iii., p. 444 (1887). 
Shillong. 


Genus T'rrracHopopA, Hampson. 
3 


244, 1. obstataria (Walker), xxi., p. 769 (1861). 
Anisodés subroseata, Walker, xxvi., p. 1582 (1862). 
A. acuta, Moore, Lep. Ceylon, iii., p. 445, pl. 215, 
fig. 9 (1887). 
Shillong. 


Genus ANnrisoDEs, Guén, 
245. A, lichenaria, Swinh., Trans. Ent. Soc., 1892, p. 8. 

Shillong. 

246. A. intermietaria, Swinh., lc. 

Shillong. 

247. A. pulverentula, Swinh., L.c., p. 9, pl. 1, fig. 8. 

Shillong. 

248. A. heydena, sp. n. 

3. Yellowish straw colour, upperside of palpi black ; wings 
irrorated with brown atoms, forewings with a very large chocolate- 
brown oval patch in the centre, a suffused, sinuous short band 
running upwards from it, hindwings with a similar central smaller 


spot, pale in its centre, a sub-basal transverse brown line, an ante- 
medial thin brown band touching the inner side of the spot, and a 


Lepidoptera of the Khasia Hills. 179 


discal brown line running evenly with the outer margin across 
both wings ; on forewings there are two small suffused brown 
patches on the margin, one at the apex and the other below it ; 
marginal points black, one on each vein and one between each 
vein. Expanse of wings, 1,4; inch, 


Cherra Punji; one example. 


249, A. dotilla, sp. n. 


3 Dark pinkish-grey, wings densely irrorated with minute 
brown atoms, a white dot at the end of each cell, those on the 
hindwings ringed with black, both wings with a discal row of black 
dots on the veins and a similar row of marginal dots, cilia pale 
grey. Underside body and wings suffused with pink, a discal 
sinuous pale brown indistinct line with pale black points across 
both wings, marginal line brown, legs pink, abdomen grey. 


Expanse of wings, 1,3; inch. 
Cherra Punji; one example. 


Genus Dirnatma, Meyrick. 


250. D. ideea (Swinhoe), Trans. Ent. Soc. 1892, p. 6. 
Shillong and Cherra Punji. 


251. D. crina (Swinhoe), l.c., p. 14, pl. 1, fig. 14. 
Shillong. 


252. D. albivertew (Swinhoe), l.c., p. 15. 
Shillong and Cherra Punji. 


253. D. hampsoni (Swinhoe), l.c., p. 14. 
Shillone. 


254, D. mecysma, sp. n. 

gd @. Dark mouse colour, wings very minutely irrorated with 
grey, an indistinct black dot at the end of each cell, both wings 
crossed by ante-medial and post-medial nearly straight brown 
lines, the latter a little outwardly bent below the middle on the 
hindwings, forewings with a discal similar line, and both wings 
with brown marginal lines, cilia grey, underside grey, lines as 
above. Expanse of wings, 2; inch. 


Cherra Punji; numerous specimens, 


180 Colonel Charles Swinhoe on the 


Genus EritHroLopnus, Swinh., 
Organopoda, Hampson. 


255. EH. fascicorpus, Swinh., Trans. Ent. Soc. 1892, p. 11. 
Shillong. 
256. H. carnearia (Walker), xxii., p. 644 (1861). 


Shillong. The cell spot in both wings is very variable 
in size, and sometimes in the forewings it is obsolete. 


Genus SyneciopEs, Swinb. 


257. S. diffusifascia, Swinh., Trans. Ent. Soc. 1891, 
p: Jd; pl: 1) fig. 9;and Ann Maco NE (oiS) sr, 
p. 150 (1893). 


Common. 


258. S. histrionaria, Swinh., Trans. Ent. Soc. 1891, p. 12. 
Common, 


Genus STREPTOPTERON, Swinh. 
259. S. posticamplum, Swinh., l.c., p. 10, pl. 1, fig. 12. 
Shillong. The type specimen is the only example 
received of this extraordinary insect. 
Genus CHrysocrAsPeDA, Warren. 
260. C. abhadraca (Walker), xxii., p. 630 (1861). 
Cherra Punji. 
Genus AcIpALIA, Treit. 
261. A. walkeri, Butler, P. Z. 8., 1883, p. 170. 
A. extimaria, Walker, xxii., p. 794, No. 280, nec. 
782, No. 252 (1861). 
Shillong and Cherra Punji. 
262. A. opsinaria, Swinh., Trans. Ent. Soc. 1892, p. 15. 
Shillong. 
263. A. similaria (Moore), P. Z,.S., 1867, p. 841. 
Shillong, 


Lepidoptera of the Khasia Hills. 181 


264. A. attentata, Walker, xxii., p. 754 (1861). 
Shillong. 

265. A. patularia, Walker, xxxv., p. 1633 (1866). 
Shillong. 

266. A. remotata, Guén., Phal., i., p. 488 (1857). 
Shillong. 

267. <A. nesciaria, Walker, xxii., p. 750 (1861). 
A. negataria, Walker, xxii., p. 701. 
Cherra Punji. 

268. A. fibulata,Guen., Phal.,i., p. 490, pl. 15, fig. 5 (1857). 
Cherra Punji. 


Genus CraspepiaA, Hiibn. 
269. C. pallivittata (Moore), P. Z. S., 1867, p. 641. 
Shillong and Cherra Punji. 


270. C. arenosaria (Moore), Lep. Ceylon, ui., p. 445 
a) 


(1887) 
Shillong. 
271. CO. addictaria (Walker), xxii., p. 749 (1861). 
Cherra Punji. 


Genus RHopostropHiA, Hiibner. 
Phyletis, Guén. 
Delocharis, Butler. 


272. R. stigmatica, Butler, Ill. Typ. Lep. Het. B. M., vii, 
p- 110, pl. 1386, figs. 19, 20 (1889). 
Shillong. 
273. R. prasonaria (Swinh.), Trans. Ent. Soc., 1892, 
p. 12. 
Tanaotrichia trilineata, Warren, P. Z. 8., 1893, p. 361, 
pl. 32, fig. 2. 
Shillong. 
274. R. pelloniaria (Guén.), Phal., ii., p. 169 (1857). 


Shillong and Cherra Punji; common. 


182 Colonel Charles Swinhoe on the 


275 R. khasiana (Moore), Desc. Ind. Lep. Atk., iu., 
p. 264 (1887). 
Cherra Punji; a very distinct species. 


276. R. similata, Moore, Desc. Ind. Lep. Atk., 11., p. 264 
(1887). 
The type came from the Khasia Hills, not in my 
collection. 


This genus links the Acidulide with the Geometride, 
vein 5 in both wings being emitted above the centre of 
discocellulars. 


‘Genus Hyrta, Steph. 
277. H. ferruginaria (Moore), P. Z. 8., 1867, p. 654. 


278. H. marginata, sp. n. 

d. Rosy grey, minutely irrorated with rosy red atoms, fore- 
wings with a black mark at the end of the cell, both wings crossed 
by indistinct sinuous red lines, ante-medial, medial, and post- 
medial, the first obsolete on hindwing, the last with brown points 
on the veins, costa of forewings and marginal line on both wings 
dark reddish brown, thick, and very prominent, cilia rosy. Ex- 
panse of wings, ,’; inch. 


Cherra Punji. 


Genus Deroa, Swinh. 
Lipomelia, Warren. 


279. D. ustata, Swinh., Ann. Mag. N. H. (6), xii., p. 151 
(1893). 


Inpomelia subusta, Warren, P. Z. 8., 18938, p. 360, 
pl. 32, fig. 24. : 


Shillong and Cherra Punji. 


Genus Somatina, Guén. 


280. S. rosacea, sp. n. 

dg. Front white, collar and abdomen rosy, thorax white, abdo- 
men with darker segmental bands; forewings greyish white with 
the interior portions suffused with rosy, a black point at end of 


Lepidoptera of the Khasia Hills. 185 


cell, a patch of darker rosy grey in the disc, limited outwardly by 
a pale whitish sinuated line which commences on the-hinder mar- 
gin near the angle, and curves inwards round the top of the cell. 
On the inner side of this line there is a brownish suffusion, a sub- 
marginal grey band arising frem a small black patch near the hinder 
angle, and curving inwards in to the centre where it is double, 
leaving the apex white, marginal line black, slightly lunular ; hind- 
Wings rosy, a more distinct black point at the end of cell, a sub- 
marginal pale sinuous line, inwardly broadly edged with brownish ; 
a marginal white band with the ground colour running into it in 
parts, and a black marginal line as in forewings, cilia of both 
wings white, interlined with rosy grey. Expanse of wings, 
1,3, inch, 
Cherra Punji; two examples. 


Genus Runeca, Moore. 
281. R. ferrilineata, Moore, Desc. Ind. Lep. Atk., iu., 
p. 202, pl. 8, ig. 13 (1887). 
Cherra Punji. 


Genus Prosiepsis, Led. 
Argyris, Guén. 

282. P. apollinaria (Guén.), Phal., ii., p. 13 (1857). 

Shillong and Cherra Punji. 

With the discal ocelli of forewings very large and 
quite round. 
283. P. delaria, Walker, xxiui., p. 808 (1861). 

Shillong. 

Very doubtfully distinct from apollinaria. 


284, P. delphiaria, Guén., Phal., ii., p. 14 (1857). 
P. vulgaris, Butler, Il. Typ. Lep. Het. B. M., vii., 
p. 43, pl. 125, fig. 2 (1889). 
Shillong. 


285. P. extrusaria, Walker, xxii., p. 637 (1861). 
Cherra Punji. 


184 Colonel Charles Swinhoe on the 


Genus Atpx, Walker. 
Orgalmia, Walker. 


Gamoruna, Moore. 
286. A. palparia (Walker), xxiii., p. 988 (1861). 
A, nigrozonata, Walker, xxvi., p. 1754 (1862). 
Gamoruna palparia, Moore, Descr. Ind. Lep. Atk., 11., 
p. 265 (1887). 
Shillong and Cherra Punji; common. 


Genus Groarapa, Nov. 


Closely resembles Alex in appearance, but in the male 
the antenne is serrate; in Alew it is bi-pectinate. The 
cell is longer, vein 5 of hindwings is free; in Alex it is 
stalked with 6, 


287. G. helweola, n. sp. 

gd. Olive-brown, without the red colour always to be found in 
A, palparia, the transverse band and other markings very similar. 
Sometimes there is a large blackish-brown spot on forewings near 
the hinder angle, but this is often absent ; the underside, however, 
is very distinctive ; the bands are more like lines, and well defined ; 
the medial band is farther away from the base in the hindwings 
than in Alex, and is well curved ; the sub-marginal band in a semi- 
dentated line, not a diffuse band ; and on the forewings there is an 
inwardly curved thin band inside the middle band, and rising from 
its lower end, runs up towards the costa. Expanse of wings, 
2 inches. 


Shillong and Cherra Punji. Many examples, but not 
nearly so common as Alex, the type female of which in 
the B. M. has been carefully examined. 


Genus T'rycopss, Guén. 
288. T. divisaria, Walker, xxiii., p. 927 (1861). 
Shillong and Cherra Punji. 


Fam. LARENTIIDA. 
Genus GonanticLea, Swinh. 
289. G. aversa, Swinh., Trans. Ent. Soc., 1892, p. 4. 
Shillong ; common. 


Lepidoptera of the Khasia Hills. 185 


290. G. occlusata (Felder), Reise Nov. Lep., pl. 182, 
fig. 16 (1874). 
Shillong and Cherra Punji. 


Genus CoLix, Guén. 
291. C. hypospitata, Guén., Phal., ii., p. 358 (1857). 
Shillong. 
292. C. ghosha,; Walker, xxiv., p. 249 (1862). 
Shillong. 


Genus Remopes, Guén. 
293. R. abnormis, Moore, Descr. Ind. Lep. Atk., iii. 
p. 2/1, pl Soll ese): 
Cherra Punji. 
294. R. decussata (Moore), P. Z.S., 1867, p. 655, pl. 33, 
fio. 10. 
Cherra Punji. 
295. R. fasciata, Moore, l.c., p. 270. 
The type came from Cherra Punji. 
296. R. mterruptaria, Moore, l.c. 
Recorded from the Khasia Hills. 
297. R. lineosa, Moore, Deser. Ind. Lep. Atk., iii., 
p. 270, pl. 8610 (1887). 
Shillong. 


Genus Carice, Walker. 


298. OC. rachiaria, Swinh., Trans. Ent. Soc. 1891, p. 492. 
Shillong and Cherra Punji. 


299. C. lunulineata, Moore, Descr. Ind. Lep. Atk., i., 
p. 263, pl. 8, fig. 26 (1887). 
Cherra Punji. 
300. C. zalska, sp. n. 

d- Ochreous grey, striated with brown ; forewings with the 
costa marked with black and pale spots, crossed by an inner in- 
distinct and incomplete brownish band, a discal, sinuous brownish 
line, double in parts, and with two or three square black spots in 
patches on its outer side above the hinder angle, where there are 
also some black and white striations marking the wing, and 


186 Colonel Charles Swinhoe on the 


extending in a band-like form up the outer margin ; hindwings 
with the outer portion darkly and thickly striated with brown, 
underside paler, with black striations and band on the outer por- 
tions of both wings. Expanse of wings, ;5; inch. 

Cherra Punji. 

A very obscure-looking little insect, allied to nothing 
I know of; received in great numbers. 


Genus PuipaLapreryx, Steph. 
301. P. plurilineata, Moore, Descer. Ind. Lep. Atk., 111., 
p. 273 (1887). 
P. nigrovittata, Warren, P. Z. §., 1888, p. 327. 
P. nigripunctata, Warren, l.c., p. 328. 
Shillong. Identical with my Thundiani examples. 


Genus ZrripAva, Walker. 
302. Z. eylinearta, Walker, xxvi., p. 1550 (1862). 


Menophra (?) rubridisca, Hampson, Ill. Typ. Lep. Het. 
Be M., vin, p. £06, p)ito0; fie.3 (169): 


Shillong ; common. 


Genus Arponis, Moore. 
303. A. chlorophilata (Walker), xxvi., p. 1768 (1862). 
Shillong. 
Genus KurirHecta, Curt. 
304. H. felicata, Swinh., Trans. Ent. Soc. 1892, p. 1. 
Shillong and Cherra Punji; many examples. 
305. H. infestata, Swinh., P. Z. 8., 1889, p. 430. 
Cherra Punji. 


306. H. dentifascia, Hampson, Il. Typ. Lep. Het. B. M., 
vili., p. 172, pl. 152, fig, 12 (189), 
Shillong. 
807. H. rigida, Swinh., Trans, Ent. Soc. 1892, p. 2. 
Shillong. 
308. H. wmfestata, Swinh., P. Z. 8., 1889, p. 430. 
Cherra Punji. 


Lepidoptera of the Khasia Hills. 187 


309. H. atroviridis, Warren, P. Z.S., 1893, p. 384, pl. 31, 
fig. 19 
Shillong. : 
310. H. griseipennis, Moore, Descr. Ind. Lep. Atk., 
p. 269, pl. 1, fig. 28 (1887). 
Cherra Punji. 
dll. H. incurvata, Moore, l.c., p. 268. 
The type came from the Khasias. 
312. H. rufifascia, Hampson, Ill. Typ. Lep. Het. B. M., 
ix Pp: [ods pls iil, ties to (E693). 
Cherra Punji. 


Genus Lygranoa, Butler. 
313. L. rectilineata, Moore, Descr. Ind. Lep. Atk., ., 
p- 272 (1887). 
The type came from the Khasia Hills. 


Genus Barpanss, Moore. 
314. B. plicata, Moore, Descr. Ind. Lep. Atk., i., p. 251, 
pl. 8, fig. 22 (1887). 
Shillong. 


Genus Lozornora, Steph. 
315. L. decorata, Moore, l.c., p. 272. 
Shillong and Cherra Punji. 
316. L. pulcherrima; Swinh., Ann. Mag. N. H. (6), xu, 
p. 224 (1893). 
Shillong. 


Genus DysretTuia, Warren. 


317. D. ocyptaria (Swinh.), Ann. Mag. N. H. (6), xii, 
p. 157 (1893). 
Shillong. 


Genus Campoaia, Guén. 


318. C. pulchella (Hampson), Ill. Typ. Lep. Het. B. M., 
vui., p. 124, pl. 158, fig. 22 (1891). 
Cherra Bai 


188 Colonel Charles Swinhoe on the 


Genus LacintopzEs, Nov. 


Antenne simple in both sexes ; palpi depressed, short, covered 
with hairs; hindwing with the lower portion of cell much 
extended, vein 3 emitted from the end, 2 at one third before end, 
5a little above middle of disco-cellular, 6 and 7 forked at nearly 
half the distance beyond upper end of cell ; forewing with vein 3 
emitted a little before end of cell, 2at one third before end, 5 a 
little above centre of disco-cellular, 6 and 7 at upper end of cell; 
mid-tibia with one pair of terminal spurs ; hind-tibia with two 
pairs. 

319. L. plurilinearia (Moore), P. Z. 8., 1867, p. 645. 

Shillong and Cherra Punji. 


There are two forms of this insect, one being without 
any of the usual dark markings on the wings. 


Genus Piemyria, Hub. 


9 


320. P. catenaria (Moore), P. Z. 8., 1867, p. 655, pl. 33, 
fig, 9. 
Shillong. 


Genus PonynzstA, Swinh. 


321. P. truncapeax, Swinh., Trans. Ent. Soc. 1892, p. 5. 
Shillong. 


Genus Cryprotosa, Warren. 


322. O. aerata (Moore), P. Z. S., 1867, p. 654. 
Shillong. 


Genus XanrHoRHOE, Hub. 
Coremia, Guén. 


323. X. conturbata (Walker), xxvi., p. 1703 (1862). 
Shillong. 


Genus Erirruor, Hib. 


324. H. exliturata (Walker), xxiv., p. 1195 (1862). 
Shillong. 


Lepidoptera of the Khasia Hills. 189 


Genus Crpartia, Treit. 


325. CO. relata, Butler, Ann. Mag. N. H. (5), vi., p, 229 


(1880), 
Shillong. 
326. C. aliena, Butler, l.c., p. 230. 
Shillong. 


327. O, fecunda, Swinh., Trans, Ent, Soc, 1891, p, 493, 
Shillong and Cherra Punji, 


328. C. scortea, Swinh,, l.c, 
Shillong, 


329, C, delecta, Butler, Aun. Mag. N. H. (5), vi., p. 229 
(1880). 


Shillong ; common. 


330, OC. curewmata, Moore, Descr. Ind. Lep. Atk., iti, 
p. 278 (1887). 


Shillong and Cherra Punji. 


33l, O. decurrens, Moore, Descr. Ind. Lep, Atk., iii, 
p. 276 (1887). 
Cherra Punji. 


332. O. bareconia, sp. n. 

¢ 2. Palpi, antenne, head and body black; forewings with 
the basal half dark blackish-brown, with its outer edge somewhat 
sinuous and bent slightly outwards above the middle, crossed in its 
centre bya pale thin band which has a thin black line in it, is 
sometimes indistinct and sometimes obsolete, a transverse brown 
line close to the outer edge of the basal half followed by a 
pale pinkish-grey band, and then a number of transverse lines 
close together right up to the outer margin, where there is a black 
patch near the hinder angle, another below the apex, and a third 
in the costa near the apex ; sometimes the outer lines are obsolete, 
and the outer portions of the wings grey with the black patches 
showing more prominently ; hindwings dark brownish-grey, with 
indistinct transverse grey bands, which however, are often 
obsolete ; underside dark grey, crossed by indistinct central and 


discal darker bands. Expanse of wings, ,°;—,° inch, 


Cherra Punji; many examples, 


190 Colonel Charles Swinhoe on the 


333. C. suwbstituta, Walker, xxxv., p. 1691 (1866). 
Shillong. 

334. C. chalybearia, Moore, P. Z. 8., 1867, p. 663. 
Shillong. 

335. C. cervinaria, Moore, l.c., p. 664. 
Shillong. 

336. OC. nivicincta, Butler, Il. Typ. Lep. Het. B. M., vii., 

p. 119; pl: 137, fe 200839). 

Shillong. 

337. C. viridata, Moore, P. Z. 8., 1867, p. 661. 
Shillong. 

338. C. aurata, Moore, l.c., p. 664. 
Shillong. 


Genus ParaLopuiaA, Warren. 
339. P. pustulata, Warren, P. Z.8., 1898, p. 371, pl. 30, 


foe. 
Cherra Punji. 


Genus Eustroma, Hiibn. 
340. E. dissecta, Moore, Lep. Ceylon, iii., p. 488, pl. 207, 
he Oe (ser): 
Shillong and Cherra Punji. 
341. H. triangulifera, Moore, Descr. Ind. Lep. Atk., i., 
p. 275 (1887). 

Shillong ; common. . 
342, H. obscurata, Moore, P. Z. §., 1867, p. 633, pl. 33, 
ieee 

Shillong ; very common. 
343. H. furva (Swinh.), Trans. Ent. Soc. 1891, p. 494. 
Shillong and Cherra Punji. 
344. H. dentifera, Moore, Descr. Ind. Lep. Atk., ii,, 
p. 275 (1887). 
The type came from Cherra Punji, 


Lepidoptera of the Khasia Hills. 191 


345, EE monana, Swinh., Ann. Mag. N. H. (6), xii, 
p. 224 (1893).° (PI. II, fig. 10.) 


Shillong. 
346. H. multifaria (Swinh.), P.Z. 8., 1889, p. 429, pl. 44, 
fig. 9. 


Cherra Punji. 


347. H. aurigena (Butler), Ann. Mage. N. H. (5), vi., 
p. 280 (1880). 
Cherra Punji. 


Genus Scorosia, Steph. 


348. S. sordidata, Moore, Descr. Ind. Lep. Atk., i1., 
p. 274 (1887). 
Shillong and Cherra Punji. 


Genus Ganparitis, Moore. 
349, G. flavata, Moore, P. Z. 8., 1867, p. 660. 
Cherra Punji. 


Genus Psoroscorosta, Warren. 
350. P. miniosata, Walker, xxv., p. 1354 (1862), 
Shillong. 


Genus ANTHyYRIA, Swinh, 
301. A. grataria (Walker), xxii, p. 663 (1861). 
Shillong. 
302. A. ole, Swinh., Trans. Ent. Soc. 1892, p. 2. 
Shillong. 
303. A. lunulosa (Moore), Lep. Ceylon, i., p. 450, 
pl. 200, figs. 5, 5a (1887). 
Shillong and Cherra Punji. 


Genus Pomasta, Guén. 
354. P. moniliata, Warren, P. Z.8., 1893, p. 367, pl. 32, 


fie. 9 
Type, Khasia Hills, in coll. Elwes, 


192 Colonel Charles Swinhoe on the 


Genus AstHENna, Hiibn. 
355. A. ochracea, Warren, P. Z. S., 1888, p. 231. 


A. ochracea, Elwes, P. Z. 8., 1893, p. 264, pl. 32, 
fic, 10 ¢. 


Recorded from Shillong. 


Genus PsitocAmpocia, Hampson. 


356. P. rapistriaria (Swinh.), Trans. Ent. Soc. 1890, 
p: 210, spl Gees 
Shillong. 
307. P. deviaria (Walker), xxu., p. 664 (1861). 
Shillong. 


Genus Raopomerra, Meyrick. 


358. R. sacraria (Linn.), Nat. i., 2, 863, 220 (1766). 
Cherra Punji. 


Fam. BOARMIID/AA. 
Section I. URAPTERYGIN Ai. 


Genus Urapteryx, Leach. 


309. U. multistrigaria, Walker, xxxy., p. 1535 (1866). 
Shillong and Cherra Punji. 


360. U. sciticaudaria, Walker, xxvi., p. 1480 (1862). 
Shillong. 


361. U. picticaudata, Walker, xx., p. 12 (1860). 
Shillong and Cherra Punji. 
This species, the type of which is in O.M. from Borneo, 


is quite distinct from the yellow form; sciticaudaria is 
uniformly much larger, and with different markings. 


362. U. primularis, Butler, I]. Typ. Lep. Het. B. M., vi., 
p. 49, pl. 113, fig. 4 (1886), 
Shillong. 


Lepidoptera of the Khasia Hills. 193 


Genus T'ainopreryx, Butler. 
303. T. crocopterata (Kollar), Kasch von Hiigel, iv., 
p. 453 (1848). 
Shillong and Cherra Punji; very common. 
364. T. pretoraria (Felder), Reise Noy. Lep., pl. 122, 
fig. 13, g (1893). 
Shillong. 
360. T. nebulosa, Butler, Journ, Linn. Soc., 1883, p. 203. 
Shillong and Cherra Punji. Quite distinct from the 
preceding. I have a fine series of both of them without 
any intermediates. 
Genus XEropreryx, Butler. 


366. X. columbicola (Walker), xx., p. 11 (1860). 
Shillong. 
Genus Sremopreryx, Butler. 
367. S. rufivinctata (Walker), xxvi., p. 1747 (1862). 
Shillong and Cherra Punji; very common. 
Section II, DEILINIINA. 
Genus Barra, Steph. 
368. B. margarita (Moore), P. Z. 8., 1867, p. 647. 
Corycia alba, Moore, Descr. Ind. Lep. Atk., il., p. 261 
(1887). 
Shillong. 
369. B. platylewcata (Walker), xxxv., p. 1628 (1866). 
Cherra Punji. 
370. B. indistincta (Moore), Descr. Ind. Lep. Atk., iii., 
p-. 261 (1887). 


Shillong and Cherra Punji. 
371. B.imamata (Walker), xxil., p. 755 (1861). 
Shillong and Cherra Punji. 
Genus Derninta, Hiibn. 
372. D. luteifrons, sp. n. (PI. II., fig. 14.) 


@. Pure white ; front luteous ; wings crossed by three indis- 
tinct dentated grey lines ; ante-medial, medial, and discal at even 
TRANS, ENT, SoC, LOND. 1894.—ParT I. (MARCH.) N 


194, Colonel Charles Swinhoe on the 


distances apart ; fore legs with ochreous tibie ; hind legs with 
brown bands on the tarsi. Expanse of wings, 13 inch. 


Cherra Punji. Two examples. 


Genus Perera, Herr Schiiff. 
378. P. medardaria, Herr Schiff, Exot. Schm., pl. 94, 
fig. 534 (1856). 
Shillong and Cherra Punji. 
374, P.fasciata (Moore), P. Z. 8., 1867, p. 634, pl. 32, fig. 8. 
Shillong. 
5. P. riobearva (Walker), xx., p. 129 (1860). 
Shillong and Cherra Punji; common. 
376. P. capitata (Walker), xxiii., p. 1019 (1861). 
Alana rubiginata, Walker, xxxv., p. 1568 (1866). 
Cherra Punji. 


377. P. albopunctata (Swinh.), Trans. Ent. Soc., 1891, 


or 
ol 


p. 491 
Shillong. 
378. P. cervinaria Pa: Descr. Ind. Lep. Atk., i1., 
p. 233 (1887). 


Cherra Punji. 


Genus Tasta, Walker. 
Dissophthalmus, Butler. 
379. T. micaceata, Walker, xxvi., p. 1570 (1862). 
Dissophthalmus iridis, Butler, Ann. Mag. N. H. (5), 
vi, p. 219 (1880). 
Cherra Punji. 


Genus Myrreta, Walker. 
Orthocabera, Butler. 
Microniodes, Hampson. 
380. M. planaria, Walker, xxili., p. 831 (1861). 
Shillong. 
381. M. ocernaria (Swinh.), Ann. Mag. N. H. (6), xii, 
p. Lo2 (1693). (Plank, die.7s;) 
Shillong, 


Lepidoptera of the Khasia Hills. 195 


882. M. brunneiceps (Warren), P. Z. S., 1893, p. 387, 
pl. 31, fig. 23. 
Shillong and Cherra Punji. 


Genus CABrropEs, Guen. 
383. C. achromaria, Guén., Phal.i., p. 136 (1857). 

Shillong and Cherra Punji. 

384. C. costalis (Moore), P. Z. 8., 1867, p. 616. 

Shillong. 

Genus Hererosrecant, Hampson. 
385. H. subtessellata (Walker), xxvi., p. 1648 (1862). 

Shillong, Cherra Punji. 

386. H. lala (Swinh.), Trans. Ent. Soc., 1892, p. 13. 

Shillong. 

Genus Srecanra, Guén. 
387. iS. latifasciata (Moore), Descr. Ind. Lep. Atk., iii, 
p. 200 (1887). 
Cherra Punji. 
388. S. peralba, sp.n. (Pl. IL., fig. 5.) 

g 2. Of a uniform white colour, tinged with grey. Above 
there are very few markings. Costal line of forewings flesh colour, 
marked with black; and on the hind wings the cell spot and black 
band of the underside is slightly visible ; on the underside there is 
a minute black dot at the end of cell on forewings ; and on the 
hindwings the black spot and black discal band are very promi- 
nent ; legs flesh-colour. Expanse of wings, ¢ 3%, 2 1, inch. 

Cherra Punji; two males, one female. 


Genus Micronissa, Swinh. 


889. M. dephinaria, Swinh., Ann. Mag. N. H. (6), xii, 
p. 152 (1893). 
Shillong. 


Genus Piuropss, Guén. 


390. P. flavescens, Butler, Ann. Mag. N. H. (5), vi, 
p. 223 (1880). 


Shillong ;.common. 


196 Colonel Charles Swinhoe on the 


391. P. exquisata, Butler, Ann. Mag. N. H. (5), vi, 
p. 223 (1880). 


Shillong ; a fine series, 


392. P. subcaudata, Butler, l.c., p. 224. 
Shillong; many examples. 


393. P. prasina, Swinh., Trans. Ent. Soc., 1892, p. 13, 
pl i, fe. 10: 


Shillong and Cherra Punji. 


394. P.costatus, Butler, Ill. Typ. Lep. Het. B. M., vi., 
p- 93, pl. 114, fig. 4 (1886). 


P. triangularis, Warren, P. Z.8., 1893, p. 388. 
Shillong and Cherra Punji; very common. 


This is a variable species, and Mr. Warren could 
not have had a sufficient number before him when de- 
scribing triangularis; none of the points of difference 
mentioned by him hold good. I have examples from 
Solon, near Simla, and a long series from Shillong and 
Cherra Punji. The costal band of forewimgs varies in 
width, the triangular central tooth varies in size; in one 
example it is continued in the form of a yellow bar right 
across the wing to the hinder angle, and similarly the 
antemedial transverse line in some examples runs into 
the yellow band just before the first tooth, but in others 
it runs into the tooth. 


395. P. lamisca, sp. n. 


¢. Bright ochreous yellow, thorax (except the collar) and 
abdomen chocolate-brown ; wings patched and marked with the 
same colour ; forewings with a patch at base extending also down 
the abdominal margin of hindwings, as in all the species of the 
Cyclaria group ; a large discal patch on forewings with dentated 
margin all round, and joined to the basal patch by an irregular 
band ; on hindwings two bands of small spots, discal and sub- 
marginal, with a diffuse patch between them below the middle. 
Expanse of wings, 1,3; inch. 


Cherra Punji; three examples, 


Lepidoptera of the Khasta Hills. 197 
Section III. ENNOMINA. 


Genus Dauima, Moore. 
Panisala, Moore. 
Metoxidia, Butler. 
Hololoma, Warren. 
396. D. intricata, Warren, P. Z. 8., 1893, p. 396. 
Shillong. 
397. D. patnaria, Felder, Reise Nov. Lep., pl. 122, 
fir 12,4 (E873), 
Shillong ; common. 
398. D. truncataria (Moore), P. Z. 8., 1867, p. 620. 
Cherra Punji. 
399. D. patularia (Walker), xx., p. 247 (1860). 
Shillong and Cherra Punji. A very variable species, I 


have them in all colours, from dark purplish brown to 
pale yellowish grey. 
400. D. calaminia (Butler), Ann. Mag. N. H. (5), vi, 
p. 121 (1880). 
Shillong; many examples. 


Genus Hypocurosis, Guén. 
Omiza, Walker. 
Marcala, Walker. 
Celenna, Walker. 
Phenix, Butler. 


401. H. pachiaria (Walker), xx., p. 247 (1860). 
Shillong and Cherra Punji. 

402. H. hyadaria (Guén.), Phal. ii., p. 587 (1857). 
Marcala ignivorata, Walker, xxvi., p. 1764 (1862). 
Shillong and Cherra Punji. 

403. H. venusa, sp. n. 
$- Of a uniform dark pink brown, with a greenish tinge, 

antenne black, forewings with the costal line ochreous, dotted 

with brown, a dark pink upright line or thin band from the hinder 
margin before the middle, which bifurcates at the end of the cell, 


the outer branch outwardly elbowed before reaching the costa 
hindwing with the costal space pale pinkish, underside pale 


198 Colonel Charles Swinhoe on the 


ochreous red, unmarked, darkest on forewings with the hinder 
marginal space of that wing whitish. Expanse of wings, 1,%; inch. 

Cherra Punji; one example. 

The general colouration above is somewhat similar to 
that of H. hyadaria, Guén.=ignivorata, Walk., but the 
markings are different to any other species of this genus 
yet recorded. I have very fine series of all the different 
species of Hypochrosts mentioned in this paper; all the 
females are more or less like each other, but can easily be 
separated ; the males are very distinct, and I have never 
seen any intermediates. 

404. H. abstractaria (Walker), xxxv., p. 1559 (1866). 

Marcala irrorata, Moore, Descr. Ind. Lep. Atk., ii1., 

p. 282 (1887). 

Shillong and Cherra Punji; common. A somewhat 
variable insect; many of the examples have black 
patches on the hinder margin near the angle of various 
sizes, some of them very large. 

405, H. obliquaria (Moore), l.c. 

Shillong and Cherra Punji; very common. 

406. H. festivaria (Fabr.), Hnt. Syst., iii, 2, 152, 84 
(1793). 
Geometra saturataria, Walker, xxii., p. 519 (1861). 
Cherra Punji. 
407. H. iris (Butler), Ann. Mag. N. H. (5), yi., p. 122 
(1880), 
Cherra Punji. 
Genus Acurosis, Gueén. | 
408. A. quadraria, Warren, P. Z. 8., 1893, p. 409. 
Cherra Punji. 
Genus Prionia, Hiibner. 
Osicerda, Walker. 
Oclesdera, Walker. 
Ismisca, Walker. 
Zomia, Moore. 
409. P. incitata (Walker), xxiv., p. 1085 (1862). 
Zomia miscella, Swinh., Trans. Ent. Soc., 1891, p. 144. 
Shillong and Cherra Punji. 


Lepidoptera of the Khasia Hills. 199 


410. P. squalidaria, Hiibner, Zutr., iv., p. 40, figs. 787, 
788, 9 (1882). 
P. squalidaria, Cotes and Swinh., Cat. Moths of India, 
iv., No. 3691 (1888). 

Ismisca cyclogonata, Walker, xxvi., p. 1765 (1862). 
Shillong ; a fine series. 

411. P. costimaculata (Moore), P. Z. 8., 1867, p. 650. 
Shillong. 


412. P. trinotaria (Moore), lc. 
Cherra Punji; one example, much’smaller than usual. 
413. P. intexta (Swinh.), Trans. Ent. Soc., 1892, p. 145, 
; pl 3; fos ie. 
Shillong. 
Genus Herpromiza, Warren. 
414. H. flava (Moore), Descr. Ind. Lep. Atk,, iii., p. 233, 
Shillong. pl. 8, fig. 5 (1887). 
415. H. sanguiflua (Moore), l.c., fig. 4. 
Shillong. 
416. H. cruentaria (Moore), P. Z. 8., 1867, p. 616. 
Cherra Punji. 


Genus Napacara, Walker. 
417. N. inordinata, Walker, xxiv., p. 1094 (1862). 
Shillong. The type from Sarawak isin the O.M. I 
have received two males from Shillong which are 
identical with Bornean and Maylayan specimens. 


Genus Hinicrinra, Hubner. 
418. E. flava (Moore), Descr. Ind. Lep. Atk., i1., p, 233, 
pl. 8, fig. 2 (1887). 
Shillong. 
Genus SpitopeRA, Warren. 
Hrinnis, Warren. 
419. S. umbrata, Warren, P. Z.S., 1893, p. 403. 
Cherra Punji. 
Closely allied to S. gracilis, Butler, from Japan, 
uniformly smaller, and differing in the disposition of the 
bands. 


200 Colonel Charles Swinhoe on the 


420. S. hepaticata, sp. n. 

¢. Chocolate-brown, wings smeared with whitish in parts, a 
black dot at the end of each cell, forewings with a straight 
transverse brown line one-third from base, an irregular sinuous 
brown line from costa of forewings near apex to hinder margin, 
one-fourth from the angle, continued on the hindwings, where 
it is discal and curved, corresponding to the curves of the outer 
margin, the portion of the wings outside this line is darker and 
suffused towards the apex of forewings with a red tint, and there 
is a white smear on the outer margin below the apex and another 
smear in the disc below it. Underside paler, uniform in colour, 
suffused with grey, the cell dots distinct and both wings crossed 
by a discal sinuous chocolate line. Expanse of wings, 1,5; inch. 


Cherra Punji; two examples. 


421. S. combusta, Warren, P. Z.8., 1893, p. 415. 
Cherra Punji. 
422. 8. obliquilinea (Moore), Descr. Ind. Lep. Atk., iii., 
p. 229 (1887). 
Leptomiza straminea, Warren, P. Z. S., 1898, 
p. 407, 9. 
Shillong and Cherra Punji. 


The type of Warren’s genus Leptomiza is calcearia, 
Wk. Of the three new species he describes in the 
same paper all are from female types, and it is impossible 
to tell what genus they really belong to until the males 
are examined ; out of one hundred and seventy-four new 
species described in this paper, no less than fifty-four 
are erected on female types, and of many of them he 
appears to have had only single specimens before him. 


Genus Fascrentina, Walker. 

423. F. chromotaria, Walker, xx., p. 215, ? (1860). 
Geometra usta, Walker, xxxv., p. 1602, ¢ (1866). 
Shillong and Cherra Punji; common. 

424, F. plagiata (Walker), xxxv., p. 1601 (1866). 

F. viridis, Moore, P. Z. 8., 1867, p. 79, pl. 7, fig. 4. 
Shillong. 


Lepidoptera of the Khasia Hills. 201 


425. F. subsignata, Warren, P.Z.S., 1893, p. 399. 


F. curtaca, Swinh., Ann. Mag. N. H. (6) xu, p. 220 
(1893). 


Shillong. 


426. F. dacoda, Swinh., l.c., p. 221. (Pl. I1.,.fig. 7.) 
Shillong and Cherra Punji. 


427. F. cyanifera, sp.n. (Pl. IL, fig. 13.) 

d ¢. Chocolate-brown suffused with ochreous, and smeared 
and striated in parts with white ; forewings with two indistinct 
brownish transverse straight bands, first one-third from base, 
second in the middle ; a thin white line, edged with brown on both 
sides, and more or less sinuous, extending from a white sub- 
apical patch on costa of forewings to hinder margin, one-fifth from 
the angle, and continued across the disc of the hindwings, where 
it is bent at its centre ; inside this line on the hindwings is a broad 
chocolate-brown band, and outside it on both wings are several 
white smears ; on the underside the body, legs, and wings are 
ochreous, wings striated with brown, suffused with white and yellow 
on forewings, uniformly bright ochreous on hindwings; on the 
forewings, a white line edged with brown runs from the costa of 
forewings where it is deeply elbowed outwards, and. then nearly 
straight down to the hinder angle, one-third from the angle, a broad 
reddish-brown band on its innerside, narrowing downwards, and all 
the wing outside the line of the same colour and smeared with white 
in parts ; the hindwings are crossed by two discal brown lines, the 
inner one bent in its middle. Expanse of wing, ¢ 1;4;, 2 1-5, inches. 


Cherra Punji; one pair. 


The female only differs from the male in having rounder 
wings. | 


Genus Oxsonozpa, Walker. 
428. O. clelia (9) (Cram.),. Pap. Exot., iii, p. 172, 
pl. 288, figs. B, c. (1780). 
O. rajaca (¢), Walker, xx., p. 219 (1860). 
O. pallida, Butler, Ann. Mag. N. H. (5), vi., p. 125 (1880), 


Cherra Punji. 


202 Colonel Charles Swinhoe on the 


Genus Hyposipra, Guén. 
Lagyra, Walker. 

429. H. talaca (Walker), xx., p. 59, ¢ (1860). 

Shillong. 
430. H. aquilaria (Walker), xxvi., p. 1485 (1862). 

Shillong. 
431. H.umbrosa (Swinh.), Trans. Ent. Soc. 1890, p. 205, 

pl. 7, fig. 4. 
Macaria aquilaria (nom preoc.), Walker, xxvi., 
p. 1562 (1862). 
Shillong. 
Genus Hyatinetra, Nov. 

¢. Differs from Hyposidra in having ciliated antenne, 
not pectinated as in that genus; veins 3 and 6 of both 
wings emitted from the ends of the cells, the cells being 
closed by a hyaline mark bent outwards in the middle. 
432. H. megaspila (Moore), P. Z. §., 1867, p. 616. 
Shillong. 

Genus Garmus, Moore. 

3. G. discolor, Warren, P. Z.S., 1893, p. 400. 


Genus Psyra, Walker. 
434. P. cuneata, Walker, xxi., p. 485 (1860). 
Shillong and Cherra Punji. 


Genus CrocaLus, Treit. 
Odontopera, Steph. 
Corotia, Moore. 
Niphonissa, Butler. 


435. C. translineata (Walker), Char. Undescr. Lep. Het., 
p- 98 (1869). 
Odontopera nemea, Swinh., Trans. Ent. Soc., 1891, 
p. 485, pl. 19, fig. 5. 
Shillong. 
Walker’s type is in the Devon and Exeter Museum, 
and Mr. Warren, who has seen it, says it is identical 


with my type of nemea; Walker described his species as 
a Deltoid of the Genus Selenis. © 


Lepidoptera of the Khasia Hills. 205 


436. C. cervinaria (Moore), P. Z. 8., 1867, p. 625, 
pl. 32, fig. 10. 

Shillong. 

437. C. decorata (Moore), l.c., p. 621, pl. 82, fig. 9. 

Shillong. 

438. C. codra (Swinh.), Trans. Ent. Soc., 1891, p. 486, 
pl. 19) fig. 3. 

Shillong. 

439. C. murina (Swinh.), l.c., p. 487, pl. 19, fig. 13. 

Shillong. 

410. C. heydena, sp. nu. 

S$. Dark ochreous grey, forewings densely irrorated with 
minute dark greyish-brown atoms, a brown dot at the end of the 
cell, indications of an inner brownish outwardly-curved band at 
one-third from the base, a nearly straight blackish-brown thick line 
from the costa near apex to the hinder margin one-fourth from the 
angle, edged with whitish on the outer side,-this line slightly curves 
inwards below its centre ; black points in the interspaces close to the 
outer margin, cilia brown with pale tips, hindwings pale ochreous- 
grey, unmarked except for a pale grey dot at end of cell. Under- 
side pale ochreous-grey, a blackish spot at end of each cell, and the 
discal line showing through the wing. Expanse of wings, 1,75 inch. 

Shillong ; many examples. 

Allied to C. bilineata, Swinh., which, however, is a 
larger insect, has a discal band on hindwings and has 
two teeth below its semi-falcated apex of forewing, 
a character entirely wanting in all the specimens I have 
yet seen of this species, the outer margin of the forewing 
being produced above the middle and evenly concave 
between that and the apex. 


Genus Merrocampa, Latreille. 
441. M. haliaria (Walker), xxii., p. 518 (1861). 
Shillong and Cherra Punji. 


Genus Herrronocua, Led. 
442, H. patalata, Felder, Reise Nov. Lep., pl. 152, fig. 9 
(1874). 
Marcala varians, Swinh., Trans. Ent. Soc., 1891, 
p: 487. 
Shillong. 


904 Colonel Charles Swinhoe on the 


Genus Crocorta, Hiibn. 
443. C.niguzaria, Walker, xx., p. 129 (1860). 


Shillong; many examples. 


Genus ANTHYPERYTHRA, Swinhoe. 


444, A. hemearia, Swinh., Trans. Ent. Soc., 1891, p. 485, 
pl. oy ges: 
Shillong. 


Genus Hypgeryruea, Guér. 
445, H. lutea, Cram., Pap. Exot., iv., pl. 370 figs. c, D. 
(1781). | 
Common. 


Genus PseuperyrHra, Nov. 
Differs from Hyperythra in the absence of the sexual 


patch of androconia on forewing, the cell being longer in 
both sexes, 


446. P. pheeniz (Swinh.), Trans. Ent. Soc., 1890, p. 489. 
Shillong ; a long series. 


447, P.khasiana (Swinh.), Ann. Mag. N. H. (6), xi, 
p. 221 (1898). 

Shillong; many examples: the females of the above 
two species are rather difficult to distinguish from the 
females of Hyperythra lutea; they have, however, large 
purplish spots all along the outer side of the discal band 
on both wings, whereas in lutea these spots are duller 
coloured and more restricted ; the cell also is shorter in 
lutea : from each other they can be separated by the 
difference in the colouration of the underside. 


Genus ScarDAMIA, Guén. 
448. S. metallarvia, Guén., Phal., i., p. 89 (1857). 
Shillong and Cherra Panji. 


Genus PreratoruyGa, Nov. 


Antenne of male with paired ciliations to the tips ; of the female 
with very minute ciliations ; palpi upturned, covered with hairs ; 
hindwings with vein 4 curving downwards before end of cell, 2 
emitted from middle of cell, 5 and 6 from the ends, forewing with 


Lepidoptera of the Khasia Hills. 205 


vein 3 from end of cell, 5 emitted from 6 beyond end of cell, 
mid tibia with two terminal spurs, hind tibia with four. 
449, P. xrata (Moore), P. Z. 8., 1867, p. 643. 

Shillong, 


Genus Hyruuia, Nov. 

Q. Antenne dilate, palpi upturned, last joint short, depressed, 
smooth ; hindwings with vein 3 emitted before end of cell, 6 at 
the end, 2 beyond the middle ; forewing with vein 3 before end 
of cell, 5 in the centre of discocellulars, 6 bent at its origin before 
upper end; hind tibiz with two pairs spurs, one long and one short 
in each pair. 

450. S. dirempta (Walker), xxii., p. 595 (1861). 

Shillong and Cherra Punji: I have this species from 
several parts of India and from Singapore, all females, 
and have never seen a male. 


Genus PrricaLuia, Stephens. 
451. P. gynopteridia (Butler), Ann. Mag. N. H. (5), vi., 
p. 123 (1880). 
Shillong and Cherra Punji; many examples of both 
sexes. 
452. P. angulifascia (Moore), P. Z. 8., 1878, p. 851, 
pl. os tie, ie 
Shillong ; several males and one female: In this species 
vein 6 of the hindwings is emitted a little further inwards 
from the end of the cell then in gynopteridia, and the 
antenne of the female has the pectinations very short and 
invisible to the naked eye. 


Genus Hprone, Dup. 
453. H, adustata, Moore, l.c., pl. 8, fig. 20. 
Type Khasia Hills in coll. Staudinger: not known 
to me. 


Genus Synzara, Guén. 
Borbacha, Moore. 
Parasynegia, Warren. 
454. S. pardaria (Guén.), Phal., 1, p. 420 (1857). 
Shillong, 


206 Colonel Charles Swinhoe on the 


455. S.camptogrammaria (Guén.), Phal., i., p. 420 (1857). 
Caberodes infixaria, Walker, xxvi., p. 1513 (1862). 
Shillong and Cherra Punji. 

456. S. diffusaria (Moore), P. Z. 8., 1867, p. 641. 
Anisodes punctifera, Butler, Ann. Mag. N. H. (5), vi., 

p. 220 (1880). 7 
Cherra Punji. 

457. S. pluristriaria (Walker), xxvi., p. 1581 (1862). 
Shillong and Cherra Punji; common. 

458, Synegia gopterana, sp. n. 
gd. Ochreous, sparsely irrorated with red, thorax with a red 


band in front ; abdomen with segmental bands, wings with a black 
dot at the end of each cell, hindwings with a short basal red band, 
and a corresponding thick red band near the base of the abdomen, 
both wings crossed by a red band from the centre of the abdominal 
margin of hindwings, where it is double, to the apex of forewings, 
where it joins a band which curves from the apex to the costal 
third ; on the forewings there is another curved band attached to 
the upper half of the medial band and a red mark against the centre 
of the outer margin, and on the hindwings there are some irregular 
red lines in the disc and a straight red band from the apex to the 
anal angle. Expanse of wings, 1,2; inch. 
Cherra Punji; two examples. 


Allied to S. lidderdalii, Butler. 


Genus Coryaica, Walker. 
459. C. arnearia, Walker, xx., p. 231 (1860). 
Shillong. 
460. C. caustolomaria, Moore, Descr. Ind. Lep. Atk., 
iii., p. 231 (1887). 
Cherra Punji. 


Section IV. MACARIIN i, 
Genus Luxraria, Walker. 
Bithia, Walker. 
461. L. phyllosaria (Walker), xx., p. 82 (1860). 
Li. alfenusaria, Walker, l.c., p. 232, 
Shillong. 


Lepidoptera of the Khasia Hills. 207 


462. L. contigaria (Walker), xxii., p. 754 ¢ (1861). 
Acidalia turpisaria, Walker, xxiii., p. 771 ¢ (1861). 
A. tephrosaria, Moore, P. Z. S., 1867, p-. 643 ¢. 

Very common. 

463. L. subrasata (Walker), xxiii., p. 773 (1861). 
Shillong and Cherra Punji; a fine series. 

Forewings very narrow; underside ochreous-brown with the 
usual markings brown, and with a white patch at apex of fore- 
wing. 

464, L. submonstrata (Walker), xxiu., p. 772 (1861). 
Shillong and Cherra Punji; many examples 
The smallest species of the genus, of a pale grey 

colour. 


465. L. fasciosa, Moore, Descr. Ind. Leap. Atk., iii., 
p. 254 (1887). 


Shillong and Cherra Punji; common. 
466. DL. exclusa (Walker), xxi., p. 320 (1860). 
I. exclusa, Cotes and Swinh., Cat. Moths of India, iv., 
No. 4006 (1888). 
Acidalia inewactata, Walker, xxxv., p. 1600 (1866). 
Shillong. 
467. L, obliquata, ee Descr. Ind. Lep. Atk. jell, 
p. 254 (1887). 
Shillong and Cherra Pani ; Many examples. 


Genus Macartia, Curtis. 
Acadra, Herr Schaff. 
Hvarzia, Walker. 
Azata, Walker. 
Gubaria, Moore. 
Gonodela, Boisd. 
468. M. emersaria (Walker), xxiii., p. 925 (1861), 
Shillong and Cherra Punji; common. 


469. M. oliva, sp.n. (PI. IL, fig. 16.) 
&. Olive grey, with slightly darker irrorations ; wings crossed 
by olive brown lines, all bent in on to the costa of forewings, Ist 


208 Colonel Charles Swinhoe on the 


one third from base, on forewings only, 2nd medial, the bent 
portions near costa only distinguishable, 3rd discal; medial on 
hindwing, there is also a nearly straight line on hindwings, from 
apex to‘anal angle ; a black quadrate patch divided by the veins 
on discal line of forewings above the centre, with some black 
markings near it, and three or four pure white spots below the 
apex ; a brown smear like a transverse band on both wings before 
the middle ; underside with irrorations and bands prominent and 
black. Expanse of wings, 1%; inch. 

Cherra Punji; many examples. 

Allied to M. temeraria, Swinh. ‘The pattern above 
somewhat resembles M. pryeri, Butler, from Japan. 


470. M. temeraria (Swinh.), Trans. Ent. Soc., 1891, 
p- 492. 
Shillong and Cherra Punji; very common. 


A71. M. postvittata Walker), xxiii., p. 759 (1861). 
M, honoraria, Walker, l.c., p. 928. 
M. permotaria, Walker, l.c., p. 929. 
Shillong; three examples. 


472. M. vasudeva, Walker, xxiu., p. 933 (1861). 

Shillong. 

473. M. khasiana, Moore, Descr. Ind. Lep. Ath., iii., 
p. 269 (1887). 

Shillong and Cherra Punji; the common form of the 
sufiata group in the Khasias, but I have never received 
it from any other locality. 

A474, M. azataria (Swinh.), Ann. Mag. N. H. (6), xii., 
p. 154 (1893). 
Shillong ; five males. 


A475. M. apataria (Swinh.), l.c., p. 222. 
Shillong and Cherra Punji; twelve males. 


476. M. metagonaria (Walker), xxvi., p. 1518 (1862). 

Common ; in the B. M. Coll. this is made a synonym of 
S. elvirata, Guén., but elvirata belongs to the suflata 
group. I have it from the Shan States; it is the largest 
species of that group, 


Lepidoptera of the Khasia Hills. 209 


477, M. xanthonora, Walker, xxiii., p. 935 (1861), 
Shillong ; many examples. 


478. M. inchoata, Walker, xxiii., p. 931 (1861). 
M. nora, Walker, l.c., p. 934. 
M. neonora, Walker, l.c. 


Very common ; neonora is from the Philippines, it has 
the central white band very broad, but I have an 
example from the Khasias identical with the type. The 
type specimen of inchoata is a female ; some females of 
this extreme form look very distinct, but among the many 
hundreds in my museum, trom many localities, there are 
all three forms with ali the gradations between them. 


479. M. fasciata (Fabr.), Syst. Ent., p. 629 (1775). 


Shillong; appears to be more rare in the Khasias than 
any other species of this group, only three or four 
examples having been received. 


480. M. acutaria, Walker, Char. Undescr. Lep. Het., 
p. 100 (1869). 
Shillong and Cherra Punji; many examples. There 


are two distinct forms of this species, one much darker 
than the other. 


481. M. perspicuaria, Moore, P. Z. 8., 1867, p. 647. 
Shillong and Cherra Punji; common. 
482. M. odataria (Swinh.), Ann. Mag. N. H. (6), xii, 
p, tot, (0893). ((PlcH., fies 17.) 
Shillong; five examples. 
483. M. trilinearia (Moore), Descr. Ind. Lep. Atk., 111, 
p. 263 (1887). 
Shillong. 
484. M. quadraria (Moore), Lep. Ceylon, it., p. 469, 
pl. 205, fig. 1 (1887). 
Azata ferruginata, Moore, l.c., p. 470, fig. 2. 
Shillong; many examples. 


485. M. subfasciata (Hampson), Ill. Typ. Lep. Het. B. M., 
vii., p. 112, pl. 151, fig. 20: (1891). 
Shillong; nine examples. 
TRANS. ENT. SOC, LOND. 1894,—PparT. I. (MARCH.) 0) 


210 Colonel Charles Swinhoe on the 


Genus T'epHRINA, Guén, 
486. T. atmala, sp. n. 


d. Dark slaty-brown, forewings with the costa tinged with 
ochreous, a large black spot at the end of the cell, three transverse 
black lines antemedial, medial, and discal, the first acutely bent 
outwards below the costa, second curving outward from the costa 
round the discal spot, third curving a little outwards from the 
costa, sinuate, pale edged outwardly and continued across the mid- 
dle of the hindwings ; underside paler, striated with whitish, with 
medial and discal darker thin bands, which on the hindwings are 


antemedial and medial. Expanse of wings, 1,2; inch, 


Cherra Punji; one example. 


Genus CassymA, Guén. 
Huteea, Walker. 
Nustidava, Walker. 

487. C. heteronewrata, Guén., Phal. ii., p. 19 (1857). 
Butea personaria, Walker, xx., p. 217 (1860). 
Nustidava unilineata, Walker, xxvi., p. 1573 (1862). 
Shillong and Cherra Punji. I have it also from Sikkim. 


. 


Genus Krananpa, Moore. 
A488, K. semihyalina, Moore, P. Z. 8., 1867, p. 648. 
Shillong; a fine series. 


Genus ZaMARADA, Moore. 
489. Z. translucida, Moore, Lep. Ceylon, ii, p. 432, 
pl. 197, tie, 2 (be 7). 

Cherra Punji. 

490. Z. cosmiaria, Swinh., Ann. Mag. N. H. (6), xii., 
p. Lbo (1893)5~ (Ply fies 3.) 

Shillong and Cherra Punji. 

491. Z. (2) marginata, Warren, P. Z. §., 1893, p. 388, 
pl. 82, fig, 22. 
Cherra Punji. 


Genus ZrHEBA, Moore. 
492. Z. marginata (Walker), xxxv., p. 1561, 9 (1866). 
Hydata spectabilis, Butler, P. Z, 8., 1877, p. 474, 


Lepidoptera of the Khasia Hills. 211 


Z. aureata, Moore, Descr. Jud. Lep, Atk., iii., p. 263, 
4 (1887). 


Cherra Punji. 


Section V. ABRAXIN AH. 


Zerenine. 
Genus Cistripia, Hiibner. 
Halthia, Mén. 
Obeidia, Walker. 
493. C. nigripars (Swinh.), Trans. Ent. Soc., 1892, p. 16, 
pl toes 
Shillong; one example. 
494, C. lucifera, sp. n. 


d 2. Head and body ochreous, thorax with brown spots, 
abdomen with brown segmental bands, wings white, with a broad 
band on all the margins, the bands are ochreous, spotted with 
brown in some examples, but in others the band is brown marked 
with ochreous. In some examples, especially in the females, the 
bands are macular, much as in the Chinese species, C. tigrata, Guén. 
On the paieiias there is also a brown macular band. Expanse of 
wings, ¢ 2%, 2 3 inches. 

Types, India, ¢ 9 in O. M. 

Shillong; one female. I have it also from Sikkim. 
Differs from C. tigrata in its smaller size, and the colour — 
of the wings being pure white instead of ochreous. 


Genus ParicrEropEs, Warren. 
495. P. tenebraria (Moore), P. Z. S., 1867, p. 653. 
Shillong. 


Genus Icrrropes, Butler. 
496. I. hamiltonia (Swinh.), Trans. Ent. Soc., 1892, p. 17, 
pl.) fel. 
Shillong. 
497. I. lapsariata (Walker), xxiv., p. 1121 (1862). 
Cherra Punji. 
498. I. fasciata, sp.n. (Fl. IL, fig. 1a 9. 
3 @. Pevctus, palpi beneath and shafts of the antenne ochreous, 


plumes and upper side of palpi black, abdomen ochreous with 
black spots, wings white with broad grey transverse bands, one at 


Ab? Colonel Charles Swinhoe on the 


the base where there are some ochreous marks, an incomplete band 
in the middle, obsolescent in the hindwings, followed by a broad 
discal band across both wings, and an almost equally broad marginal 
band with a white sinuous line running down itscentre. Body and 
legs below ochreous. Expanse of wings, 2 inches. 


Cherra Punji. I have it also from Kurseyone. 


Genus CatcuLa, Moore. 
499, C. exanthemata, Moore, Descr. Ind, Lep. Atk., iii., 
p. 266 (1887). 
Shillong. 
Genus Apraxas, Leach. 
500. A. leopardina (Kollar), Hiig. Kasch., iv., p. 490 
(1848). 

A. martaria, Guén., Phal. ii., p. 205 (1857). 

A, intermedia, Warren, P. Z. 8., 1888, p. 324. 

Shillong and Cherra Punji; very common: it is im- 
possible to separate the above three. Amongst the 
hundreds [I have received from the Khasias there are 
typical examples of all three, and very many intergrades ; 
it 18 @ species very variable in both pattern and size. 


501. A. khasiana, Swinh., Trans. Ent. Soc., 1892, p. 17. 

Shillong; five examples. 

Genus Perenia, Guén. 
502. P, felinaria, Guén,, ‘Phal.j i. p.216, pl. 19; fet 
(1857). 

Common; I have two or three curious sports of this 
species from Cherra Punji. 

503. P. belluaria, Guén., l.c., p. 217. 

Very common. 

504. P. guttata, Felder, Reise Nov. Lep., pl. 180, 
fig. 15 ¢ (1874). 

P. submissa, Warren, P. Z. 8., 1893, p. 391. 

Shillong ; three examples. I have it also from Simla 
and Kurseyong, the antenne of the male is quite 
different in this species to that in felinaria or bell- 
waria, having well separated thick spine-like branches 
densely ciliated. 


Lepidoptera of the Khasia Hills. 213 


005. P. coryneta, sp. n. 

g. White, pectus, top of head and thorax ochreous ; abdomen 
grey tinged with ochreous ; the two latter with black spots as in 
Percnia ; forewings ochreous at the base, both wings with spots 
much as in P. felniaria, but there are black streaks at the base of 
veins 3, 4, 5 and 6 of the forewings, and the sub-marginal row of 
spots are in pairs, except those in the 2nd and 8rd interspaces ; 
and these spots being somewhat near the margin they leave a white 
discal band between them, and the central duplex row ; in the hind- 
wing the central band of spots are in pairs in the middle, being 
near the origin of veins 3 and 4, and 6 and 7, these being emitted 
from the lower and upper ends of the cell. Expanse of wings, 
28. inch. 


Fao) 


Shillong; one example. 


Genus Mition14, Walker. 


506. M. pulchrinervis, Felder, Reise Nov. Lep., pl. 104, 
fig. 6 9 (1868). 
M. latwitta, Moore, P. Z. 8., 1872, p. 570. 
Shillong. 


Section VI. BOARMIIN/AK. 


Genus Amratica, Moore. 

507. A. recwrsaria (Walker), xxi., p. 374 2 (1860). 
Boarmia ferrolavata, Walker, xxvi., p. 1536 ¢ (1862). 
B. solivagaria, Walker, xxxv., p. 1586 ¢ (1866). 

A. fortissima, Moore, Descr. Ind. Lep. Atk., iii, 

p. 245 4 (1887). 

Shillong; a fine series. ‘The above is a very good 
instance of the futility of erecting types on female 
examples in the Geometers: the above three females all 
vary inter se, but are all undoubtedly one species, easily 
proved by the examination of a long series of this 
species. 

Genus Eusysa, Hiibn. 
Amphidaris, Treit. 
508. H. regalis (Moore), Descr. Ind. Lep. Atk., iii, 
p. 234 (1887). 
Shillong. 


214. Colonel Charles Swinhoe on the 


Genus Cusi1ats, Moore. 
509. CO. bengalaria (Guén.), Phal.,i., p. 210, pl. 4, fig. 2 
(1857). 
Shillong ; many examples of both sexes. 
510. OC. contectaria (Walker), xxvi., p. 1529 (1862). 
Shillong and Cherra Punji. 
511. C. angulata (Hampson), Ill. Typ. Lep. Het. B. M., 
vill, p: LO7, pl. 150, fie, 7.1300), 
Cherra Punji. 


Genus Buzura, Walker. 
512. B. swppressaria (Guén.), Phal., i., p. 210 (1857). 
B. multipunctaria, Walker, xxvi., p. 1531 (1862). 
B. strigaria, Moore, P. Z. 8., 1879, p. 416. 
Shillong and Cherra Punji; common. 


Genus CuHoropna, Walker. 
613. C. erebusaria, Walker, xxi., p. 314 (1820). 
Shillong and Cherra Punji. 
514. C. metaphearia (Walker), xxvi., p. 1482 (18 62). 
Hrebomorpha semiclusaria, Walker, l.c., p. 15: 
#. pretextata, Felder, Reise Nov. Lep., pl. 196, fig. 16 
(1874). 
Shillong. 
615. OC. testaceata (Moore), P. Z. S., 1867, p. 615. 
Cherra Punji. 
Genus Hresomorrua, Walker. 
Vindusara, Moore. 
616. EH. fulgurita, Walker, xxi., p. 495 (1860). 
EH. xanthosoma, Felder, Reise Nov. Lep., pl. 126, 
fig, 17 (1874). 
Shillong ; many examples. 
617. E. fulguraria, Walker, xxi, p. 495 (1860). 
Shillong; seven examples. 


Lepidoptera of the Khasia Hills. 215 


518. E. compositata (Guén.), Phal. ii., p. 207 (1857). 


Common. 


Genus OprHaLMopns, Guén. 
519. O. herbidaria, Guén., Phal., i., p. 283 (1857). 
O. diwrnaria, Guén., Lc., p. 284. 
Boarmia pertusaria, Felder, Reise Nov. Lep., pl. 125, 
fig. 17 (1874). 
Shillong and Cherra Punji. 
520. O. infusaria, Walker, xxi., p. 448 (1860). 


Shillong and Cherra Punji; very common. 


521. O. pulsarva, Swinh., Trans. Ent. Soc., 1891, p. 489. 
Shillong and Cherra Punji; many examples. 
522. O. cordularia, Swinh., Ann. Mag. N. H. (6), xi.., 
p. 155 (1893). (PI. IT., fig. 4.) 
Shillong and Cherra Punji. 


Genus AmBiycHiA, Guén. 
523. A. angeronaria, Guén., Phal., i., p. 215, pl. 4, fig. 9 
(1857). 
Shillong. 


Genus Exeuxos, Guén. 
524, H. hymenaria, Guén., Phal.,i., p. 285, pl. 16, fig. 4 
(1857). 
A. pardicelata, Walker, xxvi., p. 1544 (1862). 
E. preumbrata, Warren, P. Z. 8., 1893, p. 433 9. 


Shillong ; common. Warren’s species is erected on a 
female from Perak: in the Oxford Museum and in 
my own museum are examples from several parts of 
India, Borneo, Sumatra and Singapore, with many vari- 
ations of females bearing many of the characters 
described by Warren. 


525. EH. scolopaiea, Drury, Exot. Ins., ii, pl. 22 fig. 1, 
App. ii. (1773), | 
Shillong ; common. ° 


916 ' Colonel Charles Swinhoe on thé 


Genus Boarmia, Treit. 

526. B. admissaria, Guén., Phal. i., p. 239 (1857). 

B. pleniferata, Walker, xxvi., p. 1538 (1862). 

Aleis vicina, Moore, Descr. Ind. Lep. Atk., iii., p. 243 

(1887). 

Shillong. A splendid series of this very variable 
species. 
027. B. perspicuata, Moore, P. Z. 8., 1867, p. 630. 

Shillong. 


028. B. ahenaria, Walker, xxi., p. 370, ¢ (1860). 
B. gelidaria, Walker, xxvi., p. 1537, ? (1862). 
Chogada fraterna, Moore, Descr. Ind. Lep. Atk., iii., 
p. 245 (1887). 

Shillong and Cherra Punji; very common; a very 
widely distributed species: I have it from Sylhet, 
Sikkim, Khasias, Andamans, Nilgiris, and Ceylon. 
£29. B. propulsaria, Walker, xxi., p. 385 (1860). 

B. contiquata, Moore, P. Z. 8., 1867, p. 631. 

Shillong and Cherra Punji; common. 


° 


939, B. semiclarata (Walker), xxiv., p. 1029 (1862). 
Shillong ; many examples. 


531. B. semialba (Moore), Descr. Ind. Lep. Atk., i., 
p. 241 (1887). 
Cherra Punji. 


532. B. latifasciata, Warren, P. Z. 8., 1893, p. 427. 
Cherra Punji. 


084, B. ratotaria, sp.n. (PI. IL, fig. 18.) 

¢. Brown; the colour caused by minute brown irrorations 
densely packed on a grey ground; wings with a lunular-shaped 
ringlet at the end of each cell, and crossed by several dark-brown 
lines, 1st one-third from base, and indistinct ; 2nd medial, out- 
wardly dentated, regularly curved on hindwing outside the ringlet, 


Lepidoptera of the Khasia Hills. 217 


double on forewing, the two lines gradually widening on to the 
costa; 3rd discal somewhat close to the margin, more or less 
outwardly dentated, and edged in places with whitish on its outer 
side, marginal sinuous line dark brown; underside pale brown, 
discal ringlets and central thick black transverse line prominent on 
both wings. Expanse of wings, 1,°; inch. 

Shillong and Cherra Punji; many examples. Allied 
to B. gemmaria, Brahm., but differs in the absence of 
the discal marks, evanescent bands above, and band on 
hindwings more elbowed outwards. 


Genus Racotis, Moore. 


535. R. boarniaria (Guén.), Phal. i., p. 282 (1857). 


Shillong and Cherra Punji. There are two forms, one 
darker and smaller than the other. It is a very common 
species in the Khasias. 


Genus XanpramEs, Moore. 
536. X. dholaria, Moore, P. Z. S., 1867, p. 634. 
Shillong and Cherra Punji. 
D837. X. latiferaria (Walker), xxi, p. 445 (1860). 
Shillong. Six males. The type came from N. China, 


but these specimens appear to be identical with the 
type. 


Genus Mepastna, Moore. 
538. M. strivaria (Guén.), Phal. i., p. 217 (1857). 
Shillong. 


539. M. creataria (Guén.), l.c. 

Shillong and Cherra Punji; common. 
540. M. mawraria (Guén.), lc., p. 218. 

Shillong and Cherra Punji. In Cotes and Swinhoe’s 
“ Moths of India,” No. 3264, M. parisnatei (Walker) is 
put as a synonym of the above. But this is not correct ; 
parisnatet is uniformly much larger than mawraria, and 
has a large pale-yellowish spot in the centre of the outer 
margin of the forewings in both sexes, besides other differ- 
ences. I have both sexes from the Chin Hills, and many 
examples of both sexes of mauraria from the Khasias. 


918 Colonel Charles Swinhoe on the 


541. M. nigrovittata (Moore), P. Z. 8., 1867, p. 626. 
Shillong and Cherra Punji. 


Genus Catiocasra, Nov. 


¢. Antennz bipectinated to the tips, pectinations densely 
ciliated ; hindwing with vein 2 emitted from the centre, 3 from a 
little before end of cell, vein 6 from the upper end ; forewing with 
vein 2 from the centre, 3 from before the end of the cell, 6 bent 
upwards beyond the upper end of the cell, emitting vein 5 at a 
little distance beyond ; hind tibia with two pairs of spurs. 


542. C. similis (Moore), Descr. Ind. Lep. Atk., 111, 
p. 235 (1887). 
Shillong. 
543. C. basistrigaria (Moore), P. Z. 8., 1867, p. 626. 


Shillong; one female. 


Genus Psrupancurona, Moore. 
d44. P. separata (Walker), xxi., p. 381 (1860). 
Boarnia retractaria, Walker, I.c., p. 386. 
B. intectaria, Walker, xxvi., p. 1535 (1862). 
Shillong and Cherra Punji; many examples. 
545. P. reparata (Walker), xxi., p. 380 (1860). 
Shillong ; five examples. 


546, P. lectularia (Swinh.), Trans. Ent. Soc., 1891, 
p. 489, pl. 19, fig. 4. 


Shillong; one example. 


Genus Carascra, Hiibn. 
547. C. eolaria (Guén.), Phal., i., p. 294 (1857). 
Shillong and Cherra Punji; common. 


648. C. ventraria (Guén.), Phal., lc. 
Shillong and Cherra Punji; common. 


Genus Caroria, Moore. 
549, C. sublavaria (Guén.), Phal., i., p. 256 (1857), 


Lepidoptera of the Khasia Hills. 219 


CO. olivescens, Moore, Descr. Ind. Lep. Atk., ii., 
p. 244 (1887). 
Shillong and Cherra Punji. A fine series; typical 
examples of sublavaria and olivescens look very distinct, 
but I have many intergrades. 


Genus Darisa, Moore. 


550. D. mucidaria (Walker), xxxv., p. 1581 (1866). 
Shillong and Cherra Punji. 


Genus Lassapa, Moore. 
Hol. DL. albidaria (Walker), xxxv., p. 1582 (1866). 
Shillong. 
Genus Hemeropuira, Steph. 
Menophra, Moore. 
552. H. delineata (Walker), xxi., p. 387, 2 (1860). 
H. canidorsata, Walker, xxxv., p. 1578 (1866). 
Shillong. 
553. H. retractaria, Moore, P. Z.8., 1867, p. 627, pl. 32, 
a 


fig. 
Shillong. 
504. H. humeraria, Moore, /.c., p. 627. 
Shillong. 


555. H. contubernalis (Moore), Descr. Ind. Lep. Atk., iii, 
p. 238 (1887). 
Shillong. 
556. H. vialis (Moore), Descr. Ind. Lep. Atk., i., p. 238, 
pl. 8, fig. 9 (18387). 
Cherra Punji. 
Genus Hrrasa, Moore. 
557. H. scripturaria (Walker), xxxv., p. 1590 (1866). 
Scotopteryx permuscosa, Warren, P. Z.8., 1893, p. 432. 
Cherra Punji. 


920 Colonel Charles Swinhoe on the 


Genus RutreLLerona, Nov. 


Antenne of male with short paired bristles ; hindwing with cell 
open, vein 2 at less than one-third from end, rather close to 3 ; 
forewing with veination normal, vein 5 in centre of disco-cellulars ; 
mid tibia with one pair terminal spurs ; hind tibia with two pairs, 
the inner pair the longer. 

598. R. cessaria (Walker), xxi., p. 383, g¢ (1860). 
Angerona preclaria, Walker, xxvi., p. 1499, ¢ (1862). 
A. pallicostaria, Moore, P. Z. 8., 1867, p. 620. 
Shillong and Cherra Punji; common. 

099. R. figlina (Swinh.), Trans. Ent. Soc., 1890, p. 205, 

pli 7, tig. 0: 

Shillong and Cherra Punji. 


Genus Gyaproma, Nov. 


¢. Antenne fasciculate; palpi upturned, first two joints 
covered with long hairs, last joint short and depressed ; wings long, 
rather narrow ; forewing with costa slightly arched, outer margin 
much rounded; hindwing with outer margin produced and 
scalloped, both wings with veins 3 and 6 emitted before ends of 
cells ; hind tibia thick, with two pairs of short stout spurs and 
fringed with long hairs. 


560. G. testacearia (Moore), P. Z. 8., 1867, p. 623. 
Shillong and Cherra Punji; in great numbers. 


Genus Smrraca, Moore. 
d61. S. transcissa (Walker), xxi., p. 380 (1860). 
Shillong and Cherra Punji. 


Genus Astacupa, Moore. 


562. A. cineracea, Moore, Descr. Ind. Lep. Atk., i., 
p. 244 (1887). 
Shillong. 
563. A. ampla, Moore, l.c. 
Cherra Punji. 


Lepidoptera of the Khasia Hills. 221 


Genus Ecrropris, Hiibn. 
564, H. dentilineata (Moore), P. Z. 8., 1867, p. 631. 
Shillong and Cherra Punji. 


565, EH. pallidaria (Moore), Descr. Ind. Lep. Atk., i, 
> .p.237. (0880) > (RI fies 12.) 
Pscudocoremia dendrellaria, Swinh., Ann. Mag. N. H, 
(6), xi., p. 156 (1893). 
Shillong. 
566. E. pannosaria (Moore), P. Z. 8., 1867, p. 629. 


Shillong. 


567. EH. simplaria, sp. n. 

¢d. Grey, irrorated with black atoms; abdomen with black 
dorsal bands, the last two or three duplex ; wings crossed by black 
sinuous and semi-dentate transverse lines, ante-medial, medial, and 
post-medial, these lines more or less furnished with black points, 
marginal lunules black; cilia yellowish-grey, with black streaks 
opposite the veins. Expanse of wings, 1, inch. 


Cherra Punji. 


568. H. planaria, sp. n. 

¢. Grey, striated with brown; the striations thick on the 
apical portions of the forewings, making them much darker than 
the rest of the wings ; a black lunule at the end of each cell; a 
sinuated aute-medial transverse brown line more distinct on the 
hindwings; another similar medial line, bent outwardly on fore- 
wings before reaching the costa, an indistinct, pale, very sinuous 
discal line and marginal black festoon ; cilia yellowish-grey, with 
dark grey patches; underside grey, with the medial line very 
distinct. Expanse of wings, 1,3; inch. 

Cherra Punji and Shillong. 

Allied to H. ochrifasciata, Moore, Lep. Atk., i., 
p. 240, but quite distinct. 


Genus Ascotis, Hiibn. 

569. A. selenaria (Schiff.), Wien. Verz., p. 101. 
Boarmia reciprocaria, Walker, xxi., p. 366 (1860). 
B. imparata, Walker, l.c., p. 372. 

B. promptaria, Walker, l.c,, p. 379, 


299 Colonel Charles Swinhoe on the 


Ophthalmodes cretacea, Butler, Ann. Mag. N. H., ii., 
p. 373 (1879). 
Shillong and Cherra Punji; common. A very widely- 


spread species occurring in Africa, India, China and 
Japan, 


Genus Anonycata, Warren. 
570. A. grisea, Butler, P. Z. S., 1883, p. 172. 
Shillong and Cherra Punji. 


Genus ARrIcHANNA, Moore. 
571. A. tramesata, Moore, P. Z. 8., 1867, p. 658, pl. 33, 
fig. 2. 
Shillong. 
5/72. A. marginata, Warren, P. Z. S., 1893, p. 423. 
Shillong. 


573. A. transfasciata, Warren, P. Z. S., 1892, p. 425. 
Cherra Punji. 


Genus Scoropreryx, Hiibn. 
O74, 8S. albistellaria, Warren, P. Z. 8., 1893, p. 432. 
Type Khasia Halls in collection Elwes, 


Genus Orspasta, Nov. 


Antenne of both sexes simple; hindwing with vein 4 much 
bent upwards towards end of cell, 2 emitted beyond the middle, 3 
before the end, 6 at the upper end ; forewing with vein 2 at one- 
third before end, 3 before the end of the cell, 5 from just above 
centre of disco-cellular, 6 before upper end of cell ; bind tibia with 
one pair of stout terminal spurs. 


575. O. spurcataria (Walker), xxvi., p. 1498 (1862). 


Shillong ; one female. 


Genus PsiLaucis, Warren. 
576. P. breta, Swinh., P.-Z. 8., 1889, p. 426: 
Shillong ; in Ill. Typ. Lep. Het. B. M., ixs:.p. 36; 


Hampson makes breta a synonym of inceptaria, Walker, 
the type of which isa unique and very much rubbed 


Lepidoptera of the Khasia Hills. 228 


female example from Flores, in the Oxford Museum. 
This is mere conjecture; Flores is a long way from 
India, and the type specimen is too much rubbed for 
identification : it does not even resemble breta, and until 
we can find a male we cannot be certain it even belongs 
to the same genusas breta. On tuisrubbed female Warren 
states he has erected his genus Psilalcis; but he has 
erected his genus on a male, and as the male of tnceptaria 
is not known, that species cannot stand as the type of 
the genus, 


EXPLANATION OF PratE II, 


1. Ocinara cyproba, p. 154. 

2. Eupterote axesta, p. 157. 

3. Zamarada cosmiaria, p. 210, 

4, Opthalmodes corduiaria, p. 215. 

5. Stegania peralba, p. 195. 

6. Absala dorcada, p. 170. 

7. Fascellina dacoda, p. 201. 

8. Myrteta ocernaria, p. 194. 

9. Hpisothalma ocellata, p. 172. 
10. Eustroma monana, p. 191. 
11. Icterodes fasciata, p. 211. 
12, Ectropis pallidaria, p, 221. 
13. Lascellina cyanifera, p. 201. 
14. Deilinia luteifrons, p. 193. 
15. Tanaorhinus kina, p. 174. 
16. Macaria oliva, p. 207. 

17. Macaria odataria, p. 209. 
18, Boarmia ratotaria, p. 216, 


VII. An Entomological Excursion to Corsica. By 
Gzorce C. Cuampion, F.Z.S. 


[Read Feb. 28th, 1894.] 


My friend, Mr. R. 8. Standen, having determined to 
make an entomological expedition to the mountains cf 
Corsica, I, at the last moment, decided to accompany 
him, and, accordingly, we started together on May 28th 
last, arriving at Ajaccio on the morning of May 30th, 
and at the hotel at La Foce de Vizzavona, our proposed 
head-quarters, the same evening. Here we remained 
about a month, being joined later on by Mr. A. H. Jones, 
Colonel Yerbury, R.A., Mr. Lemann, and others, Mr. 
Raine of Hyéres having preceded us by some weeks. 
Altogether, we were quite a large party, all, however, on 
Lepidoptera intent, myself and Colonel Yerbury excepted. 
Mr. Standen has already (Ent. 1893, pp. 236—238, and 
pp. 259—263) given us an account of his experiences 
with the Lepidoptera (Rhopalocera), for most of which we 
appeared to be too early, the season in the mountains 
being very backward compared to the semi-tropical coast 
region. | devoted myself almost exclusively to the 
Coleoptera and Hemiptera-Heteroptera; and it is 
proposed here to give an account of the species met 
with, adding to my list some others (chiefly Longicornia 
and Buprestidz) subsequently obtained by Colonel Yer- 
bury—who remained till August,—which he kindly 
handed over to me on his return. The Hemiptera- 
Heteroptera have been entirely determined by Mr. E. 
Saunders. Our head-quarters at Vizzavona, on the 
summit of the Col between Bocognano and 'Tattone, on 
the main road between Ajaccio and Bastia, at an 
elevation of about 4,000 feet, and commanding a splendid 
view of the lofty Mont d’Oro, proved to be a capital 
centre for collecting. In the immediate vicinity of the 
hotel, though it is not visible till the summit of the Col 
is reached, is a splendid forest, composed almost entirely 
of beech and Corsican pines (Pinus laricio), some of the 
trees being exceedingly lofty, the beech ascending 
TRANS. ENT, SOC. LOND. 1894.—PART I. (MARCH.) P 


226 Mr. George C. Champion on an 


higher up the mountain slopes than the pines. The 
beeches are more or less covered with moss, indicating a 
humid climate, though the coast region is excessively dry 
and arid at this season. The forest entirely fills the head 
of the valley on the northern side of the Col, and in it 
there was an abundance of fallen timber just ready for the 
Coleopterist ; this being partly due to the fact that a 
large number of trees had been felled for the purposes 
of the new railway, which passes in a long tunnel right 
under the forest. Lower down, on either side of the 
Col, to Bocognano on the one side, and Tattone on the 
other, there are plenty of magnificent chestnuts and 
other trees, with green meadows and small patches of 
cultivation. Below Bocognano, right down to Ajaccio, 
the hill-sides are in many places covered with a 
dense bush or “ macchie,” chiefly composed of arbor- 
escent heath, arbutus, evergreen oak, myrtle, and Cistus, 
with, in the opener places,.an abundance of aromatic 
Labiateo and other plants. At Ajaccio itself there is the 
usual semi-tropical vegetation of the Mediterranean 
region, the abundance and thriving condition of the in- 
troduced Opuntias, etc., recalling to my mind visions of 
distant tropical climes. ‘he mountain slopes for some 
distance above the limits of the forest are clothed in 
many places with a dense, almost impenetrable, growth of 
dwarf alder, such as I have seen nowhere else, the alders 
coming into flower directly the snow melts, the ground 
beneath them at this time being bright with crocuses, 
though no gentians were scen. ‘The mountain summits 
are more or less serrated, excessively rugged, and barren, 
and it is not easy to find workable spots, nor, indeed, to 
get about, many of the ridges being impassable without 
along detour. There is also a scarcity of water; even 
in some of the valleys the streams are small, the Gravone 
itself being lost in the marsh and sand before reaching 
the Bay of Ajaccio. In the forest at Vizzavona not 
many flowering plants were to be seen, beyond foxgloves, 
a beautiful Pancratium, an asphodel, a Cyclamen and a 
Sambucus ; but lower down at T'attone, etc., there were 
plenty of Umbelliferze, Orchidacez, Composite, etc. 
Though Monte d’Oro and Monte Retondo were within 
comparatively easy distance of Vizzavona, I did not 
attempt an ascent of either of them, contenting myself 
with working to the summit of the Col between the hotel 
and Ghisoni, to an elevation of a little over 6,000 feet. 


Entomological Hxecursion to Corsica. 227 


From Vizzavona, excursions were made to Tattone, 
Vivario, Corte, Bocognano, Tavera, Ajaccio, etc., the rail- 
way helping a good deal for this purpose ; but most of the 
time was spent in the mountains. At Corte, about half- 
way between Vizzavona and Bastia, in the northern central 
part of the island, to which we made a pleasant three 
days’ excursion, the heat notwithstanding, there is more 
water, the united streams, the Restonica and Tavignano, 
forming a river of considerable size, the banks of which 
were productive in small Coleoptera. Of the cork-oaks, 
of which there are forests in some parts of the island, 
we saw very few in any of the places visited. At 
Bocognano we received a good deal of assistance and 
kind hospitality from Dr. Trotter, this place producing 
many of the species peculiar to the ‘“macchie” and to 
the low country. Still, my list contains but a meagre 
assortment of these latter, it being already much too 
late in the season for most of them, everything being by 
this time parched up near the coast. One thing struck 
me as peculiar, viz., the scarcity of Cryptocephalus ; a 
similar amount of work in the south of I'rance or Spain 
would have produced many species. 


Corsica contains a large number of Coleoptera 
peculiar to itself, upwards of two hundred being given 
in the last European Catalogue (v. Heyden, Reitter, and 
Weise, 1891) from Corsica alone, the Staphylinidee and 
the Rhynchophora each having over thirty peculiar 
species ; and perhaps a still larger number of species are 
common to Corsica and Sardinia, and which do not 
extend on to the Italian mainland. One of the most 
characteristic genera, and one most in evidence, is 
Percus (a close ally of Plerostichus), which is represented 
by several peculiar species, its other members inhabiting 
Italy, Sardinia, Sicily, the Balearic Islands, and Spain. 
One abundant insect in the island, Cetonia awrata, 
affords an instance of the extraordinary amount of 
variation possible within the same species, the different 
varieties occurring together indiscriminately, and quite 
irrespective of a varied geographical distribution, or of 
a difference in altitude of the locality. Two very in- 
teresting papers have already been published on the 
Coleoptera .of the island: (1) ‘‘ Kin entomologischer 
Ausflug in die Berge Siid-Corsica’s,” by G. Dieck (Berl. 
ent. Zeitschr., 1870, pp. 397—404). This exceedingly 


228 Mr. George C. Champion on an 


successful collector paid special attention to the 
Pselaphidee, Scydmzenidee, Dytiscids, etc.; he gives a 
most interesting account of his entomological experi- 
ences in the southern part of the island, where he made 
excursions with M. Reveliére, a well-known Coleopterist 
residing at that time in Corsica, and to whom we are 
largely indebted for our knowledge of its insect-fauna. 
It is to be hoped that some day Dieck’s paper will be 
translated into English and published here, as has 
already been done with his paper on Andalusia (¢f. Ent. 
1888, pp. 38 e¢ seg.). (2), “Notes on some Corsican 
Insects,” by the Rev. 'T. A. Marshall (Hnt. Monthly 
Mag., vii., pp. 225—228). In this paper Mr. Marshall 
gives a most accurate description of the general features 
of the parts of the island he visited, with a list of all 
the Coleoptera he had captured and been able to name; 
but no Dytiscide or Staphylinidee are mentioned, nor 
many of the smaller species. Amongst my captures are 
many species not noticed by either Dieck or Marshall, 
and I propose to give here a list of all the species I have 
been able to identify, adding to my list, as I have stated 
above, some others subsequently obtained by Colonel 
Yerbury, chiefly at Tattone, where a saw-mill was started 
soon after I left. MM. Fauvel and Bedel, the well- 
known French Coleopterists, have kindly assisted me in 
determining many of the more obscure species, the 
former with the Staphylinide, the latter with the 
Carabidee, Curculionids, etc. Representatives of about 
510 species were obtained in all, including a good many 
of the peculiar Corsican forms. 


Before commencing my list, it is perhaps as well to 
give a short account of the general run of species met 
with in the different localities visited. 

At Vizzavona, in the forest, under the bark of the 
decaying beeches, were to be found Hndophlweus 
spinulosus, Coxelus pictus, Brontes planatus, Bolitochara 
lunulata, ete., in plenty; and Colydiwm elongatum, 
[pidia quadrinotata, two species each of Paromalus and 
Jerylon, Litargus bifasciatus, Orchesia undulata, Melan- 
drya caraboides, Melasis buprestoides, Sipalia scabripennis, 
Phleeocharis corsica, Placusa, sp. n. (?), etc., sparingly : 
in the older trees, with looser bark, Pterostichus 
ambiguus and Pristonychus carinatus, both commonly ; 
and in those with plenty of moss and fungi on the bark, 


' Entomological Excursion to Corsica. 939 


Platyrrhinus latirostris, Peltis grossa, Diaperis boleti, 
Triplax rufipes, ete., the first and last-named in numbers. 
Under the bark of the dead pines, especially of those 
still standing, Helops swperbus, a very fine species 
peculiar to the island, and H. ceruleus, Menephilus 
curvipes, Uloma culinaris and U. perroudi, Adelocera 
carbonaria, Hlater preustus and EH. pomonx, and Brachy- 
temnus porcatus, more or less commonly, and Tenebrioides 
maroccanus, Nudobius collaris, ete., sparingly ; on the 
bark of these trees Chalcophora mariana and Dictyoptera 
sanguinea were sometimes to be met with. In recently 
fallen pines Acanthocinus griseus, Pogonocherus perroudi, 
Pissodes notatus, two species of Hrnobius, Salpingus 
nitidus, Helops genei, etc., were to be found on the 
branches ; and in or under the bark, in addition to plenty 
of Scolytidx and the abundant Clerus formicarius, Platy- 
soma oblonguwm, two species of Plegaderus, Nemosoma, 
Aulonium bicolor, two species of Corticeus, Clerus 
quadrimaculatus, Xylita revelicrer, etc. One specimen 
of Lucanus tetraodon was found in a rotten pine stump 
(numerous others were subsequently obtained by Colonel 
Yerbury); and the gigantic larva or pupa of the large 
Prionid, Ergates faber, sometimes occurred in similar 
situations, or under the bark of the lofty standing dead 
pines. ‘Three species of the characteristic Carabideous- 
genus Percus were not uncommon under stones, logs, 
ete., two of them also occurring on the barer mountain 
sides ; a Telephorus (precox) frequented the foxgloves ; 
and Morimus lugubris was abundant all over the place, 
it being especially fond of a heap of cut beech logs near 
the hotel. On the mountain sides a pretty little 
Malacoderm, Malachius longicollis, frequented almost 
every flower; an Asida, a Thylacites, some Harpalt, 
etc., occurred beneath stones, and Meloe brevicollis was 
occasionally found crawling on the grass. Higher up 
in the mountains a Brachyderes (analis*) frequented the 
growing pines; and still higher, on the Col between 
Vizzavona and Ghisoni, Otiorrhynchus guttula swarmed 
on the alder catkins on sunny calm days, and with it a 
single specimen of the rare Ceuthorrhynchus duvalv was 
obtained; under stones in these high places an Asida 
was abundant, and a second Otiorrhynchus (corsicus), not 
uncommon, also one of the species of Percus found lower 


* Originally recorded from Ajaccio, but certainly in error, 


930 Mr. George C. Champion on ait 


down; and on the snow, an Omophlus. In a very 
shallow mountain pool, supplied by the melting snow, 
Hydroporus griseostriatus was plentiful, and on its banks 
were two species of Bembidiwm, also in plenty. 

Below Vizzavona, on the road to Bastia, at 'Tattone, in 
the meadows, Cetonia awrata, showing extraordinary varla- 
tion in colour—black, blue, green, coppery, green with 
coppery thorax and scutellum, etc.,— abounded on flowers, 
and with it were C. floricola and C. morio, T'r iehine 
zonatus, Trichodes, Bostrychus capucinus, Leptura hastata 
and L. fulva, Strangalia bifasciata, Stenopterus rufus, 
Agapanthia cardwi, Clytus rhamni, Clytanthus massi- 
liensis, Bhagonycha Corsica, Malachius sardous, various 
Gidemer «, etc. In the roads, etc., were found a handsome 

variety of Cicindela campestris, not uncommonly, and Calo- 

soma sycophanta, occasionally ; also about stercore, Sisy- 
phus, Ateuchus, Gymnopleurus, Oniticellus, Onthophagus 
schrebert, Geotrupes ¢ geminatus—abundant everywhere, 
even on the higher parts of the mountains where the cattle 
are pastured in summer, —Hister imequalis, H. pustu- 
losus, H. major, H. sinwats, ete. 

By working downwards on the road to Ajaccio, from 
Bocognano to Tavera, representatives of a good many 
interesting species were obtained. In the “ macchie,” 
by beating the arborescent heath, evergreen oak, etc., 
the rare Cleonus tabidus, Caulostrophus delarouzei, 
two species of Nanophyes, Metallites parallelus—one of 
the commonest weevils in the mountains—Simicronyx 
corsicus, Pachybrachys scriptus, Stylosonvus depilis, two 
or three species of Clythra, etc. By sweeping the 
scattered flowering plants (most of which were very 
aromatic) on the slopes and in the meadows lots of 
Coleoptera came to light, as a Livus, two species of 
Cardiophorus, in plenty, an Athous, a Xyletinus, two 
species of Danacea and various other small Malacoderms, 
Throscus corsicus, numerous species of Bruchus, Olibrus, 
and Meligethes, Hadrotoma variegata, Silaria suturalis, 
various CUsdemere, Gymnetron, Anthrenus, Omophlus, 
Anthicus corsicus, Hispa atra, ete. 

At Ajaccio, Pimelia sardoa was abundant all round the 
sandy shores of the bay on both sides of the town, 
though more dead than living examples were seen, 
where also, Calathus, Pedinus, Colpotus, Crypticus, 
Dendavus, Tentyria, Asida, Phaleria, Chrysomela, etc., 
were represented by one or more species; Ammoph- 


Entomological Excursion to Corsica. 231 


thorus rufus abounded at the roots of plants; Capnodis 
tenebricosa flew about in the hot sun; Saprinus semi- 
punctatus abounded in a dead horse, etc. Cetonia morio, 
two species of Anthaxvia, an Acmexodera, Stenopterus 
ater, Phytecia virescens, a Coptocephala, a fine large 
Bruchus (longicornis), Troglops silo, Colotes, etc., oc- 
curred on flowers ; Corebus rubi, commonly on bramble 
bushes; Anthicws corsicus, swarming on a variety of 
bushes and trees on the slopes at the back of the town ; 
Pachypus cornutus (¢), on the wing in the town towards 
evening; Bembidiwm kiistert rarely, and Cicindela 
littoralis commonly, at Campoloro, on the sand at the 
mouth of the Gravone, etc. 

At Corte, at an elevation of about 1,500 feet, some 
interesting small species were to be found on the banks 
of the Tavignano, as Lionychus sturmi, Tachys apristoides, 
Perileptus, the very minute Cryptohypnus meridionalis, 
Anthicus longicollis, two small blue Philonthi, Lathro- 
bium labile, Scopxus, Medon, etc. In the mountain 
ravines, Sisyphus, Ateuchus, Akis punctata, etc., were 
common about stercore on the roads ; and at a trickling 
mossy waterfall two Steni (levigatus and strigosus) 
occurred sparingly. In the meadows and about the 
riverside near the town a good many species already 
met with at Bocognano were to be had by working, 
with the addition of Zonitis mutica and Larinus latus 
on thistle-heads, Malachius rufus on Umbelliferous 
flowers, etc. 


List OF THE SPECIES MET WITH, 50 FAR AS AT PRESENT 
DETERMINED. 


[Those marked with a * are given from Corsica only in 
the Catalogue of v. Heyden, Reitter, and Weise. } 


Cicindela campestris, L., var. connata, Heer, not un- 
commonly on the main road at Vizzavona, Corte, etc.; one 
dark specimen, perhaps belonging to the var. funebris, 
Sturm, Corte; C. littoralis, F., commonly, on the sandy 
beach at the head of the Bay of Ajaccio, at the mouth of 
the River Gravone. 


Carabus morbillosus, F., on the beach at Ajaccio ; 
C. genet, Thoms.,* fragments of a specimen found at 
Vizzavona. Calosoma sycophanta, L., a few specimens, 
Vizzavona. Leistus spinibarbis, F., Vizzavona. Bem- 


332 Mr. George C. Champion on dn 


bidium bipunctatum, Gyll., and B. agile, Duv.,* both 
commonly, on the banks of a shallow pool in the 
mountains, at about 5,500 feet elevation, on the Col 
between Vizzavona and Ghisoni; B. rufescens, Guér., 
Vizzavona; B. kistert, Schaum, on the coast, Ajaccio ; 
B. concinnum, Steph., river banks, Corte; B. nitidulum, 
Marsh., Vizzavona; B. preustwm, Dej., and B. fasciola- 
tum, Duftschm., Bocognano. Yachys apristoides, Rott., 
a few specimens beneath shingle on the banks of the 
River Tavignano, at Corte. Tachyta nana, Gyll., rarely, 
under bark, Vizzavona. Perileptus areolatus, Creutz, on 
the banks of streams, Bocognano and Corte. Synuchus 
nivalis, Panz., Vizzavona. Calathus circumseptus, Germ., 
sea-coast, Ajaccio. Pristonychus carinatus, Chaud.,* 
commonly in the beech-forests at Vizzavona, beneath 
loose bark. Pterostichus (Adelopterus) ambiquus, 
Fairm.,* commonly, with the preceding. Percus reichei, 
Kr.,* P. corsicus, Dej.,* and P. loricatus, Dej.,* under 
stones, old logs, etc., in the beech-forests, also in bare 
places on the mountains, Vizzavona, the first-named 
commonly, the others more rarely ; P. ramburi, Lap.,* 
under rotten Opuntias, Ajaccio. Amara ovata, I, 
A. familiaris, Duftschm., and A. trivialis, Gyll., Vizza- 
vona; A. striatopwnctata, Dej., on the mountains, 
Vizzavona. Harpalus belliert, Reiche,* commonly, at 
Vizzavona, in company with Percus; H. consentaneus, 
Dej., H. distinguendus, Duftschm., H. rubripes, Duftschm., 
and H. honestus, Duftschm., Vizzavona. Ophonus pwmilio, 
Dej., Ajaccio. Anisodactylus binotatus, F., Vizzavona. 
Stenolophus tewtonus, Schr., Vizzavona. Chlenius 
vestitus, Payk., Ajaccio. Lebia cyanocephala, L., Ajaccio. 
Lionychus sturmi, Gené, rarely, beneath shingle, on the 
banks of the Tavignano, Corte. Dromius meridionalis, 
Dej., and D. quadrisignatus, Dej., Vizzavona, the latter 
not uncommonly, under bark of beech. Brachinus 
sclopeta, F'., Ajaccio, on the beach. 

Hydroporus griseostriatus, Deg., commonly, and 
H. planus, F., singly, in ashallow pond on the mountains, 
supplied by the melting snow, on the Col between 
Vizzavona and Ghisoni, at an elevation of about 5,500 
feet; H. memnonius, Nicol, and H. corsicus, Wehncke ?*, 
singly, in a slightly running stream, Vizzavona. Agabus 
bipustulatus, L., Viazzavona; A. binotatus, Aubé, not 
uncommon, in a small sluggish stream at Vizzavona ; 
A. conspersus, Marsh., a single specimen, on the Col 


Entomological Excursion to Corsica. 933 


between Vizzavona and Ghisoni, not differing from 
examples from the English coast. Gyrinws natator, L., 
Corte and Vizzavona. 


Anacena globulus, Payk., Hemisphera infima, Pand., 
and Limnebius perparvulus, Rey (?), in shallow pools on 
the sandy banks of streams, Bocognano; L. picinus, 
Marsh. (?) and Laccobius, sp., Corte. Spheridium 
scarabxoides, L., and S. marginatuwm, F., Vizzavona. 
Cyclonotum orbiculare, ¥., Corte. Helophor us, two or 
three species, Vizzavona. Hlmis oblongus, Kuw., Bocog- 
nano. Parnus intermedius, Kuw., in plenty, on the banks 
of a stream, Bocognano. 


Ocalea castanea, Er. (?), Vizzavona. Aleochara spissi- 
cornis, Er., Vizzavona; A. mesta, Grav., Corte. Ocyusa 
picta, Rey,* Haploglossa pulla, Gyll., Homalota nigri- 
cornis, 'Thoms., Oxypoda referens, Rey, and O. fusina, 
Rey, ‘fortuitously, Vizzavona. O. alternans, Grav., in 
fungus, Vizzavona. Phlewopora reptans, Care. WP He 
sita, Rey, and P. corticalis, Grav., Ischnoglossa prolixa, 
Grav., and J. corticina, Er., Leptusa fumida, Er., and 
L. (Sipalia) scabripennis, Rey,* Placusa, sp. un. (2), 
Homalota cuspidata, Hr., Conurus pubescens, Payk., 
sparingly, under bark of beech or pine, Vizzavona. 
Homalota fragilis, Kr., on the banks of the Tavignano, 
Corte. Bolitobius trimaculatus, var. kraatzi, Pand., i 
plenty in cit on an old dead beech, Vizzavona. 
Gyrophena sp. (?), G. manca, Er., and G. boleti, Linn., 
in fungus on beech, Vizzavona. Bolitochara obliqua, 
Er., in plenty, and 'B. lunulata, Payk., rarely, under 
bark of beech, Vizzavona. (wuedius impressus, Panz., 
Corte. Ocypus olens, Miill., and O. cyaneus, Payk., in 
the road, Vizzavona. Philonthus alcyoneus, Er., and 
P. luxurians, Er., beneath shingle on the sandy banks of 
the 'Tavignano, Corte. P. nigritulus, Grav., Corte. 
Actohius prolivus, Kr., banks of streams, Bocognano and 
Corte. Lathrobiwm labile, Er., rarely, ag the Philonthi 
at Corte, on the banks of the Tavignano. Xantholinus 
fulgidus, Fabr., Corte. Nudobius collaris, Er., a few 
specimens under beech-bark, Vizzavona. Medon nigri- 
tulus, Kr., banks of streams, Bocognano. Scopeus 
minutus, Hr., banks of the Tavignano, Corte. Stenus 
levigatus, Rey, about running water, on mossy stones, 
Vizzavona and Corte; S. strigosus, Fauv.,* in company 


234 Mr. George C, Champion oi aii 


with the preceding, Corte; S. ater, Mann., ean S. 
asphaltinus, Er., Viezavona and Bocognano; S . fusci- 
cornis, Hr., and WS. erichsoni, Rye, Wizzivona, Tr oqoph- 
leus arcuatus, Steph., Bocognano; 7. corticinus, Grav., 
Ajaccio; J. pusillus, Grav., Vizzavona. Homalium 
jlorale, Payk., H. fossulatwm, Kr., H. pusillum, Grav., 
and H. vile, Er., Vizzavona; H. concinnum, Marsh., 
Bocognano. Anthobium tempestivum, Myr., Vizzavona. 
Phleocharis corsica, Fauv.,* under bark of beech, very 
rare, Vizzavona. 


Bryaxis revelierei, Saulcy (7), on the banks of streams, 
Corte and Bocognano. Huplectus punctatus, Muls., 
under bark, Vizzavona. Neuraphes dubius, Reitt.,* 
under bark, Vizzavona, very rare. 


Silpha granulata, Thunb., in the road, Vizzavona. 
Tnodes humeralis, Kug., in fungoid growth on beech, 
Vizzavona. Arthrolips sp., Corte and Ajaccio.  T'richo- 
pteryx, sp., Vizzavona. Sphexrius acaroides, Waltl, on 
the sandy banks of the Tavignano, Corte. Cybocephalus, 
sp. (?), Corte. Phalacrus coruscus, Panz., Bocognano. 
Olibrus stierlini, Flach, Ajaccio and Bocognano; O. 
biplagiatus, Guill., Ajaccio; 0. afinis, Sturm, Bocog- 
nano. T'riplax rufipes, F., in plenty, in fungus on beech, 
Vizzavona. Hndomychus coccineus, L., under bark an 
beech, Vizzavona. Cryptophagus dentatus, Herbst, C. 
distinguendus, Sturm, and C. scanicus, L., Vicnevane: 
Enicmus minutus, L., Vizzavona; H. testaceus, Steph., 
and H. hirtus, Gyll., one specimen of each under beech- 
bark, Vizzavona; EH. transversus, Oliv., Corte. Lathri- 
dius productus, Rosenh., Ajaccio, one specimen. Corti- 
caria pubescens, Gyll., C. obscura, Bris. (?), and C,. 
gibbosa, Herbst, Vizzavona. Litargus bifasciatus, F., 
under bark of beech, Vizzavona. Mycetophagus quadri- 
pustulatus, L., including the var. bipustulatus, Schilsky, 
in fungus on beech, Vizzavona. Ipidia quadrinotata, 
I’., rarely, under bark of beech, Vizzavona. Meligethes 
zneus, F'., and three other spp., Bocognano. Pityophagus 
Jerrugineus, L., Rhizophagus depressus, F., and fh. 
Jerrugineus, Payk., under bark of Pinus laricto, Vizza- 
vona. Nemosoma elongatum, L., one specimen, on the 
bark of a fallen pine, Vizzavona. Tenebrioides maroc- 
canus, Reitt., not very rarely, under bark of pines, 
Vizzavona. Peltis grossa, L., one specimen, in moss on 


Entomological Hxcursion to Corsica. 235 


the trunk of a dead beech, Vizzavona. Ooaelus pictus, 
Sturm, abundant, Hndophleus spinulosus, Latr., rather 
commonly, Colydiwm elongatum, F., rarely, Ditoma 
crenata, F., rarely, and Brontes planatus, L., in plenty, 
beneath the bark of decaying beeches, Vizzavona. 
Aulonium bicolor, Herbst, commonly, in the burrows of 
Scolytide, in the bark of pines, Vizzavona. Cerylon 
histeroides, F., and Q. ferruginewm, Steph., under bark of 
beech, Vizzavona. Lamophleus pusillus, Schonh., and 
Silvanus unidentatus, Oliy., rarely, under bark, Vizza- 
vona. Dermestes vulpinus, F., Ajaccio. Hadrotoma 
variegata, Kust., on flowers, not rare, Ajaccio, Bocog- 
nano, Vizzavona, ‘attone, Vivario. Anthrenus varius, 
Pe, and, A. sp? ), common, on flowers, Ajaccio, Bocog- 
nano, etc. Limnichus incanus, Kies., Corte. 


Platysoma oblongum, F., Plegaderus saucius, Er., and 
P. sanatus, Truqui, all three not uncommonly, beneath or 
in the bark of pines, apparently preying on the larva of 
Scolytidxe, Vizzavona. Hister major, L., H. inequalis, 
Oliv., H. 4-maculatus, L., H. cadaverinus, Hoftm., H. 
sinuatus, F., and H. purpurascens, Herbst, on the roads, 
in dung, etc., Vizzavona; H. pustulosus, Gené, Bocog- 
nano. Saprinus senupunctatus, F., commonly, and NV. 
furvus, Hr., rarely, under a dead horse, on the beach, 
Ajaccio; SS. chalcites, Ill., under dung in the road, 
Ajaccio. Paromalus parallelopipedus, Herbst, and P. 
flavicornis, Herbst, under bark of beech, Vizzavona. 


Lucanus tetraodon, Thunb., a single ¢, found in a 
rotten pine-stump, and another example seen on the 

wing, flying about the top of a tree, June 27th, Vizza- 
vona; subsequently found in some numbers by ‘Colonel 
Yerbury. Dorcus parallelopipedus, L., in rotten beech, 
etc., Vizzavona, not uncommon. Afeuchus pius, Il., in 
the roads, Bocognano and 'attone; A. laticollis, L., 
abundant about dung, Corte, Apa Vizzavona, 
Ajaccio, ete. Sisyphus schefferi, L., abundant, with the 
preceding.” Gymnopleurus sturmi, Macl., Ajaccio; G. 
mopsus, Pall., Vizzavona. Onthophagus amyntas, Oliy., 
O. tawrus, Schr., O. wuchicornis, L., O. fracticornis, 
Preyssl, and O. schreberi, L., Oniticellus flavipes, F., 
Vizzavona, Corte, etc. Aphodius erraticus, L., A. 
pusillus, Herbst, A. scrofa, F., A. hemorrhoidalis, L., 
Vizzayona. Greotrupes geninatus, Gené, abundant every- 


236 Mr. George C. Champion on ait 


where in the mountains, at an elevation of 2,500 feet 
and upwards, Vizzavona; G. hypocrita, Serv., Ajaccio ; 
G. stercorarius, Linn., Vizzavona. Oryctes yrypus, Ill., 
Ajaccio, one ¢. Pachypus cornutus, Oliv., one ¢, flying 
in the town of Ajaccio, towards evening. Rhizotroqus 
bellicrt, Reiche, four specimens, found by Colonel Yer- 
bury in July, Vizzavona. Polyphylla fullo, L., Vizza- 
vona. <Anomala junit, Duftschm., Ajaccio. Cetonia 
aurata, L., very abundant on flowers at Tattone, Vizza- 
vona, “eam. Corte, etc., and exceedingly variable, 
green, blue, coppery, or black, often green with the 
thorax and scutellum coppery ; C.jloricola, Herbst, rarely, 
in company with the preceding ; C. opaca, Gory, and C. 
morio, F., Ajaccio, the latter very abundant on thistle 
flowers, and difficult to extract therefrom, also at Corte, 
etc.; C.hirtella, L., and CO. stictica, L., Vivario, Vizza- 
vona, Corte, ete. Trichius gallicus, Heer (abdominalis, 
Dej.), Ajaccio, and 1’. zonatus, Germ., rarely, Bocognano 
and Vizzavona. 


Chalcophora mariana, Lap., a few specimens on the 
trunks of pines, Vizzavona; subsequently found by 
Colonel Yerbury in some numbers with various other 
Buprestide at a new saw-mill at Tattone. Capnodis 
tenebricosa, Herbst, not uncommonly on the coast at 
Ajaccio, flying about in the hot sun. Aneylochira 
rustica, L., A. flavomaculata, F., and <A. octoguttata, L., 
ut the saw mill, 'Tattone, Colonel Yerbury. Melanophila 
appendiculata, F., and Phenops cyanea, F., rarely, pine 
forests, Vizzavona. Anthavia inculta, Germ., and 
A. grammica, Lap., not rarely, on flowers, Ajaccio ; 

. millefolit, F., Ajaccio. Ptosima 9-maculata, F., on 
bushes, Ajaccio. Acmexodera adspersula, Hl., not un- 
commonly, <A. teniata, F., rarely, on flowers, 
Ajaccio. SE lineata, ¥., Ajaccio. Chryso- 
bothrys affinis, F., saw-mill, Tattone, Colonel voce 
Corebus rubi, L., on bramble bushes, Ajaccio and Corte. 
Agrilus viridis, L., Vizzavona; A. roscidus, Kies., 
Ajaccio; A. soliert, Gory (7), Corte and Ajaccio. Trachys 
pygmexa, F., on flowers, Ajaccio. Throscus carinifrons, 
De Bony., not rarely, by sweeping bracken towards 
evening, in the pine forests, Vizzavona; J. corsicus, 
Reitt.,* three examples by sweeping, Bocognano and 
Vizzavona. Melasis buprestoides, L., in dead beech, 
Vizzavona. Adelocera carbonaria, Schrank, commonly, 


Entomological Excursion to Corsica. 237 


under bark of dead pines, especially where charred by 
fire, Vizzavona. later preustus, F., commonly, under 
bark of pines, Vizzavona, varying greatly in the amount 
of black at the apex of ‘the elytra; H. pomonx, Steph., 
four specimens, with the preceding, inseparable from 
others from England (from the New Forest). Orypto- 
hypnus meridionalis, Lap., rarely (and very difficult to 
secure), on the banks of the Tavignano, Corte. Cuardio- 
phorus ulcerosus, Gené, and OC. eleonore, Gené, com- 
monly, by sweeping, Bocognano, Athous rhombeus, 
Oliv., one specimen, Vizzavona; A. depressifrons, 
Desbr. (?),* a few examples, by Pweepine: Corte 
and Vizzavona; A. corsicus, Desbr. (?), one specimen, 
Corte. 


Helodes genei, Guér., damp places, Corte, Vizzavona, 
Bocognano.  fydrocyphon  deflexicollis, Miill., Corte. 
Cyphon, sp. (?), Vizzavona. Dictyoptera sanguinea, Scop., 
pine forest, Vizzavona, rarely. Telephorus precoz, 
Gené,* not uncommon, on flowers of foxglove, forest 
of Vizzavona. Rhagonycha corsica, Reiche, on flowers 
of Umbelliferze, etc., Vizzavona, Bocognano, Vivario, 
Corte. Malthodes, 3 spp. (?), Vizzavona. Troglops silo, 
Kr., three specimens, on the coast at Ajaccio. Charopus 
concolor, F., by sweeping, Bocognano. Oolotes macu- 
latus, Lap., on flowers, Corte and Ajaccio; C. punctatus, 
Hr., rarely, on the sandy beach at Ajaccio. Sphinginus 
apicalis, Perr., a few specimens on flowers, Vizzavona. 
Attalus (Antholinus) sericans, Hr., and sp. (?), Corte and 
Bocognano. Malachius rufus, Oliv., Corte; M. sardous, 
Kr., Bocognano; M. bipustulatus, L., Vizzavona and 
Corte ; M. (Oogynes) longicollis, Kr., commonly, on the 
flowers of low plants on the mountains, between 
2,000-5,000 feet, Bocognano and Vizzavona; M. cyani- 
pennis, Kr., Corte. Dasytes tibialis, Muls., rarely, in 
flowers, pine forest, Vizzavona; D. “fusculus, Iil., and 
D. plumbeus, Miill., commonly, Vizzavona, Ajaccio, ete. ; 
D. pilicornis, Kies. (?), one specimen, Bocognano. 
Dolichosoma nobile, Kies., common, on the coast at 
Ajaccio. Haploenemus impressus, Marsh., on flowers, 
Bocognano and Vizzavona; H. wxanthopus, Kies.,* 
Vizzavona; H. cribricollis, Muls.* (?), Corte and Ajaccio. 
Danacexa picicornis, Kiist., and sp. (?), commonly, on 
flowers, Vivario, Vizzavona, Bocognano and Ajaccio. 
Clerus ‘formicarius, lig abundant on cut pines, Vizza- 


238 Mr. George C. Champion on an 


vona; CU. 4-maculatus, Schall., one specimen, on a fallen 
pine, Vizzavona. 'richodes alvearius, F., Vivario and 
Vizzavona. Plinus fur, L., in the Hotel Dependance at 
Vizzavona; P. dubius, Sturm, Vizzavona; LP. lichenum, 
Marsh., Vivario and Corte. Dryophilus densipilis, Ab., 
four specimens, by eeceping , Corte. Hrnobius parens, 
Mals., and Hrnobius, sp. (?), by beating dead pine tops, 
Vizzavona. Xestobium rufovillosum, Deg., Vizzavona. 
Xyletinus laticollis, Duftschm., rarely, by sweeping, 
Bocognano. Metholeus cylindricus, Germ., Ajaccio. 
Lasioderma bubalus, Fairm., Corte. 

Bostrychus capucinus, L., fortuitously, Vivario and 
Vizzavona. Cis boleti, pi C. setiger, Mell., C. hispidus, 
Gyll, and /C. sp. (?), Vizzavona, 


Pachychila servillet, Sol., and Tentyria ramburi, Sol., 
on the sandy shores of the Bay of Ajaccio. Akis 
punctata, Thunb., on the roads, not rarely, near Corte. 
Asida carinata, Sol., abundant on the Col between 
Vizzavona and Ghisoni, at about 6,000 feet, under stones, 
also occasionally on the lower slopes, and on the roads 
about La Foce; A. corsica, Lap., two specimens, on the 
coast at Ajaccio. Pimelia sardoa, Sol., crawling about 
Siete along the shore of the Bay of Ajaccio. 

Crypticus gibbulus, Quens., common, under stones, 
Ajaccio. Dendarus tristis, Rossi, Ajaccio, Vivario, and 
Vizzavona. Pedinus meridianus, Muls., Ajaccio and 
Corte. Colpotus godarti, Muls. (one ¢), Ajaccio, under 
stones. Opatrum pusillum, F., and O. rusticum, Ol., 
Ajaccio. Ammophthorus rufus, Luc., abundantly, Pha- 
laria oblonga, Kiist., commonly, and P. cadaverina, F., 
rarely, along the shores of the Bay of Ajaccio. Diaperis 
boleti, i., one specimen, in fungus on beech, Vizzavona. 
Corticeus castaneus, F., and C. pini, Panz. (?), under 
bark of pines, common, forest of Vizzavona; C. bicolor, 
Oliv., under bark of beech, Vizzavona. Uloma culinaris, 
Linn., rarely, and U. perroudi, Muls., and Menephilus 
curvipes, ¥'., more or less commonly, under the bark of 
pines, Vizzavona. Helops superbus, Muls., one of the 
finest species of the genus, and apparently confined to 
the mountain-forests of Corsica, and H. ceruleus, L., 
both common, beneath the loose bark of pine stumps, 
or of dead standing pines, Vizzavona; H. genei, Allard, 
four specimens, beaten from branches of a fallen pine, 
Vizzavona. ILsomira corsica, Muls., Bocognano; J. 


Entomological Excursion to Corsica, 239 


murina, L., Vizzavona. Omophlus lepturoides, F., 
Ajaccio and Corte; O. picipes, F., Ajaccio and Vizza- 
vona. Lagria hirta, L., Vizzavona. Orchesia undulata, 
Kr., in fungus on beech, Vizzavona. Xylita revelieret, 
Muls., four examples, in the crevices of the bark of a 
fallen pine, Vizzavona. Melandrya caraboides, L., not un- 
common, under bark of beech, etc., these specimens have 
the elytra bluer in colour than ‘those from England. 
Salpingus nitidus, Chevr. (= reyt, or exsanguis, Abeille), 
four specimens, by beating dead pine branches, Vizza- 
vona ; originally recorded from Algeria. Rhinosimus 
plamirostris, Vizzavona. Notoxus cornutus, F., Bocog- 
nano. Anthicus longicollis, Schmidt, on the sandy banks 
of streams, beneath stones, Corte and Bocognano; A. 
fenestratus, Schmidt, Corte; A. corsicus, Lap., swarming 
on bushes (including olive) on the hill-sides near Ajaccio, 
also at Bocognano. Mordella aculeata, L., M. fasciata, 
Fabr., and M. bipunctata, Germ., Ajaccio and Corte. 
Mordellistena pumila, Gyll., Ajaccio, Anaspis thoracica, 
L., A. pulicaria, Costa, and A. spp., Vizzavona, eee 
ete. Silaria suturalis, Em., Corte, Vivario, Bocognano, 
and Vizzavona. Meloe brevicollis, Panz., two specimens, 
on the mountain slopes above Vizzavona, at about 
4,500 feet elevation. Zonitis mutica, Scriba, abund- 
ant, on thistle-heads, Corte. Cidemera podagrarix, L., 
O. barbara, F., O. flavipes, F., O. nobilis, Scop., and 
O. lwrida, Marsh., Ajaccio, Bocognano, Corte, etc. 
Stenostoma rostratum, #., on the beach, Ajaccio. 
Otiorrhynchus guttula,* Fairm., swarming on the 
catkins of the alder, on the summit of the Col between 
Vizzavona and Ghisoni, occasionally found lower down ; 
O. intrusus,* Reiche, Bocognano; O. corsicus,* Fairm., 
a few specimens under stones, in the same locality as 
O. guttula. Peritelus parvulus,* Seidl., one specimen, 
Vizzavona ; P. (Meira) lutiscrobs,* Desbr., two specimens, 
by sweeping bracken in the pine-forest of Vizzavona. 
Phyllobius pellitus, Boh., swarming on alders, Vizza- 
vona. Metallites parallelus,* Chevr., commonly, by 
beating arborescent heath, pines, and other trees, Bocog- 
nano, Vizzavona, etc. Polydrusus leucaspis, Boh., Corte. 
Strophosomus coryli, F., Vizzavona. Caulostrophus 
delarouzet, Fairm., by beating various trees, Bocognano 
and Vizzavona. Brachyderes analis, Desbr.,* in plenty 
on pines at the upper limit of their growth on the 
mountain slopes above Vizzavona. Thylacites fritillum, 


240 Mr. George C. Champion on an 


Panz., Ajaccio, and 7’. lapidarius, Gyll., Bocognano and 
Vizzavona, under stones. Cleonus (Plagiographus) 
tabidus, Oliv. (pelleti, Fairm.), one specimen, beaten 
from arborescent heath, Bocognano. Livxus scolopax, 
Boh., not rare, Ajaccio and Bocognano ; L. cribricollis, 
Boh., Vizzavona. Larinus latus, Herbst, on thistles, 
Corte ; L. genei, Boh., Corte, Bocognano, and Ajaccio; 
L. turbinatus, Gyll., Corte. Hypera pedestris, Payk., 
Bocognano; H. trilineata, Marsh., Corte. Pissodes 
notatus, F., commonly on pines, Vizzavona. Smicronyx 
corsicus,* Fairm., not uncommonly, by beating arbores- 
cent heath, Bocognano and Tattone. Anoplus roboris, 
Suffr., Vizzavona. Brachytemnus porcatus, Germ 
common, under pine-bark, Vizzavona. Acalles setuli- 
pennis,* Desbr., and A. henont, Bedel, Vizzavona; the 
latter hitherto only known from Algeria. COwliodes 
exiguus, Oliv., Ajaccio. Ceuthorrhynchus echit, By 
Ajaccio and Bocognano; OC. duvali, Bris., one specimen, 
beaten from alder-catkins, on the summit of the Col 
between Vizzavona and Ghisoni; OC. smaragdinus, Bris., 
Corte; CO. cyanipennis, Germ., Corte and Ajaccio. 
Ceuthorrhynchidius troglodytes, F'., Bocognano. Baris 
imsularis, Desbr.,* one specimen, Ajaccio ; B. cerulescens, 
Scop., Corte. Balaninus elephas, Gyll., Vizzavona ; B. 
turbatus, Gyll., and B. glandiwm, Marsh., Bocognano. 
Sibinia arenarix, Steph., Ajaccio; S. attalica, Gyll., not 
uncommonly, Bocognano. Miccotrogus suturatus, Per- 
ris (?),* one specimen, Bocognano. Orchestes erythropus, 
Germ., var. tricolor, Kies., ‘and OQ. fag, L., Vizzavona. 
Mecinus circulatus, Marsh., Bocognano ; M. pyraster, 
Herbst, Corte. Gymnetron pascuorwm, Gyll., Bocog- 
nano; G. tetrum, F., on Verbascum, Vizzavona; G. 
lanigerum, Bris. (?), Bocognano. Miarus campan ule, ies 
Vizzavona. OCtonus hortulanus, Marsh., Ajaccio, Bocog- 
nano, Corte. Nanophyes niger, Waltl, Bocognano and 
Vizzavona; N. nitidulus, Gyll., Bocognano. Apion 
flavofemoratum, Herbst, A. viciw, Payk., Ajaccio; A. 
frumentarwm, 1a, Al eneum, Woe Al inalies, Tede 
levicolle, Kirby, A. brevirostre, Herbst, and other 
species, Corte; A. nigritarse, Kirby, and A. violaceum, 
Kirby, Vizzavona; A. tubiferum, Gyll., and A. subparal- 
lelum, Desbr. (2), Bocognano. Attelabus cuwrculionoides, 
L., Bocognano, Vizzavona, Corte. Platyrrhinus latiros- 
tris, ., not uncommon in fungus on old decaying beech- 


co) 
trees, forest of Vizzayona. Scolytus rugulosus, Ratz., 


Entomological Hxcursion to Corsica. 241 


Ajaccio. Myelophilus minor, Hart., in pine-bark, Vizza- 
vona; these specimens are unusually large, and have 
strongly tuberculate elytra. Hylastes ater, Payk., and 
H, attenuatus, Er., on the pines, Vizzavona. Cryphalus 
numidicus, Hichh., Tomicus sexdentatus, Boern., T. 
suturalis, Gyll., T. longicollis, Gyll., and T. proximus, 
Hichh., Pityophthorus glabratus, Hichh., and Pityogenes 
bistridentatws, Hichh. (?), in or under bark of Pinus 
laricio, Vizzavona. Orypturgus cinereus, Herbst, in the 
small branches of the same trees, Vizzavona. Xyleborus 
sawesent, Ratz., rarely, and Taphrorychus bicolor, Herbst, 
commonly, on beech-logs, Vizzavona. 


Bruchus longicornis, Germ., common, Ajaccio and 
Vivario; B. atomarius, L. (?), Corte; B. dispar, Germ., 
Bocognano; and various other species of the genus. 
Spermophagus cardut, Boh., Corte. 


Spondylis buprestoides, L., saw-mill at Tattone, Colonel 
Yerbury, three examples. Hrgates faber, L., fragments of 
dead specimens only, in the rotten pine-trees at Vizza- 
vona, in June; one male bred from a pupa after return 
to England ; female subsequently obtained in numbers 
at Tattone by Colonel Yerbury; the enormous ¢ larve 
sometimes found under bark of dead standing pines. 
Leptura rubra, L., L. fulva, Deg., L. scutellata, F., 
and J. hastata, Sulz., Vizzavona. Strangalia auru- 
lenta, F., S. maculata, Poda, Vizzavona; S. bifasciata, 
Mill., Corte and Ajaccio. Stenopterus rufus, L., Vizza- 
vona, and S. ater, L., Vivario, Corte, and Ajaccio, on 
Umbelliferous flowers. Cerambyx scopolvi, Fuessly, 
Criocephalus rusticus, L., Hylotrupes bajulus, L., Vizza- 
vona and T'attone, also found by Colonel Yerbury. 
Clytus arietis, L., and C. rhamn, Germ., Vizzavona 
and Corte. Xylotrechus arvicola, Oliv., Vizzavona. 
Clytanthus massiliensis, L., common on flowers, Ajaccio, 
Bocognano, Vivario, Corte. Morimus lugubris, F., 
common, in the beech-forest at Vizzavona, about logs, 
ete. Acanthocinus griseus, F., about twenty specimens, 
on a fallen pine, Vizzavona. Pogonocherus perroudi, 
Muls., a few specimens, in company with the preceding, in 
the pine-forest at Vizzayona. <Anzsthetis testacea, L., 
one specimen, Vizzavona. Agapanthia cardw, L., Vizza- 
vona. Phytecia virescens, F., Ajaccio, Bocognano, and 
Corte; P. ephippium, F., Ajaccio. 

TRANS. ENT. SOC, LOND. 1891.—PART I. (MARCE.) Q 


242 Mr. G. C. Champion on an Excursion to Corsica. 


Labidostomis taxicornis, F., and L. centromaculata, 
Gené, not uncommon, Ajaccio, Bocognano, and Corte. 
Lachnea sexpunctata, Scop., abundant, on bushes, in the 
hot sun, Ajaccio, Bocognano, Vizzavona, Corte, etc. 
Clythra leviuscula, Ratz., Corte. Titwhaa biguttata, Oliv., 
var. dispar, Luc., Corte. Coptocephala scopolina, L., com- 
mon, flying in the hottest sun, Ajaccio. Cryptocephalus 
pygmeus, F., Ajaccio, Bocognano, and Vizzavona; C. 
morext, L., and C. ocellatus, Drap., Vizzavona, rarely. 
Pachybrachys scriptus, H.-S., and Stylosomus depilis, Ab. 
(nec minutissimus, Germ.), not uncommon, by beating 
arborescent heath, etc., Bocognano. Colaspidea oblonga, 
Blanch., Ajaccio. Chrysomela banksi, F., and C. hemo- 
ptera, L., Ajaccio; C. quadrigemina, Suffr., Bocognano. 
Lina populi, L., Vivario. Prasocwris (Hydrothassa) 
pumila, Reiche, Vizzavona. Lwperus flavipes, L., Vizza- 
vona. Galeruca corsica, Joann.,* Vizzavona. Galerucella 
luteola, Mull., Corte. Podagrica intermedia, Kutsch., 
Corte. Crepidodera rubicunda,* Perris, Corte. Haltica 
sp., Longitarsus spp., Bocognano. Psylliodes sp., Corte. 


Hispa atra, L., Bocognano. Cassida equestris, F., 
Vizzavona; C. hemispherica, Herbst, Bocognano. 


Epilachna chrysomelina, F., Ajaccio. Coccinella oblon- 
goguttata, L., and C. 18 guttata, L., on the pines, Vizza- 
vona; C. 11-notata, Schneid., Bocognano and Corte; C. 
revelierit, Muls., Corte; C. conglobata, L., Vizzavona; 
C. 7-punctata, L., C. 22-punctata, L., Ajaccio, ete.; C. 
bipunctata, L., C. variabilis, Il., and C. 14-punctata, L., 
Vizzavona, ete.  Chilocorus bipustulatus, L., Ajaccio. 
Heochomus 4-pustulatus, L., Vizzavona, Bocognano, ete. 
Platynaspis villosa, Fourcr., Bocognano. Scymnus fas- 
ciatus, Fourcr., S. damryi, Weise,* and spp., Bocognano, 
Vizzavona, etc. 


VIII. A list of the Hemiptera-Heteroptera collected by 
Mr. Champion in Corsica in June, 1893, with a 


description of one new species. By Hpwarp 
SaunpErs, F.L.S8. 


[Read Feb. 28th, 1894.] 


Odontoscelis fuliginosa, Linn., Ajaccio. 

Odontotarsus grammicus, Linn., Vivario and Vizza- 
vona. (2500—4000 ft.) 

Lurygaster maura, Linn., Vizzavona. 

Ancyrosoma albolineatum, Fab., Ajaccio, Vizzavona. 

Graphrosoma lineatum, Linn., Ajaccio, Vizzavona. 

Brachypelta aterrima, Foerst., Corte. 

Sciocoris sulcatus, Fieb., Corte, Bocognano. 

Dryoderes marginatus, Fab., Corte. 

Aflia acuminata, Linn., Vizzavona, Vivario. 

Peribalus vernalis, Wolff, Corte. 

Carpocoris fuscispinus, Boh., Vizzavona, Ajaccio. 

Palomena vir idissima, Pod: Wassepona! 

Pentaloma 3 juniperina, Linn., Bocognano. 

Piezodorus incarnatus, Germ. »  Bocognano and 
Vizzayona. 

Rhaphigaster grisea, Fab., Ajaccio. 

Hurydema ornatum, L., Corte. 


< decoratum, H.-S., Corte, Vizzavona, Ajaccio. 
pictum, H.-8., Ajaccio. 
- oleraceum, Linn., Vizzavona. 


Hlasmostethus interstinctus, Linn., Vizzavona. 
Spathocera lobata, H.-S., Bocognano, Vizzavona. 
Centrocoris spiniger, Fab., Bocognano, Vizzavona. 
Syromastes marginatus, Linn., Corte, Vizzavona. 
Verlusia rhombea, Linn., Ajaccio, Bocognano, 
Vizzavona. 
ra sulcicornis, Fab., Bocognano. 
Gonocerus insidiator, Fab., Bocognano. 
Ceraleptus squalidus, Costa, Vizzavona. 
Loxocnemis dentator, Fab., Vivario. 
Coreus affinis, H.-S., Bocognano, Corte, Vizzavona. 
»  denticulatus, Scop., Corte, Vizzavona. 
Strobilotoma typhecornis, Fab., Corte and Vizzavona. 
TRANS. ENT. SOC. LOND. 1894.—PaRT I. (MARCH.) 


244 Mr. H. Saunders on Hemiptera- Heteroptera 


Campiopus lateralis, Germ., Vizzavona, Ajaccio. 

Therapha hyoscyami, Linn., Bocognano, Vizzavona. 

Corizus crassicornis, Linn., * : 
Vivario. 

capitatus, Fab., Vizzavona. 

parumpunctatus, Schil., Vizzavona and Bo- 


cognano. 


2) 
2) 


Megalomerium meridionale, Costa, Ajaccio. 
Lyxus militaris, Fab., Ajaccio, Corte, Bocognano, 


Vizzavona. 
:, equestris, Linn., Vizzavona. 
ois 
ae savatilis, Scop., es 
. apuanus, Rossi, ra 


Lygxosoma reticulatum, H.-8., Vizzavona, Vivario. 
Nysius thymi, Wolff, Bocognano. 

Cymus melanocephalus, Fieb., Ajaccio, Vizzavona. 
Ischnorhynchus geminatus, Fieb., Vizzavona. 
Geocoris lineola, Ramb., Ajaccio. 

Heterogaster affinis, H.-S., Vizzavona. 


Microplax interrupta, Fieb., Vizzavona. 


sy albofasciata, Costa, ,, 


Metopoplax ditomoides, Costa, Vizzavona. 

Macroplax fasciata, H.-S., Bocognano, Ajaccio, Vizza- 
vona, Vivario. 

Rhyparochromus pretextatus, H.-8., Vizzavona. 

Stygnus pedestris, Fall., Corte. 

Peritrechus geniculatus, Hahn, Vizzavona. 

Trapezonotus agrestis, Fall., Vizzavona, Vivario. 

Calyptonotus rolandri, Linn., Ajaccio. 


Aphanus inarimensis, Costa, Bocognano, Vizzavona. 


»  douglasi, Fieb., Vizzavona.* 


collected in Corsica by Mr. Champion. 245 


Beosus erythropterus, Brullé, Vizzavona, Corte. 
»»  luscus, Fab., Corte. 


Eremocoris fraternus, Horv. (?), Vizzavona. 

Scolopostethus cognatus, Fieb., Vizzavona and Bocog- 
nano. 

Gastrodes ferrugineus, Linn., Vizzavona. 

Pyrrhocoris apterus, Linn. , Ajaccio. 

Serenthia leta, Fall., Wien one 


Monanthia cardui, Linn., Ajaccio, Corte, Vizzavona. 
og Pe var. cognata, Fieb., Vivario.* 
y 5; awriculata, Costa, var. (?), Vizzavona, 
Corte. 
angustata, H.-S., Bocognano. 
wolfit, Fieb., Bocognano, Ajaccio, Corte. 
nassata, Put. , Ajaccio. 


PP) 
39 


33 


Aradus cinnamomeus, Panz., Vizzavona, Bocognano. 
kruepert, Reut., Vizzavona. 

varius, Fab., Vizzavona. 

depressus, Fab., Vizzavona. 


bP) 
3) 


3) 


Gerris najas, De Geer, Vizzavona. 
»,  thoracica, Schum., Vizzavona. 
»  gibbifera, Schum., Corte. 


Harpactor erythropus, Linn., Ajaccio, Corte, Vivario, 
Vizzavona. 


33 wracundus, Poda, Corte. 


Coranus xgyptius, F., Ajaccio. 
Nabis lativentris, Boh., Corte, Vizzavona. 


rugosus, Linn., Vizzavona. 
ferus, Linn., Vizzavona. 


9 
3) 


Salda cocksii, Curt., var. geminata, Corte. 


Piezostethus obliquus, Cost., Ajaccio. 
Scoloposcelis angusta, Reut., Vizzavona. 
Elatophilus nigricornis, Zett., " 
Anthocoris confusus, Reut., a 
Triphleps nigra, Wolff, Ajaccio. 
horvathi, Reut. (?), Corte. 


2) 


246 Mr. H. Saunders on Hemiptera-Heteroptera 
Miris levigatus, Linn., Vizzavona. 


Lopus lineolatus, Brullé, Vizzavona. 
»» mat, Rossi, Vizzavona. 


Phytocoris femoralis, Fieb., Bocognano. 


Calocoris vandalicus, Rossi, Corte. . 


we sewpunctatus, Fab., Ajaccio, Vizzavona, 
Corte. 

6 bipunctatus, Fab., Bocognano, Vizzavona, 
Corte. 

re trivialis, Cost., Bocognano. 

a ventralis, Reut., Corte. 


Lygus pratensis, Fab., Vizzavona. 
& pastinace, Fall., Vizzavona, Ajaccio. 
»  cervinus, H.-S., 
Poeciloscytus g yllenhali, Fall, Corte. 
Camptobrochis punctulata, Fall. , Vizzavona, Bocognano. 
Capsus laniarius, Linn., Corte. 
Pilophorus cinnamopterus, Kb., Bocognano, Ajaccio. 


Strongylocoris obscwrus, Ramb., Bocognano. 


Orthocephalus champroni, Saund.* 
(See description at the end of the list.) 


Dicyphus hyalinipennis, Klug, Ajaccio. 
“2 geniculatus, Fieb., Bocognano. 


Globiceps sphegiformis, Rossi, Corte. 
Heterotoma diversipes, Put., Corte, Ajaccio. 


Placochilus seladonicus, Fall., Ajaccio. 
Pachyxyphus lineellus, M. & R., Corte, Vizzavona. 


Macrotylus longirostris, Fieb., Bocognano. 


Psallus mollis, M. & R., Vizzavona, Bocognano. 
»  anchorifer, Fieb., Bocognano, Ajaccio. 


Criocoris crassicornis, Hahn, Corte. 


Ooriaa semistriata, Fieb., Vizzavona. 


collected in Corsica by Mr. Champion. 247 


Orthocephalus championt, sp. n. 

Piceo-niger nitidiusculus, levis, pilis nigris, pilisque squami- 
formibus albis, facillime divellendis, vestitus ; capite transverso, 
oculis rotundatis, prominulis, vertice inter eos quam oculo ( ¢) fere 
duplo, (2) duplo et dimidio latiore, postice acute carinato, autennis 
pilosulis, articulis primo et secundo incrassatis, testaceis, hoc versus 
apicem piceo nigro, duobus sequentibus simul sumptis longiore, 
maris lateribus subparallelis, femine fusiforme ; tertio quartoque 
filiformibus, tertio basi testaceo, quarto duplo longiore ; prothorace 
levi, trapezoidale, planiusculo, angulis anticis rotundatis, lateribus 
rectis, callis discretis ; scutello Jeevi ; hemielytris (¢) corio interne 
plus minus late pallido-testaceo, membrana infumata, venis nigro- 
piceis, basi areolisque pallidis, (? brach.) abdomine brevioribus, 
apicibus oblique truncatis, corio margine interno basi arcte pallido ; 
pedibus testaceis, femoribus piceo nebulosis, tibiis nigro punctatis, 
setis spiniformibus ornatis ; subtus nigropilosus et albo squamiger, 
Long.: ¢ 44 mm., 9 4mm. 


Hab. Bocognano, Corsica, on aromatic plants. 


This is a beautiful addition to the Huropean fauna, 
which I have pleasure in naming after its discoverer. 
I have sent examples of it to Dr. Reuter, who has kindly 
examined them, and agrees in considering them as 
belonging to an undescribed species allied to bivittatus, 
Fieb., though at once distinguishable by the form, etc., 
of the antenne, 


gia ton sat 
OM, e Oe ; 


© ay 
alee 
a. 
le Pts, 


( 249 ) 


IX. On the Phylogeny of the Pierine, as illustrated by 
their Wing-markings and Geographical Distri- 
bution. By Freverick A. Drxey, M.A., M.D., 
F.ELS., Fellow of Wadham College, Oxford. 


[Read February 7th, 1894.] 
Pruares, IT, LV.) Vic 


TABLE OF CoNTENTS. 


Page 
I. INTRODUCTORY . : : p : : . 249 
II. Tot WING-MARKINGS f . 202 
1. The submarginal series of dark oe . 252 
2. The dark marginal area or spots . . 262 
3. The light series between 1 and 2 : = 269 
4, The discoidal spots. : 269 
5. The markings on the underside of tite 
hindwing : : ars) 
a. The yellow meee Aree f Be ris: 
b. The pinkish basal patch . ‘ ey 
c. The red basal areas . , ‘ = 290 
III. PHYLOGENETIC CONCLUSIONS . 4 : zoo 
1. The evidence of the Wing-markings . . 289 
2. The evidence of Distribution . : Bees 4 
IV. INDEX OF SPECIES MENTIONED . ; ‘ . 330 


I. Inrropuctory. 


In the present paper my object has been to apply to 
the study of the subfamily Pierinz the methods which I 
used in investigating the phylogenetic relations of another 
group of Lepidoptera, as recorded in a former communi- 
cation to this Society.* I have in the first place 
endeavoured to work out the homology of the various 


* “On the Phylogenetic Significance of the Wing-markings in 
certain genera of the Nymphalide,” Trans. Ent. Soc. Lond., 1890. 


TRANS. ENT. SOC. LOND. 1894.—PartT U1. (JUNE). R 


250 Dr. Frederick A. Dixey on the 


markings to be found throughout the group, and have 
then attempted to state the phylogenetic conclusions to 
which this evidence appears to lead. Though my 
principal aim has been the elucidation of the Pierine 
wing-markings, which so far as | am aware have never 
before been systematically studied from this point of 
view, I have not ignored those other features that are 
usually known as “ structural” ; and I have also tried to 
estimate the bearing upon phylogenetic questions of the 
facts relating to the present distribution of the Pierine 
in space. ‘lhe serious limitations under which anyone 
who wishes to construct a phylogeny for such a group as 
this must necessarily labour have been elsewhere acknow- 
ledged; it is of course manifest that little or no help 
can be expected from embryological or palzontological 
sources. There are, however, many compensating features 
to be reckoned on the other side; and in view of certain 
comments that have at times been passed on previous 
work of this kind, I may perhaps be allowed to quote a 
few sentences from the writings of one of our leading 
naturalists, which seem to me to state the special ad- 
vantages afforded by these and similar researches with 
a force and cogency that it would not be easy to gainsay. 
After pointing out the pre-eminent value of the Diurnal 
Lepidoptera to the student of distribution and variation, 
the writer I refer to proceeds as follows—‘‘ But besides 
their abundance, their universal distribution, and the great 
attention that has been paid to them, these insects have 
other qualities that especially adapt them to elucidate the 
branches of inquiry already alluded to. These are the 
immense development and peculiar structure of the wings, 
which not only vary in form more than those of any other 
insects, but offer on both surfaces an endless variety of 
pattern, colouring, and texture. . . . This delicately 
painted surface acts as a register of the minutest differ- 
ences of organization,—a shade of colour, an additional 
streak or spot, a slight modification of outline continually 
recurring with the greatest regularity and fixity, while the 
body and all its other members exhibit no appreciable 
change. The wings of butterflies, as Mr. Bates has well 
put it,* ‘serve as a tablet on which Nature writes the 


* See “ The Naturalist on the Amazons,” 2nd edit., p. 412. 


Phylogeny of the Pierinee. 251 


story of the modifications of species’; they enable us to 
perceive changes that would otherwise be uncertain and 
difficult of observation, and exhibit to us on an enlarged 
scale the effects of the climatal and other physical con- 
ditions which influence more or less profoundly the 
organization of every living thing.”* Whether any par- 
ticular investigator is or is not competent to undertake 
this kind of research may be open to question, but I think 
that the study itself needs no further vindication than 
these words of Mr. Wallace. 

In treating of the details of the wing-markings I have 
tried to be intelligible and explicit, even at the cost of 
being somewhat lengthy. I have, however, been obliged 
to content myself with selected examples; a complete 
treatment would run to a vast length, and in most cases 
the lines here laid down may be easily applied to the 
insects not mentioned in this paper. Some of the iden- 
tifications of markings may at first sight seem startling ; 
for instance, that of the straight orange-coloured line 
crossing the disc of the wings in Dercas with the series of 
black spots in the females of Ganoris ; but I believe that 
in every such case it will be found that the homology can 
be certainly traced, step by step. It is hardly necessary 
to point out the importance of being on one’s guard 
against the misleading suggestions of merely superficial 
resemblances, and also of giving due weight to the 
disturbing influence of mimicry. The details of this 
latter process, when minutely examined in a favour- 
able group such as the Pierinx, are seen to be of so 
complicated and interesting a character, that I may 
hope to be excused for having treated of some of 
these details with considerable fulness, and even for 
having ventured to introduce a new term in order to 
distinguish a special case of the phenomenon known as 
“ Convergence.” t 

The generic names employed in this paper are those 
of Mr. Butler in his well-known “ Revision” (Cistula 
Entomologica, vol. i., 1870, p. 88), supplemented by sub- 
sequent memoirs from the hand of the same author, and 


® “On the Papilionide of the Malayan Region,” Trans. Linn. 
Soc., xxv., p. 1 (1866). 
+ See page 298. 


252 Dr. Frederick A. Dixey on the 


by the arrangement at present adopted in the National 
Collection under his charge. I confess that { am not 
always in agreement with this arrangement, but the 
importance of a definite and accessible standard is obvious, 
and no better one could be found for my purpose. Hven 
if the genera are not in every instance distinguished by 
characters of real ‘‘ generic value,” they are usually 
natural groups, and in most cases of great convenience in 
practice. The list of genera and species which I have 
appended may be found useful as indicating the exact 
insects which are mentioned in the course of the paper, 
and also as giving an idea of the scope and contents of 
those genera whose names may still be somewhat un- 
familiar. A paper of this kind is not the place in which 
to introduce changes of nomenclature or to discuss 
questions of synonymy ; and as regards classification I 
have done no more than throw out a few suggestions 
which, perhaps, may one day be taken up. 


II. Tae WinG-MARKINGS. 
1, The submarginal series of dark spots. 


Comparing together the three species of the subfamily 
Pierine that are commonest in this country, viz., Ganoris 
brassice, G. rapx, and G. napi, we find that the females 
of all three resemble one another in the possession of a 
series of black spots on the upper surface of the forewing. 
One of these spots (Fig. 16, 8 8) occurs between the 
second and third median nervules ; another (ibid., S 10), 
between the first median nervule and the submedian 
nervure ; and the third (ibid., S 11) between the 
submedian nervure and the inner margin of the wing. 
The latter spot, which varies considerably in the amount 
of its development, takes the form of a streak nearly 
parallel with the inner margin. In the males, this series 
of spots is evanescent. The whole series is normally 
absent in G. brassice ¢, and in some specimens of 
G. rapes, and G. napi 3; the males of the two latter 
species, however, generally have the spot between the 
second and third median nervule more or less developed, 
though the other two are not present. Turning to the 
underside of the wing in these species, we find the first 
two spots usually present in both males and females; 


Phylogeny of the Pierinee. 253 


the third is but faintly visible in the female, and in the 
male not visible at all. These three spots are all that 
can be seen in ordinary specimens of G. brassicx, rape, 
and napi ; but in some specimens of G. napi, and less 
distinctly in some of G. rapx, the series is continued 
towards the anterior border of the wing by a fourth spot 
placed just behind the junction of the third and fourth 
subcostal nervules, and traversed by the first discoidal. 
The spots that have now been referred to can be seen in a 
very well-marked form in G. gliciria. 

These indications of a submarginal series of black 
markings become more pronounced in Synchloe daplidice. 
Here, in the female, a submarginal row exists in easily 
recognisable form, three of the constituent spots of 
which are clearly homologous with those already des- 
cribed in G. napi, rape, and brassice (Fig. 15). 
In this region of the wing nearly every interspace be- 
tween nervules contains a spot; the spots tending to 
become confluent, especially towards the costal margin. 
There is generally no spot between the first and second 
median nervules, and often none between the submedian 
nervure and the inner margin of the wing. In the male, 
the anterior portion of the series is distinct enough, the 
large spot, however, between the submedian nervure and 
first median nervule is absent from the upper surface, 
though regularly present beneath. The female S. daplidice 
also shows a continuation of the same series on the 
hindwing, well-marked at the costa, but usually becoming 
less plain as the anal angle is approached. In the male 
SS. daplidice, one term of the series is generally all that is 
present in the hindwing; and turning back to G. brassice, 
rape, and napi, we find the same spot invariably 
present on the upper, and often on the lower surface of 
the costa in the hindwing of both sexes (Figs. 15, 16, 
S 14—18). 

Other species of the genus Synchloe present the same 
submarginal series in greater completeness. S. callidice? , 
for instance, has the whole series well developed on the 
forewing, and in most specimens on the hindwing also. 
A comparison with S. daplidice will show that the series 
is really homologous in the two cases, and will, moreover, 
demonstrate that in S. callidice the smallest spot of the 
series belongs to the interspace between the first and 
second median nervules, which in S. daplidice is usually 


254 Dr. Frederick A. Dixey on the 


not furnished with a spot at all; and that the largest and 
best-marked member of the group, namely that between 
the second and third median nervules, corresponds with 
the spot which we saw to be most persistent in the males 
of G. brassice, rape, and napi. This spot, moreover, as 
well as the next most persistent (that between the 
first median and submedian), is usually picked out with 
black on the underside of both S. callidice and S. daplidice, 
where the markings corresponding to those of the upper 
surface are, as a rule, chiefly composed of green or yellow 
scales with but small admixture of black. The hindwing 
of WS. callidice 2 shows a continuation of the series; 
though here the markings as they approach the inner 
margin tend to assume the appearance of chevrons rather 
than of spots, which tendency is still more marked on the 
underside. 

The South American genus TYatochila presents the 
same series on both fore and hindwings, the females as a 
rule most completely (as in 7. autodice). The male of 7’. 
theodice, however, exhibits the whole series quite plainly ; 
and in both of these species the markings in question 
tend to assume the form of a line of chevrons, the apices 
of which point towards the margin of the wing, instead of 
towards the base, as in SN. callidice. 

I propose to distinguish this series of spots as 8 
(for Submarginal), numbering the constituent markings 
according to the interspaces they respectively occupy. 
In all the forms as yet mentioned the full number of 
marginal interspaces, from the costal nervure to the anal 
angle, is in the forewing 11, in the hindwing 8. Another 
space may be reckoned in the hindwing, that, namely, 
between the costal margin and the costal nervure. The 
corresponding space in the forewing is exceedingly narrow, 
and never carries a separate spot. We may take then 
the number of spaces potentially containing spots as 11 
and 9 in the fore and hindwing respectively, and may 
designate all possible terms of the series as 8S 1—20, 
beginning at the costa of the forewing. None of the 
species above referred to exhibits the whole range of spots 
from | to 20 in a discrete condition, sone being generally 
absent and others fused ; the series may, however, be seen 
complete on the upper surface of the forewing and lower 
surface of the hindwing in many specimens of Callidryas 


philea and C. thalestris 2 (Fig. 22,5 1—11). 


Phyloyeny of the Pierinee. 255 


In certain genera (Hronia, Huchloe, etc.), the presence 
of a fifth subcostal nervule gives rise to an additional 
interspace in the forewing; this, however, causes no 
difficulty, inasmuch as the extra interspace so formed is 
never occupied by a distinct spot. If necessary, the 
additional subcostal interspace may be referred to as 
5a; by which plan we shall avoid throwing the number- 
ing out of correspondence with that adopted in the other 
genera (see Fig. 23). In those genera (Pereute, Delias, 
etc.) which possess only three subcostal nervules in the 
forewing, I shall for convenience consider S 4 as 
absent. 

Having identified the series S in the species named, 
we shall find no difficulty with the remaining members of 
the genera Synchloe and Tatochila, all of which possess it 
in a condition of greater or less development. In all these 
cases, if the sexes differ, the female invariably presents 
the series in greater completeness ; though such of the 
spots as are present in the male are sometimes more 
distinct. ‘Thus in SV. hellica ?, the whole range occurs 
with the exception of S 9 (this being also the spot 
which is regularly small or absent in SV. callidice and 
S. daplidice 2), and the spots show a great tendency to 
fuse into a submarginal band, especially towards the 
costa of the forewing. In the male, however, all are 
absent from the hindwing, and S$ 9—11 also from the 
forewing; § 8 is distinct; S 7 hardly visible; and S 2—5 
are fused into a single spot which is more distinct than the 
corresponding patch in the female. 

The same submarginal series can be traced with equal 
certainty in the genus Belenois. Here again the series 
is as a rule more complete in the females, while the 
individual spots are more distinct m the males. This can 
be well seen for example in B. mesentina (Fig. 15), where 
the males show a submarginal band on the forewing which 
can without difficulty be resolved into a series of five spots, 
representing § 2—3,5,6, 7and 8. On the upper surface 
of the females the spots show a greater tendency to fuse 
with one another, and with the dark area of the apex and 
hindmargin ; they are nevertheless in most specimens 
recognizable on the upper surface, and nearly always 
fairly distinct beneath. Much the same is the case with 
the submarginal series on the hindwing; this is constantly 
present in a distinct form on the under surface of both 


256 Dr. Frederick A. Dixey on the 


sexes, and also to a variable extent on the upper surface 
of the male. The whole range (except the last member, 
S 20) occurs constantly on the upper surface in the 
female, but shows a strong tendency to fuse with the 
dark marginal band of the wing. In both male and 
female of this species the conspicuous and persistent spot 
S 8 occupies a prominent position, being pushed back as 
it were towards the centre of the wing, and so causing an 
indentation in the submarginal row. In B. severina the 
series is generally less well-marked than in B. mesentina, 
being to a great extent fused with the dark marginal 
band; S8 10 and 11, however, which are absent in the 
latter insect, are mostly visible in B. severina @. 
B. calypso is noticeable as having the present series 
particularly well-marked on the underside of the hind- 
wing ; in the female it is also well in view on the upper 
surface. Pinacopterya larima, again, has the series well 
developed on the hindwing, and recalls some species of 
Synchloe and Ganoris by the prominence given to § 8, 10, 
and 11. In several species of Belenois, as for instance 
B. teutonia 2 and B. coronea, the fusion of the spots of 
series S with one another and with the dark apical and 
marginal area is almost or quite complete. (See B. 
peristhene, Fig. 14). 

The genera Appias, Catophaga, and Hiposeritia pre- 
sent us in many of their members with the same series, 
more or less developed. In H. ‘lalage, for instance, S 
1—9 are fairly well marked on the forewing (most 
distinctly on the underside); while some spots of the 
series are also in most cases visible on the hindwing. 
In the greater number of species, however, the series 
is so fused with the dark apical and marginal area as 
to be hardly distinguishable (as in C. zamboanga® ), or is 
altogether obsolete (as in most specimens of A. nero). 
It is noticeable that the spot S 8, to which I have before 
drawn attention, is usually the last to disappear from the 
forewing; and that in many cases where S no longer 
exists as a separately recognizable series, the place of S 
8 is indicated by a prominent projection inwards of the 
fused marginal area (Figs. 9,10, 11). The underside of 
the hindwing often retains traces of the submarginal 
series when the upper surface of one or both wings has 
entirely lost them (C. lankapura ? and some specimens of 
A. nero). 


Phylogeny of the Pierinee. 257 


In the genus Delias the same assemblage of spots 
meets us again, though scarcely in a very definite form. 
The well known D. ewcharis (Fig. 5) shows the series on 
both surfaces of each wing, the spots being to a great 
extent merged with one another; the band formed by 
their fusion is nevertheless quite distinct, and its in- 
dentations correspond with the usual disposition of the 
spots on the wing in those species where they exist in a 
more discrete condition. ‘The series is also visible in D. 
hyparete (underside) ; and the same range of markings is 
present as a sinuous band in the Australian D. aganippe 
(best seen in the female, and on the underside); in less 
distinct form in D. harpalyce, and still recognizable, 
though now almost completely merged, in the closely 
allied D. nigrina. A comparison of the upper surface of 
the wings of the last named insects with those of other 
species of the genus Delias, such as D. belladonna (Fig. 4), 
D. pasithoe, and D. thisbe, serves to suggest the manner 
in which the submarginal series (in common with other 
features) may have arisen—namely, as a survival of an 
original dark ground-colour, which has become broken 
up, first into irregular bands and finally into spots, by 
the appearance and gradual enlargement of touches of 
a lighter colour in the spaces between the nervules. 
Further reference will be made to this point when other 
markings, whose history appears to be similar to that of 
the present series, come under consideration. 

Metaporia agathon (Fig. 7) presents the series in 
much the same condition as D. ewcharis, the band is, 
however, somewhat thicker and less defined. It is easy 
to trace the same feature through many species of the 
genera Pontia and Huphina; thus in P. soracta it is met 
with in the hindwing as a series of faintly-developed 
acute-augled chevrons, pointing outwards (as in Tato- 
chila). In the forewing, 8 3, 5—9 are easily visible. It 
is noticeable that S 8 is in this insect exceptionally 
small ; it retains, however, its usual tendency to break the 
line of the series by advancing further than the others 
towards the centre of the wing. In Pontia crategi the 
whole series has vanished ; but in Huphina coronis, H. 
phryne (Fig. 8) and others, it is well marked; the most 
persistent spots being, as in other genera, S 6, 8 and 10; 
and the same general differences between the sexes, in 
relation to the series, being observable here as in Synchloe 


258 Dr. Frederick A. Dixey on the 


and Ganoris. Indications of the series, more or less pro- 
nounced, can also be seen in H. timnatha, H. aspasia, 
H. judith, H. nama, and other species. 

The strong resemblance to WDelias belladonna borne 
by Prioneris thestylis is very probably due to mimicry, 
as suggested by Wallace (“ Pieridze of the Indian and 
Australian Regions,” Trans. Ent. Soc. Lond., 3rd series, 
iv., pp. 309, 383). But from the presence of traces of the 
submarginal series in other species of Prionerts (as in P. 
autothisbe and P. clemanthe), it seems likely that it is 
rather the retention than the origin of the submarginal 
series in P. thestylis that is attributable to this cause. 
Material for the production of a likeness of D. belladonna 
was no doubt already to hand in the existence of an 
homologous series of markings in both insects. Other 
species of Prioneris, not having the same reason for 
keeping close to the original design, have strayed away 
more or less widely from it. 

Mr. Wallace has also pointed out the agreement in 
some respects between Delias and the South American 
genus Huterpe (ibid., p. 344). If, with Mr. Butler, we 
divide Swainson’s genus Huterpe into Pereute, Leodonta, 
Catasticta and Huterpe proper, we shall find that while 
all the normally coloured species of the group are 
demonstrably linked with one another and with Delias 
by their general system of marking and scheme of 
coloration, two of the genera (Perewte and Leodonta) 
further suggest a near relationship with Delias by the 
peculiarities of their neuration; there being in these 
instances only three instead of four subcostal nervules 
in the forewing (Butler, ‘‘ Revision of the Genera of 
the Subfamily Pierine,” Cistula Hntomologica, vol. 1., 
1870, pp. 39, 40, etc.). The appearance of several 
species of this group of genera (notably Huterpe tereas) 
has been profoundly altered by mimicry; but putting 
aside such forms as these, we shall find no difficulty 
in tracing a common pattern throughout most of the 
remaining members of the group. ‘his pattern is not 
dissimilar from that which characterizes those species of 
Delias which we have seen to be probably the older 
(e.g., D. belladonna, D. pasithoe and D. thisbe). To this 
part of the subject I shall return later, at present it will 
be sufficient to point out the distinct presence of the 
submarginal series S in such examples as Catasticta 


Phylogeny of the Pierine. 259 


nimbice, C. anattis, C. bithys, and to a less degree in 
Leodonta dysoni and LL. tellane, ‘The close resemblance 
between the remarkable Pierine Hucheira socialis and 
C. bithys, in respect of the marking of the upper 
surface, may warrant us in identifying the corresponding 
portion of the wing in the former insect with the sub- 
marginal row S undoubtedly present in the latter 
(Migsyt, 2). 

The series § having now been traced through several 
genera, until it, so to speak, loses itself in such gene- 
ralised, and apparently primitive forms as those pre- 
sented by Delias belladonna, Catasticta bithys and Hu- 
cheira socialis, it will be advisable to return to the genus 
Synchloe, and seek thereabouts for a new starting-point 
from which to pursue the same series through its 
developments in a fresh assemblage of genera which 
have not at present been noticed. Such a starting- 
point may be found in a comparison of S. daplidice 9 
with the female of almost any species of the genus 
Colias. A short examination will show that the same 
submarginal series exists in the latter genus. A good 
species for the purpose is C. hyale (Fig. 2), im which 
both sexes exhibit the series distinctly. The consti- 
tuent spots broaden out towards the costa and tend to 
become fused with one another and with the black patch 
at the apex. ‘This tendency is carried further in many 
species of Colias, particularly in the males, where, as in 
C. edusa, marginal and submarginal markings are fused 
together into a deep dark border. The females, how- 
ever, preserve the spots of the submarginal series in a 
more discrete condition. In most species of Colias 8 is 
well developed on the under surface, the component 
spots being usually black on the forewing, and on the 
hindwing orange or pinkish. In a few species, such as 
C. phicomone and C. paleno, the series is obscured or 
absent. Meganostoma, which resembles Colias so closely 
in other respects, shows also the submarginal series on 
the under surface; usually as a row of somewhat faint 
pinkish dots, which are clearly identical with S in 
Colias. 

Through Meganostoma the passage is easy to Gonep- 
teryx and Amynthia. In G. rhamni and G*. cleopatra no 
vestige of S is to be seen on the upper surface; on the 
lower surface, however, of many of the males and most 


260 Dr. Frederick A. Dixey on the 


of the females, it is visible in the hindwing, and near 
the apex of the forewing, as a series of minute brownish 
dots; similar to, but as a rule even smaller than, those 
of Meganostoma. Rhodocera leachiana and Amynthia 
morula show exactly the same feature, the spots being 
dark brown or black, and a vestige of the series 
is present in A. clorinde. In the closely-allied genus 
Dercas, 8 is curiously modified into a brown or orange 
streak passing in an almost straight direction from the 
apex of the forewing to the anal angle of the hindwing. 
In D. wallichii, 8 8 is large and prominent; but the 
remainder of the series is scarcely visible on the upper 
surface, though easily recognisable below. 

In the genera Callidryas, Catopsilia, and their allies, 
the extent to which § is developed varies within wide 
limits. Callidryas philea > has already been noticed as 
presenting, on the upper surface of the forewing, in many 
specimens, a complete set from’S 1 to 8 11 (Fig. 22). In 
this, as in other species of the genera named, it is usually 
S 7 and not S 8 that breaks the line by advancing towards 
the centre of the wing. The series is not usually continued 
on the upper surface of the hindwing, but both wings show 
it tolerably well below. S is well developed on the upper 
surface of Catopsilia flava % (Fig. 21), and is visible to 
a variable extent on both surfaces (except the upper 
surface of the male) in C. catilla, Aphrissa godartiana, 
and others. In some of these insects S on the under 
surface forms part of the characteristic brown or pink- 
ish mottling of the wing; in others, as the male of C. 
phlegeus, it is almost or quite reduced to the condition 
of minute pink spots, like those of the genus Gonepterya. 
It is worth noticing that on the underside of Phebis 
trite, S takes in the forewing the form of a straight 
streak, not unlike that in Dercas lycorias, leading diago- 
nally across the wing. On the hindwing a similar streak 
appears, which, however, dces not represent 5 as the 
apparently corresponding streak in Dercas undoubtedly 
does, but is at least partly developed from another 
system of spots which will be noticed later. Attentive 
examination will disclose the existence of traces of the 
true S between the streak in question and the hind 
border of the wing. 

In Hebomoia the series is always at least partly visible 
on the forewings, and often on the hindwings as well. 


Phylogeny of the Pierine. 261 


The component spots are usually large and distinct, as 
may be especially well seen in H. glaucippe ? (Fig. 19). 
The same insect also shows the series tolerably well on 
the under surface, where it is in other members of the 
genus, as a rule, somewhat lost in the general mottling 
of the wing. 

S is often well seen on the underside of species be- 
longing to the genus Jvias, where it assumes a condition 
very like that which obtains in Colias. In I. marianne 
it is especially well developed, and the component spots 
on the hindwing are often furnished with pale centres. 
Some members of the series are frequently also visible 
on the upper surface, as in J. marianne 9, where S 6—9 
are conspicuous (Hig. 18). 

The genera Teracolus, Swains., Idmais, Boisd., and 
Callosune, Doubl., are united by Mr. Butler as Teracolus. 
In most of the species included under this head, the 
series now being discussed is distinctly visible, especially 
in the female and on the under surface. Good examples 
of the series are furnished by 7. danae and YT. etrida. 
In T. eucharis 3 it has mostly vanished from the upper 
surface; S 8, however, remains, aS in so many other 
instances, and is, moreover, in this insect often reinforced 
by the presence of S 13 and 14, which spots have been 
also seen to persist in our common species of Ganoris. 
The female of 7. regina possesses S in a well-developed 
condition on both surfaces, S 10 being especially con- 
spicuous. ‘he same series is visible to a less degree in 
the female of 7. cone. The males of both species show 
indications of S on the lower surface, but none on the 
upper. ‘The series may also be seen in a well-developed 
state in 7. hewitsonii and T. amata (Fig. 17). 

In Huchloe the series has almost disappeared. A relic, 
however, persists near the apex of the forewing in some 
species, best seen, perhaps, in H. tagis, H. ausonia, and 
EH. hyantis ; while H. reakirtit @ shows § on the upper 
side of the forewing very distinctly; S 8 indenting the 
line as in the genus Synchloe. A comparison of the 
under surface of the hindwing in H. cardamines and 
H. ewpheno with the somewhat similar system of marking 
in S. daplidice suggests also the probability that part of 
the green mottling in Huchloe represents the submarginal 
series. ‘Ihe same series is certainly visible on the upper 
surface of the hindwing in some specimens of H. lweilla 9. 


262 Dr. Frederick A. Dixey on the 


Certain species of Nepheronia—for example, N. jobcea 
? and N. valeria 2—show the series S in a form that 
strongly recalls the condition of the same series in 
Metaporia agathon. In both Nepheronia and Hronia, 
however, S is very apt to be merged, especially on the 
upper surface, with the dark marginal area. The female 
of H. leda (Fig. 23) shows the first few spots of the 
series on the upper side of the forewing in a condition 
resembling that in some species of Callidryas, which 
resemblance applies also to the lower surface, where S 
occurs in a modified form on both wings, traces of the 
same series being likewise visible in the male. 

Summary.—Most of the members of the subfamily 
Pierine possess, in a greater or less state of develop- 
ment, a submarginal series of dark spots. The genera 
Synchloe, Belenois, Tatochila, and some species of Colias, 
amongst others, present the series in a fair state of 
completeness. In other members of Colias, and in 
several genera such as [vias, Hebomoia, and Teracolus, the 
series is often less distinguishable, owing either to fusion 
of its constituent spots with each other and with the 
dark marginal area, or to suppression of some of their 
number. Relics of the series remain in Ganoris, Huchloe, 
and other genera, being especially persistent in the 
female sex; in Gonepteryx, Dercas, Callidryas, Pheebis, 
Catopsilia, and other allied genera, the series often 
undergoes curious modifications. The aspect of the 
series in such genera as Pontia, Huphina, Metaporia, and 
Nepheronia, suggests its development from portions of 
the original ground-colour left (by the formation and sub- 
sequent fusion of lighter patches in a dark or dusky area) 
as a dark band following the contour of the wing; and, 
finally, by an extension of the process, as a submarginal 
chain of spots. The possibility of such a mode of origin 
is well illustrated by the condition of the wings in many 
species of the Eastern genera Prioneris and Delias, and 
the Western Leodonta, Catasticta, and Huchevra. 


2. The dark marginal area or spots. 


In nearly all our common Pierine we find a dark 
patch at the apex of the forewing; better marked, as a 
rule, in the females than in the males. In G. brassice 


Phylogeny of the Pierinee. 263 


it is large and crescent-shaped, extending for some 
distance along both costa and outer margin of the wing. 
In G. rape 2 it is not as a rule prolonged very far on 
the outer margin, but it still reaches some way along the 
costa. In the male of the same insect it is generally 
small, and confined to the actual region of the tip. In 
neither of these species is there much indication of 
resolution, except that in most specimens of G. brassice, 
especially in the females, the hinder arm of the dark 
crescent shows an irregularity due to the massing of 
black scales about the marginal terminations of the 
nervules. But in G.napi (Fig. 16) resolution is generally 
well marked; and it becomes evident that the dark 
crescentic or triangular area of the tip is really formed 
by the fusion of the anterior members of the series §, 
which we have just been considering, with another 
series, which may be called M, occupying the actual 
margin of the wing, and consisting, in its fully resolved 
condition, of a row of dark spots, each of which is 
traversed by the peripheral portion of one of the ner- 
vules. It is noticeable that in this species the black 
marginal spots are often found extended, especially in 
the female, in the form of a powdering of dark scales 
along the course of the nervules towards the base of 
the wing. 

Turning to S. daplidice (Fig. 15), we find the consti- 
tuent elements of the apical patch still better shown 
than in G. nap. The marginal and submarginal series 
are always distinct, being separated from one another by 
portions of the general white colour of the wing, which 
take the form of a row of white spots on the dark apex, 
usually four in number. Hither of these two species 
serves well to illustrate an important difference between 
the spots of the two series, S and M. The spots of the 
former series occupy, as we have seen, the interspaces 
between nervules; and when fusion between adjacent 
members of the series takes place, it does so by an 
extension of dark scales across a nervule. On the other 
hand, each of the spots of the latter series is centred, 
not in an interspace, but around the peripheral portion 
of a nervule or nervure. ‘Thus the spots of the two 
series tend to alternate with one another, though this 
effect is to some extent interfered with by the frequent 
fusion of some of the spots with others of the same 


264 Dr. Frederick A. Dixey on the 


series. It is further noticeable that in both of these 
insects the marginal series M is usually continued in the 
form of a row of dots smaller than those helping to 
constitute the apical patch, but still distinct, along the 
outer border of both fore and hindwing. In S. daplidice 
? (Fig. 15), the submarginal series being also distinctly 
present on the hindwing, the alternate arrangement of 
the two series is in this situation very apparent. The 
constituent spots of series M may be numbered for con- 
venience, in the forewing 1—11, in the hindwing 12—20, 
one being counted for each nervure or nervule that 
reaches the margin; and the same allowance may be 
made as was suggested in the case of S for the occa- 
sional suppression of a fourth or addition of a fifth 
subcostal nervule (see p. 255). 

Many examples of the occurrence of this marginal 
series can be found among the near allies of the species 
just named, and in all of them there is a tendency 
towards the formation of a dark apical patch by the 
union near the tip of the forewing with the anterior 
members of the submarginal group 8S. In G. gliciria 
many members of the series are usually present, often 
tending in the females to form by fusion a broad mar- 
ginal band in both fore and hindwing. Other species of 
the genus Synchloe besides S, daplidice present M in a 
fairly well-developed condition. It is usually conspicu- 
ous in the female S. callidice, S. protodice and S. hellica, 
and to a less extent in the males of the same insects ; 
where it clings longest, as in the common species of 
Ganoris, to the region of the tip. Tatochila autodice and 
T'. theodice show M in very nearly the same condition as 
the species of Synchloe last named. 

Turning to the genus Oolias, we find that the mar- 
ginal series has undergone some modification. In the 
males it is often completely fused with the submarginal 
series, the two together forming a broad outer band to 
the wing, as in U. edusa 6. The same may also take 
place in the female, as in C. paleno; but in most cases 
the two series, S and M, are in the female more or less 
distinct, being marked off from one another by areas of 
ground colour which take the form of a row of light- 
coloured spots, not unlike those present in many species 
of Synchloe (see p. 263). In some instances, as in 
J. hyale and C. phicomone, this description applies also 


Phylogeny of the Pierinee. 265 


to the males. Most species show a strong tendency for 
the spots 8 8 and M 8—9, and, in a less degree, 8 10 
and M 10—11, to become fused with one another by the 
obliteration of the separating patches of pale ground 
colour. ‘The individual members of series M are usually 
more distinct from one another on the hind than on the 
forewing. Not infrequently they are fused into a com- 
plete band in both. On the lower surface M is in this 
genus rarely if ever visible; thus offermg a strong con- 
trast to 8. 

In Meganostoma the fusion of M with S is usually 
complete on the forewing; the females, however, of some 
species show traces of a separation; a few patches of the 
yellow ground-colour, like those in the females of many 
species of Colias, appearing in the midst of the dark 
border. In some forms, as M. philippa 9, these light 
patches become much enlarged, and the two series S and 
M are as distinct from one another as in any species of 
Colias. M. cesonia usually shows the spots well on the 
upper surface of the hindwing; and in most species of 
this genus, M is often visible beneath as a row of minute 
pinkish dots on the terminations of the nervules at the 
margin. 

In Gonepteryx, Rhodocera, Amynthia, and Dercas, the 
series is not as a rule strongly developed. G. rhamni 
usually shows on both surfaces a row of marginal dots, 
dark red or brown above and pinker beneath, at the 
outer extremities of the nervules, which are undoubtedly 
relics of M. These are best marked at the tip of tho 
forewing, where they often tend to run into a thin con- 
tinuous marginal line, and are generally more apparent 
in the female. The apical band is more strongly accen- 
tuated in Rhodocera leachiana, especially in the female, 
in which sex also M is often found on the hindwing in a 
fairly developed condition. ‘Traces only of the series are 
visible in the species of Amynthia; but in Dercas M is 
reinforced by the anterior portion of §S, and helps with it 
to form a conspicuous dark area at the region of 
the tip. 

The condition of M in Hebomoia needs no special 
remark, except that on the under surface its separate 
existence appears to be sacrificed to the exigencies of 
the protective pattern. 

In Callidryas, Metura, Phebis, Aphrissa, and Catop- 


TRANS. ENT. SOC. LOND. 1894,—PaRT II, (JUNE.) S 


266 Dr. Frederick A. Dixey on the 


stlia, the development of M varies. Most of the species, 
however, show the usual tendency of M to expand into 
an apical patch, either with or without reinforcement 
from S. The members of the series are generally 
reduced on the lower surface of the hindwing to a row 
of marginal dots; and the great characteristic of the 
present series, namely that it belongs to the nervules, 
and not, like the former one, to the interspaces, is kept 
up throughout. On the upper surface of Catopsilia 
flava 2? (Fig. 21), M and S are both well developed, and 
their relation to one another closely resembles that 
which obtains in Colvas. 

The same series is often present in the genera [vias 
and Teracolus ; beg, as usual, most constant in the 
females, and most prominent on the upper surface. It 
appears occasionally as a chain of spots, as in the hind- 
wing of many specimens of 1’. one 9, and some of 
T’. regina, but more often as a simple marginal band, as 
in J. marianne (Fig. 18). This band may be com- 
pletely fused with S, as in the hindwing of T. phisadia, 
or partially so, as in 7. amata. A plain marginal band, 
in the forewing expanding at the tip and more or less 
indented opposite the branching of the median nervure, 
in the hindwing sometimes breaking into a chain of 
spots, is also the usual condition of M in the genus 
Terias. 

In Nepheronia and Hronia M is generally present as 
a well-developed marginal band sometimes more or less 
fused with S, and often sending dark prolongations 
inwards along the course of the nervules. In some 
species, however, as N. thalassina and EH. leda, it is 
mostly confined to the apical region; in the latter 
indeed it is absent from the hindwing of the male 
altogether, and barely visible on that of the female 
(Fig. 28). The series in Huchloe is generally recogniz- 
able as a row of dots, sometimes fused at the apex with 
the anterior members of 8; but often, as in H. tagis and 
HE. ausonia, preserving its independent existence. 

The condition of M in the genera Metaporia and 
Huphina recalls that in Nepheronia; the constituent spots 
being generally fused into a marginal band from which 
dark offshoots run inwards, accompanying the course of 
the nervules. A similar description will apply to Pontia, 
though in this genus the dark markings are often greatly 


Phylogeny of the Pierines. 267 


reduced. P. soracta, however, shows M with sufficient 
distinctness; and even in P. crategi M itself is in some 
instances visible, while the dark prolongations along the 
course of the nervules is a well-known and conspicuous 
feature. 

Delias and Prioneris help to show us what is in all 
probability the earliest condition of the present as well 
as of the former series. ‘The upper surface of both fore 
and hindwing in the female of D. eucharis presents once 
more the familiar pattern of Metaporia agathon and 
Nepheronia gea ?. ILthas already been pointed out that 
there is an easy passage from this system of marking 
to that which we find in D. belladonna, D. pasithoe, and 
several other species; while the same features are un- 
doubtedly visible with but slight modification in the 
American genera Huterpe, Catasticta, Leodonta, and 
Eucheira (see p. 258). A comparison of the insects just 
named, to which may be added Prioneris thestylis and 
other species of the same genus, can hardly fail to suggest 
the probability of the origin of both M and S as portions 
of the original dark ground-colour marked off from one 
another with gradually increasing distinctness by the ap- 
pearance, enlargement, and confluence of whitish patches 
in the interspaces between nervules. ‘The alternate 
character of the two series S and M, when broken 
up into spots, is seen to be a consequence of the shape 
and position of the original dusky white touches that 
first bring them into view. These touches may be seen 
for example in D. belladonna (Fig. 4) to be divisible into 
two sets, which we may call marginal and basal, both 
occupying the central areas of the interspaces. Between 
the two sets is included the portion of ground colour cor- 
responding to 8, the constituent spots of which, when iso- 
lated, will necessarily have their centres in the interspaces. 
But the marginal set of light marks tends to reach by 
tapering extremities to the edge of the wing (see Fig. 5) ; 
so that the interposed marginal patches of unaltered 
ground colour, forming the first beginnings of M, are of 
necessity focussed round points in the nervules and not 
in the interspaces, And when in consequence of the 
further development and fusion of the white or pale 
markings, the marginal relics of dark ground colour 
become reduced to a row of isolated dark spots, these 
spots are, as we have already seen in S. daplidice (p. 263) 


268 Dr. Frederick A. Dixey on the 


and other cases, always traversed by the peripheral 
portion of a nervule or nervure. ‘This fact is well 
exemplified by M. agathina, M. poppea, and several 
other species of Mylothris (Fig. 12). 

The invasion of white, which is already considerable in 
D. eucharis 2, becomes still more predominant in the 
male of the same insect, where, however, the original 
ground-colour still persists, not only in the series S and 
M, but also in the form of the dark streaks accompanying 
the nervures and nervules, a feature which we have seen 
_ to be common in many other genera, especially Pontia, 
Huphina, Metaporia, and Nepheronia. 

Without leaving the genus Delias we can find several 
examples where the intrusive white has ousted the whole 
of the original ground-colour, except at the margins and 
apex. This is the case for instance in D. descombesvid, 
D. agostina &, D. coeneus $,and D. momea. In most of 
these species the dark marginal area is plainly composed 
of S and M inamore or less complete state of fusion, 
and it is noticeable here as elsewhere that the female 
almost invariably retains a larger proportion of the 
original ground-colour than the male. 

Many species of Belenois resemble the last mentioned 
species of Delias in the tendency towards fusion of the 
marginal and submarginal series. LD. peristhene (Vig. 14) 
is exceedingly like D. momea in this respect, and in B. 
teutonia 2? and B. coronea the fusion is still more com- 
plete. But in very many species of Belenois and 
Pinacopteryx, S and M are plainly distinguishable from 
one another; and in some, as in B. calypso (as also in 
Pinacopteryx capricornus), the resolution of both into a row 
of dots is complete. The formation of a series of rings 
along the margin of the hindwing (as in Synchloe calli- 
dice 2, S. protodice 2, etc.), by the union of the horns 
of the crescents or chevrons belonging to series 8 with 
the spots of series M, is a noticeable feature on both 
surfaces of many specimens of B. mesentina $ (see 
Fig, 138); and the identity of the whole arrangement of 
the submarginal and marginal series in Belenois with that 
in Synchloe becomes perfectly evident on a comparison of 
B. mesentina with S. johnstonw or 8. hellica. 

In Appias the two series when present are not sharply 
marked off from one another. In some species, however, 
S has almost or quite disappeared, and M may persist 


Phylogeny of the Pierinee. 269 


alone as a marginal band, showing more or less ten- 
dency towards resolution (Appias lyncida $, A. celes- 
tina 6, etc.). 

Summary.—The present, like the preceding series, is 
thus seen to be a relic of the original dark ground-colour. 
It is intercepted between the margin of the wing and 
a series of pale touches (best seen in certain species of 
Delias and Catasticta) that making their appearance in 
the submarginal region of the interspaces, more or less 
isolate, indent and divide up the dark marginal area. It 
may eventually disappear altogether (as in some speci- 
mens of G. rape 3), but usually persists on the forewing, 
at least, in the region of the apex. When present, it 
shows great variation in the extent to which it is resolved 
into separate spots, and also in the amount of fusion it 
undergoes with the neighbouring series 8. As a general 
rule it is, like the last-named series, more constantly 
present in the female sex, and more completely resolved 
in the male. 


3. The light series between 1 and 2. 


The region of pale ground-colour included between 
the two dark-coloured series S and M, becomes of 
necessity more sharply defined and circumscribed as the 
constituents of each dark series become fused together 
into bands instead of remaining as rows of spots. The 
tendency so often seen of the two dark series to become 
partially fused with one another, by the extension 
between them of dark lines following the course of the 
nervules, leads, when carried far enough, to the splitting 
up of the included pale area into a series of pale spots, 
which sometimes attain a very distinct and definite 
character, especially at the apex of the forewing. These 
pale spots have already been incidentally noticed during 
the discussion of the two series between which they lie ; 
it will not be necessary to do more in this place than to 
suggest that, for the purpose of separate reference, the 
letter I should be taken to represent the whole series, 
the possible constituents being numbered {—20, in corre- 
spondence with the dark spots of series S which bound 
them on the inner aspect. (See Figs. 1, 2, 5, 21, etc.) 


4, The discoidal spots. 
In none of our three common species of Ganoris does 


270 Dr. Frederick A. Dixey on the 


there exist a true discoidal spot. The nearest approach 
is to be found in G. napi, where the disco-cellulars of the 
forewing, together with the other nervules, are in many 
specimens accompanied by a sprinkling of black scales ; 
which in some instances are sufficiently numerous in this 
particular region to give almost the appearance of a 
black discoidal spot or patch. This may be best seen in 
specimens of G. bryonix, but dusky specimens of the 
ordinary G. napi will sometimes show the tendency 
almost equally well. The same tendency is carried a 
hittle further in Tatochila autodice, while JT’. theodice 
shows the corresponding marks in a condition of strong 
development, with a slight indication of a similar feature 
on the hindwing as well. Passing to the genus Synchloe, 
we find the discoidal spot invariably present, at least in 
the forewing; and in most species large and distinct, 
especially in the females. SS. daplidice (Fig. 15) has the 
spot well developed on both surfaces; on the lower sur- 
face it is more or less shaded with green. The same is 
the case with the somewhat smaller discoidal spot in 
S. callidice. Another example of a large and conspicuous 
discoidal spot is furnished by S. hellica. 

In Colias, a discoidal spot is nearly always present on 
each surface of both fore and hindwings. On the upper 
surface of the latter it is sometimes independently 
present and sometimes only shows through from 
beneath ; but it is seldom absent altogether. The spot 
on the forewing is in this genus, as in Synchloe, usually 
dark brown or black. It may be ringed or pupilled with 
orange; in the former case the orange is of a deeper 
shade than the general ground-colour. Hxamples of 
this may be seen in C. meadit, C. cunninghamii, and 
others. In C. fieldii and C. edusina the discoidal spot in 
the forewing is unusually large; in no species of Colias, 
however, does it overpass the boundaries of the sub- 
costal nervure in front and the third median nervule 
behind. It may be very small, as in C. barbara; or 
entirely absent, as in some specimens of OC. paleno and 
C. pelidne. On the lower surface it is often centred with 
white or silver. On the upper surface of the hindwing, 
the spot when present is not dark, but varies in different 
species from the palest yellow to a full orange. It is 
distinguished from the general surface of the wing either 
by its different shade of yellow or orange (as in C. hyale) 


Phylogeny of the Pierinee. 271 


or by its freedom from admixture with the darker scales 
that often give the hindwing in this genus a compara- 
tively dusky appearance (as in some specimens of C. 
edusa). In many species a small companion is visible 
in the interspace between the second subcostal and dis- 
coidal nervure. On the under surface the discoidal spot 
is much more distinct. Its centre is silvery white, and 
it is usually encircled with a brown or orange ring, 
which in some species is sharply marked off both 
internally and externally by thin marginal lines of a 
darker colour. The companion reproduces the discoidal 
spot on a smaller scale. It always occupies an inter- 
space, whereas the chief discoidal spot is situated upon 
the second disco-cellular nervule. (Fig. 20, D 4.) 

No doubt is likely to arise as to the identity of the 
discoidal spot in Synchloe with that in Colias so far as 
the forewing is concerned. The chief difference between 
them is that in the former genus the spot is less sharply 
defined and less regular in outline ; moreover in Synchloe 
it frequently reaches forward to the costa, and it may 
extend backward to the space between the first and 
second median nervule; whereas in Colias, as we have 
seen, it does not pass the limits set by the subcostal 
nervure and the third median nervule. But there can be 
no reasonable question that the spots are homologous in 
the two cases; and a comparison of the two genera 
further suggests the probability that in Synchloe we 
have an older, in Colias a newer and more specialised 
form of the marking. ‘This conclusion accords with 
what has been already said about the marginal and sub- 
marginal series of markings, which in Colvas, especially 
in the males, have departed somewhat further from the 
original type than in Synchloe; and it will be seen to 
receive further confirmation from what is to follow. 
With regard to the discoidal spot on the hindwing, 
although it is perhaps not at first sight clear that the 
markings in the two genera are really homologous, it is 
nevertheless quite possible, on an attentive examination 
of allied forms, to trace the steps by which the passage 
from one to the other is effected. In the hindwing of 
many species of Synchloe there is no indication of a 
discoidal spot at all; in others, however, a definite 
aggregation of black scales is visible about the disco- 
cellular nervules. In some species, such as S. protodice 9, 


272 Dr. Frederick A. Dixey on the 


the nervules, though surrounded with black scales, are 
themselves covered with white, or (on the underside) pale 
yellow scales; a light-coloured centre being thus provided 
for the dark discoidal patch. In the genus Vatochila 
this light-coloured centre acquires greater importance. 
On the under surface of 7’. autodice the white scales are 
seen not only covering the disco-cellular nervules, but 
also intruding some way upon the yellow of the general 
surface; in this species, however, the pale patch thus 
formed has no dark border. But there can be little 
doubt, on a comparison of 7’. autodice with Colias palzno, 
that the whitish area surrounding the second disco- 
cellular nervule in the one is homologous with the silvery . 
patch in the corresponding region of the other; while 
the identity of the discoidal marks in Tatochila with 
those in Synchloe, and of the same marks in C. paleno 
with those in other species of Oolias, is a matter of 
certainty. ‘The brown ring surrounding the silvery 
pupil on the under surface of most species of Colias is 
indicated in C. paleno, where its general appearance 
points to the probability of its origin from the dark 
scales that have already been noticed as gathering about 
the region of the disco-cellular nervules in Synchloe. 
In C. palzno itself, however, the central pale patch (as in 
Tatochila) becomes far more prominent than the sur- 
rounding dark border. An examination of the discoidal 
spot on the underside of the hindwing in Gonepteryx 
rhamnt 9 and Amynthia merula will disclose the same 
general arrangement of a pale area traversed by the 
second disco-cellular nervule, and surrounded by a ring of 
darker scales, that we have already seen to be character- 
istic of the spot in other genera. In these insects, how- 
ever, as in I¢thodocera leachiana, the spots, though clearly 
identical with those in Colias, are in a lower state of 
development and specialization. ‘Turning again to the 
discoidal spots on the forewing, we may at first sight 
hesitate to identify the bright orange spots in G. rhamnt 
or G. cleopatra with the black spots or patches in 
Synchloe and Colias. But in Amynthia clorinde we have 
what is unmistakably an intermediate form of the spot, in 
which the vivid orange of the one closely encircles the 
deep brown or black of the other, and from which either 
the Gonepteryx or Colias type of spot might be easily 
derived. Much the same is the.case with /’. leachiana; 


Phylogeny of the Pierinee. 273 


but in this insect the black markedly predominates. 
The discoidal spot in the forewing of R. leachiana is also 
noticeable as having a small companion, of similar 
character but with a greater proportion of orange, 
the root of the interspace between the subcostal and the 
first discoidal nervure. 

The discoidal spots in Dercas, when present, corre- 
spond closely with those in the genera just named. The 
spot on the forewing of D. lycorias is like that in 
G. rhamni; it is, however, less compact, and surrounds 
both disco-cellular nervules instead of the second only, 
as in the latter insect. The spot on the upper surface of 
the hindwing is scarcely visible, but those on both wings 
beneath nearly resemble the corresponding marks in 
R. leachiana. On the underside of D. wallichit the 
_ marks are of the same character, but reduced in size ; 
from the upper surface they are almost or quite 
absent. 

In Meganostoma the discoidal spots, both above and 
beneath, so closely correspond with those in Colias as to 
call for no special remark. In Callidryas, Metwra, Pheebis, 
Aphrissa, and Catopsilia, they present also the same 
general appearance. On the upper surface of the hind- 
wing, the discoidal spot is seen as a rule faintly or not at 
all; but on the forewing there is often a well-marked 
dark patch, enclosing either the second disco-cellular, 
as usually in Catopsilia thauruma and C. catilla ¢, 
or both disco-cellulars, as in Aphrissa godartiana 2. On 
the underside the form of a silvery circular patch, 
surrounded by a pinkish or brownish ring, is almost 
universally kept up. There ig in some species (as in 
Callidryas philea) a tendency for the discoidal spot in 
the forewing to divide into two parts, one for each disco- 
cellular nervule ; while in the hindwing the small com- 
panion which has been already noticed in Colias is nearly 
always present in the interspace between the discoidal 
and subcostal nervures. One or two species (as C. 
jlorella 2) possess a second companion within the cell. 
The present series, when fully resolved, seems thus to 
consist of five spots, two belonging to the two disco- 
cellular nervules in the forewing, the third belonging to 
the interspace between the subcostal and _ discoidal 
nervures in the hindwing, the fourth to the second 
disco-cellular nervule, and the fifth to the discoidal cell, 


274 Dr. Frederick A. Dixey on the 


also in the hindwing. I propose to designate the system 
as D 1—5.* 

The last-named genera exhibit a good deal of varia- 
tion in the size of the constituent spots of D, and 
also in the breadth of the encircling darker ring. In 
C. catilla $, the latter, spreading widely out from the 
silver centres, and being reinforced in the hindwing 
by a similar development from certain of the spots of 
series S, forms a conspicuous dark red patch on the 
under surface of both primaries and secondaries. A 
similar tendency is seen in many specimens of Callidryas 
thalestris and other members of that genus. In all these 
cases the border of the discoidal spots on the under 
surface is assimilated in colour to the general character 
of the darker mottlings that form the prevailing feature 
of the wing-pattern. It is interesting to observe, in the 
case of Phebis trite, that the straight diagonal streak 
already referred to (p. 260) as crossing the under surface 
of the hindwing is made up of elements belonging to 
both series S and D, as follows: The part between the 
two subcostal nervules is S 14; that between the sub- 
costal and discoidal is D3; the next, where the streak 
intersects the second disco-cellular, is D 4; it then skips 
a space, and reappears between the median and sub- 
median nervures as § 19. That the homology of the 
different parts of this line has been correctly stated will 
be evident from a comparison of P. trite with C. eubule 
and C, thalestris, in which species the elements of the 
streak are visible in a separate condition. It is in P. 
trite particularly worthy of note that in those interspaces 
where D 3 and 4 form part of the streak, the corre- 
sponding members of §, viz., S 15, 16, and 17 (which 
are not wanted for this purpose), are visible well to the 
outer side of the streak; whereas in the other inter- 
spaces there are no marks between the streak and the 
margin, the members of S having been, so to speak, 
used up for another purpose, and having thus caused an 
apparent interruption of the usual submarginal pattern. 
In the interspace between the first and second median 


“ This use of D is open to the objection that the same letter 
has also been used for an entirely different set of markings in the 
Nymphalidz (see Trans. Ent. Soc. Lond., 1890, p. 91). But I do 
not think that this circumstance is likely to give rise to any 
practical difficulty. 


Phylogeny of the Pierine. 275 


nervules, S 18 appears as a submarginal spot of the usual 
kind. Now, inasmuch as no member of series D ever 
occupies this interspace, there is no material at hand for 
continuing the streak ; and, carrying the eye up to the 
apex of the interspace, where the streak should cross, 
we find it, in fact, deficient at that place.* This may 
seem a small point, but it is of interest as illustrating 
the absence of anything that might be called violent or 
arbitrary in the evolution of these various patterns, in 
which, indeed, the closer the examination the more 
difficult it is to find anything effected per saltwm. 

In Jzias the condition is again much the same as in 
Colias ; the discoidal spot of the forewing, however, is 
often united on the upper surface with a dark band 
running across the wing from the costa to the anal 
angle. A similar feature occurs in Hebomoia glaucippe 
(Fig. 19) and other species of the same genus. On the lower 
surface the discoidal spot varies considerably in size, and 
in a few cases (as sometimes in I. marianne) may be 
absent altogether. When it is large, as in other 
specimens of I. marianne, it is frequently centred with 
white. In the hindwing the series is absent from the 
upper surface, and represented below only by D 4, which 
may also be white-centred, or may exist as a simple 
black dot of varying size. 

In Teracolus, D is usually well marked. In some 
species, such as 7’. vesta, T’. hewitsonii, and T’. puellaris, 
it occurs on the forewing as a conspicuous black patch 
covering both disco-cellular nervules, and usually merged 
anteriorly in a dark or dusky band which extends along 
the costal border. In other species, as 7’. evanthe, T’. ione, 
T. regina, etc., it takes the form of a circumscribed black 
spot, sometimes minute or absent. On the lower surface, 
the scales on and surrounding the disco-cellular nervules 
are often whitish, thus giving Da pale centre. D in the 
hindwing, when present, is usually small; on the under- 
side of some species, as 7. danae, T. evippe, and 1". 
omphale, it is centred or shaded with orange ;—this 
tint belonging primarily to the disco-cellular nervules, 
and spreading more or less widely from them. 


* As above noted (p. 260), the corresponding streak in the fore- 
wing of P. trite, and the somewhat similar feature on both wings 
of Dercas lycorias, are entirely formed out of series 8. 


276 Dr. Frederick A. Dixey on the 


In Terias and Sphxenogona, D is constantly absent 
from the upper surface, but is usually visible below, 
where it may appear as a small black dot or pair of dots 
in each wing (7. lisa), or as a group of more or less 
regular narrow-bordered rings (7. hecabe and T. floricola), 
or in various intermediate forms (7’. deva and S. gratiosa). 
The transition from the condition of D in Colias to that 
in Terias is well indicated by Xanthidia nicippe, most 
specimens of which correspond pretty closely in the 
markings of the lower surface with the latter genus ; 
while the upper surface of the forewing bears a discoidal 
spot much like that in some species of Colias. In the 
closely allied genus Pyrisitia, D has altogether disap- 
peared (see P. proterpia). 

The genus Huchloe shows the discoidal spot on the 
forewing in a well-marked condition (Fig. 24). It may 
exist as a distinct spot, as in H. cardamines, usually 
larger in the female; or as a more or less quadrangular 
patch reaching up to the costa, as in H. lucilla. On the 
underside it is often white-centred, as in H. belemia and 
others. On the hindwing, D is in many species of Huchloe 
not visible at all. In some, however, it appears as a 
small, at times very minute, patch of black scales sur- 
rounding the second disco-cellular nervule (some speci- 
mens of H. genutia and H. cardamines); in others, as 
i. charlonia and EH. lucilla, it takes the form, on the 
under surface, of a pale circular patch, closely resembling 
D 4 of Colias paleno in the same situation. 

D in Belenois is usually well marked on the forewing, 
as a rule enclosing both disco-cellulars, and often 
prolonged anteriorly to join the outer extremity of a dark 
costal band ; this is especially the case in the female, and 
on the under surface in the male, as in B. mesentina, B. 
teutonia, B. calypso, etc. The spot is sometimes small 
and quite distinct, as in B. creona, D 1 being absent ; 
sometimes, on the other hand, it is large and almost or 
quite merged in the general dark field of the outer 
portion of the wing, as in B. coronea. In the hindwing 
there may be no discoidal mark at all, asin B. creona; 
or all the nervures may be covered with dark scales 
(as generally on the under surface of B. mesentina), 
the disco-cellular nervules not being specially dis- 
tinguished; or there may be a definite discoidal spot, 
composed of D4, as in B. mesentina 9, upper surface ; 


Phylogeny of the Pierine. PY A 


and in B. calypso, under surface of both sexes and upper 
surface of female. 

In Appias and OCatophaga a discoidal spot can scarcely 
be said to exist; but in Hiposcritia and Pieris various 
members of the series D are often well marked. H. 
lalage ¢ (Fig. 11) has a conspicuous discoidal spot on 
both surfaces of the forewing, formed by D 2; and on 
the under surface of the hindwing there appears a minute 
representative of D4. In H. pandione, D 1 and 2 com- 
pose a large spot which joins the dark costal band; while 
on the under surface of the hindwing, D 4 is sometimes 
indicated by a paling of the first disco-cellular nervule 
and the immediately surrounding area. In Pieris the 
forewing often shows a conspicuous discoidal spot, formed 
by D 1 and 2 or by D 2 alone; the hindwing as a rule 
possessing only a small dark representative of D 4 on the 
under surface, which in P. buni merely touches the 
second disco-cellular, but in P. pylotis encloses it. 

In Pontia and Huphina (Fig. 8), D in the forewing 
appears generally as a mere darkening of the wing in the 
region of the disco-cellular nervules, more pronounced 
than the darkening of other nervures which is common 
in these genera, but not forming a defimite spot. In the 
hindwing it is much less distinguishable or even absent 
(P. crategi, P. soracta, H. phryne, H. timnatha, etc.). 
In Metaporia agathon (Fig. 7) the same darkening of the 
disco-cellular area occurs, which in M. phryxe takes the 
form of a definite and conspicuous patch. A like condition 
is seen in JN. gcea, and other species of Nepheronia. 

The correspondence in pattern of the genera Delias and 
Prioneris with those just discussed, already referred to on 
p. 267, applies to the present as well as to other features 
in the marking. JDelias, moreover, shows in many 
species that paling of the disco-cellular nervules them- 
selves, and sometimes of the immediately surrounding 
area in the midst of the dark discoidal patch, which we 
have seen to occur in Synchloe and other genera, and 
which in Colias, Ivias, and their allies leads to remarkable 
modifications in the general appearance of the discoidal 
spots (see Delias aganippe, D. descombesi, D. crithoe, 
etc.). In D. belisama ¢, the nervules remain dark, 
though there is a surrounding pale area. 

Nothing special needs to be said about the genera 
Euterpe, Catasticta, Leodonta, and Euchetra. Their 


278 Dr. Frederick A. Dixey on the 
general correspondence in pattern with the old-world 
genera last named, which has been already referred to, 
involves a similar condition of the present series D. In 
none of them does any member of D assume the condition 
of a definite circumscribed spot; but in nearly all, the 
series exists as a portion of the original dark ground 
colour between the touches of intrusive white or yellow. 
In C. toca, D on the underside of the hindwing consists 
of an irregular dark ring with a yellow centre traversed 
by the disco-cellular nervules, this representing an early 
condition of the homologous ocellus in Oolias. 
Summary.—This series also, like those already treated 
of, represents a local survival of the original dark ground- 
colour. Appearing in the first place as an ill-defined 
accumulation of dark scales around the disco-cellular 
nervules, it takes the form of definite spots as the in- 
vasion of white or other pale ground tint progresses. As 
in the case of the other series, resolution is generally best 
marked in the males; the females showing a greater 
tendency to adhere to the ancestral condition. When 
fully resolved, the series consists of five spots, two for 
the disco-cellular nervules in the forewing, and one for 
the second disco-cellular in the hindwing, with two extra 
spots, also in the hindwing. In many instances the 
disco-cellular nervules, with a varying amount of neigh- 
bouring ground-substance, may appear as pale streaks or 
patches in the midst of the primitive agglomeration of 
dark scales; and in the further development of the 
pattern it may happen that either the pale or the dark 
markings alone persist; or both may be retained and 
worked up as in species of Colias, Callidryas, Ivias, etc., 
into a system of more or less elaborate ocelli. 


5. The markings on the underside of the hindwing. 


a. The yellow precostal streak.—A. constant feature in 
our common species of whites is a deep yellow or orange 
streak which occupies the margin of the precostal space 
on the underside of the hindwing, and is clearly dis- 
tinguishable from the pale yellow of the general ground- 
colour. This yellow precostal streak is conspicuous in 
other species of Ganoris, as G. gliciria, and may also be 
seen in S. daplidice, S. hellica, and many other members 
of the genus Synchloe. It is, I believe, never found in 
Pontia crategi; in the closely allied P. hippia and P. 


Phylogeny of the Pierinee. 279 


soracta, however, it is well marked. Metaporia agathon 
and M. phryze, which resemble the last-named species 
in other respects, also show a deep yellow mark in the 
same situation; but in this instance it occurs only as a 
spot bounded outwardly by the precostal nervure, and is 
not prolonged as a streak along the costa.* Many 
genera present this mark in much the same condition as 
the species of Ganoris first referred to; of which good 
examples may be seen in Nepheronia thalassina, N. argia, 
the curious Herpenia eriphia, Pinacopteryx pigea and P. 
capricornus, Mylothris chloris and M. agathina (deep 
orange in these), T'atochila theodice, Glutophrissa mar- 
garita, Appias zelmira, Phrissura sylvia, Daptonura 
lycimnia, Teracolus ione, T'. regina, T. eris, etc. 

Looking back to Synchloe hellica, we find that the 
yellow precostal streak forms in that insect only one of a 
series of yellow marks, one or two of which partially 
occupy each interspace. A similar condition obtains in 
many species of Belenois; see for instance B. mesentina 
and B. tewtonia. On the underside of the hindwing in 
these species, the dark lines following the course of the 
nervures, and the dark markings belonging to the various 
series above described, are laid down upon a ground 
colour which is usually white or pale yellow, but which in 
many specimens is relieved here and there by streaks and 
touches of a deeper yellow or orange. One of these 
streaks occupies the costa, and is plainly identical with 
the yellow precostal streak in Synchloe, Ganoris, and the 
other genera named. Although in the species of Belenois 
just referred to, as in several others, this streak is easily 
seen to form but one member of a series, it is never- 
theless more constant and persistent than any of its 
companions. ‘This is evident in those not uncommon 
specimens of B. mesentinat in which the general 
ground colour on the underside of the hindwing 
takes on so deep a yellow tinge as to approach that of 
the yellow streaks and touches. In these cases all or 
some of the latter set of markings may cease to be 
separately recognizable; the precostal streak, however, 
is always the last to disappear. In certain species of the 


* A specimen of MW. agathon in Coll. Brit. Mus. has, however, a 
few orange scales in the outer division of the precostal space. 
+ I.e., the form known as B. auriginea. 


280 Dr. Frederick A. Dixey on the 


present genus, as B. gidica, the ground colour remains 
pale ; but the precostal streak is the only member of its 
series present. In others‘again, as Bb. coronea, the black 
spots of S exist in a state of enlargement and fusion, 
while the orange touches occupy nearly the whole of 
the spaces that are left. ‘The light spcts of series I (see 
p- 269) between S and M, which in B. teutonia are only 
touched with orange, are entirely of that colour in most 
specimens of B. coronea. In B. peristhene the amount 
of light ground colour is smaller still, but the orange 
costal streak and orange series I are present as in B. 
coronea. Both of these insects, B. peristhene in especial, 
suggest at once a comparison with certain members of 
the genus Velias, particularly D.nysa and D. momea. In 
these two species the yellow precostal streak and the 
yellow or orange series I are unmistakably present. In 
D. nysa, as in B. peristhene, these constitute the only 
relief to the general dark colour of the wing; while in 
D. momea § is partially defined by the presence of pale 
touches on its inner aspect in addition to the spots of 
series I along its outer margin. 

While the species named suggest a passage in respect 
of this pattern of the underside of the hindwing between 
Belenois and Delias, there is not wanting evidence of 
the same import in the case of other species in which the 
blackening of the hindwing is a less conspicuous feature. 
On the underside of D. belladonna we cannot fail to 
recognize the counterpart of such a hindwing as that of 
B. teutonia ; the dark and light areas occupy with slight 
modification the same relative positions, and the same 
spaces of pale ground-colour that are in B. teutonia 
touched up with deep yellow, are wholly or partially of 
the like colour in D. belladonna ; among these, of course, 
being the precostal margin. ‘I'he basal portion of the 
hindwing in such species as D. ewcharis and D. hyparete 
shows a more general yellow tinge than in D. belladonna ; 
but in both of these, as in other allied species, the 
deeper-coloured precostal streak is perfectly recognizable. 
In very many species of Delias we meet with a new 
feature, much of the ordinary yellow being displaced by 
a brilliant red; but the former colour still displays a 
tendency to cling to the region of the precostal space, as 
may be seen in D. thisbe ; where, however, the precostal 
yellow hardly reaches further outwards than the pre- 


Phylogeny of the Pierinee. 281 


costal nervure. In the very similar D. pasithoe it has 
been driven even from this position. 

Prioneris exhibits a condition very similar to that of 
Delias. The underside of P. thestylis closely corre- 
sponds with that of D. belladonna, and the resemblance 
applies to the precostal streak no less than to other 
features. In some species of Prioneris, as C. clemanthe 
and P. autothisbe, the precostal yellow is again ap- 
proached, and sometimes dislodged, by a greater or less 
amount of bright red, as in D. pasithoe, D. thisbe, 
D. crithoe, and other members of the former genus. 

Leaving the red patches and other features of the 
pattern of this region in Delias and Prioneris for dis- 
cussion at a later stage, we may pass on to the 
American genera Huterpe, Pereute, Leodonta, and Catas- 
ticta. In the two last-named the arrangement of light 
and dark markings corresponds generally with that in 
Delias and Belenois, and similar touches of deep yellow 
are mostly present in the same relative positions. One 
of these, usually to be seen on the costal margin, is 
clearly homologous with the precostal streak in the 
insects already referred to; in many specimens of C. 
mimbice, C. semiramis, C. toca, and others, it bears very 
much the same appearance as in the old-world genera 
above mentioned. It may be noticed in passing that 
in many instances, such as L. zenobia, L. dysoni, and 
C. anaitis, the yellow precostal streak is found sharing 
the precostal space with a greater or smaller amount of 
brilliant red, as we have seen to be often the case in 
Delias and Prioneris. The same struggle between 
yellow and red is visible in the genera Huterpe and 
Pereute. In H. tereas the space is shared, the red keeping 
to the inner, and the yellow, when present, to the outer 
side of the precostal nervure. In P. charops the space is 
usually all yellow; in P. auwtodice and P. leucodrosvme it 
is all red. In these two genera the other yellow marks 
are reduced in number or completely abolished; P. 
charops, however, generally has a conspicuous yellow 
patch between the costal and subcostal nervures which 
undoubtedly belongs to the series. 

In the genus Pieris, which is also American, some 
species, such as P. locusta and P. habra, show on the 
under surface of the hindwing a pattern which is 
evidently only slightly modified from that seem in many 


TRANS. ENT. SOC. LOND, 1894.—PART II. (JUNE.) a 


282 Dr. Frederick A. Dixey on the 


species of Catasticta ; and in these also the red or orange 
of the costa is a prominent feature. In other species of 
Pieris a farther alteration has taken place, and the 
markings have been swept from the basal portion of the 
wing (as in P. demophile), or away altogether (P. bumiz). 
In most, however, if not all of these cases, the precostal 
streak persists, again taking a yellow rather than red or 
orange colour; and, in the absence of other yellow or 
well-defined dark marks on the wing, it comes to 
resemble very closely the corresponding feature in so 
many species of Synchloe and Ganoris. 

Summary.—The facts adduced in the course of the 
foregoing remarks seem so far to allow of little doubt 
as to the history of the yellow precostal streak. It is 
evidently a survival of a series of yellow or orange marks 
which is found, in many genera of both eastern and 
western Pierinze, on the underside of the hindwing, 
partially or wholly oceupying the paler areas left between 
the relics of the original dark ground colour. The 
greater number of these marks may disappear with a 
general lightening of the wing, or the whole wing may 
become so yellow as to render them almost or quite 
indistinguishable ; but the particular yellow streak that 
occupies the margin of the precostal space is more per- 
sistent than any of the others, and may remain, as in our 
common species of Ganoris, after every other character- 
istic of the ancestral marking has departed. It will be 
seen later that this account of the precostal streak, 
though no doubt correct as far as it goes, in all pro- 
bability needs supplementing. 

b. The red basal patch.—In both British species of 
Ooclias, there occurs at the base of the hindwing, on the 
under-surface, a pinkish patch, which occupying the 
apices of the cell and of the median and submedian 
interspace, and also in most cases asmall area at the root 
of the precostal space, is often prolonged for a short 
distance along the middle of the cell. This patch, which 
is almost always present throughout the genus Colias, is 
found in most if not all species of Meganostoma, and 
also very commonly in Catopsilia, Callidryas, Phebis, 
Aphrissa, and Metura. It is present, too, in Dercas, 
Gonepteryx, Rhoducera, and Amynthia; in these, however, 
it does not as a rule extend far from the body itself. 
Beyond the limits of these closely-allied genera this 


Phylogeny of the Pierinee. 283 


special feature does not often occur in a very clearly- 
marked condition; it is, however, present in Xanthidia 
nicippe, and in several species of Terias and Sphcnogona, 
as 7’. agave and WS. mexicana; it may also be seen 
occasionally in Teracolus, as in T’. antevippe, T. ewpompe, 
and (sometimes) 1’. erts, assuming in this genus a some- 
what diffuse appearance. In some species of Appias, as 
A. hombroni (occasionally), and A. lyncida, as also in 
Herpexnia eriphia, there are deep yellow marks near the 
root of the wing, which are probably homologous with 
part of the patch now under discussion; and in the 
genera Belenois and Tatochila touches of deep yellow 
often occur at the base of the median space, though not 
in the cell, which appear to correspond with the marks 
just referred to in Appias and Herpxnia (see B. mesentina 
and J’. autodice). ‘These yellow touches in Belenois are 
undoubtedly members of the series spoken of on p. 279, 
and so far it would seem that the well-marked pink patch 
at the root of the cell in Colias, Gonepteryzx, etc., is at 
least in part developed from a portion of that series, of 
which another member has given rise to the yellow 
precostal streak. It is certain, however, that other 
factors besides the series now indicated are chiefly, 
though probably not entirely, answerable for the com- 
position of the pink basal patch in Colias; and it is at 
least possible, as will presently be seen, that the pre- 
costal streak itself owes something to their assistance. 
These other factors are the red basal areas to which 
reference has already been made, and which will now be 
examined in greater detail. 


c. The red basal areas. ‘To begin with the under- 
side of Delias pasithoe, we find the base of the hind- 
wings occupied by a bright red patch forming about 
a quarter of a circle on each wing, the body itself and the 
immediately surrounding area of the wings being black. 
The spaces affected by the red patch are six ; namely, the 
precostal, costal, subcostal (or cell), median, submedian, 
and internal—all those in fact whose apices reach to the 
base of the wing. In D. thisbe there occurs a similar 
red patch, occupying nearly the same spaces. In this 
insect, however, there is no red in the precostal space, 
which is occupied (as above noticed, p. 280) by the ordinary 
yellow precostal streak ; while the red patch, which in this 
butterfly is visible on the upper as well as the underside 


284. Dr. Frederick A. Dixey on the 


of the wing, includes thé apex of the space between the 
first and second subcostal nervules in addition to those 
above enumerated. D. egialea shows the same condition 
as D. pasithoe; but in D. crithoe, otherwise similar, the 
precostal space is black. In D. descombesi and D. belisama 
the red is confined to the costal interspace (7.e. the space 
between the costal and subcostal nervures); while in 
D. coeneus a submarginal set of roughly-defined red spots 
or dashes continues round to the anal angle the series 
begun by a red mark in the costal interspace, corres- — 
ponding with that in the two preceding species, ‘The 
submarginal series which, so to speak, becomes infected 
by the red coloration, is undoubtedly that spoken of above 
as series I (see p. 269). ‘This latter feature of the Pierid 
pattern, somewhat indefinite in D. ccenews, becomes much 
more distinct in such species as D. nigrina, D. harpalyce, 
D. aganippe, D. eucharis, and D. argenthona. The latter 
of these has another well-marked red spot surrounding 
the second disco-cellular nervule, which represents a 
diffused reddish areain D. duris, and amore circumscribed 
patch in the same situation in D. cceneus. In D. hyparete 
S is going from the underside of the hindwing; in 
D. hierte it is gone ; but in both cases the red spots of I 
remain in their full development. D. mysis, D. isse, 
D. echo, and other species exhibit the same feature in a 
more or less conspicuous manner. 

The same red coloration is visible, as we have seen, on 
the underside of many species of Prioneris in the form of 
a basal patch ; and in one species (P. sita) it spreads to 
the submarginal spots of series I, exactly as in UD. 
eucharis. In P. autothisbe the red is found in the costal 
and subcostal interspaces ; in P. clemanthe and P. vollen- 
hovit only in the inner division of the precostal space, 
whence it spreads inwards upon the body. 

In the American genera Huterpe, Pereute, Leodonta, 
and Catasticta, the red is usually present; but as in 
Prioneris and the pasithoe group of Delias, it is mostly 
confined to the basal portion of the wing.* In H. 


* An exception to this rule is furnished by Muterpe tereas and 
FE. critias, in which species the area of light ground colour 
immediately internal to that region of the wing which corresponds 
to the submarginal series 8, is on both surfaces more or less 
deeply tinged with the same red colour as that of the basal 
patches. ‘his feature is due to mimicry, the two species named 


Phylogeny of the Pierinee. 285 


tereas, Hi. critias, C. bithys, and C. anaitis, it most com- 
monly occurs in the inner division of the precostal and 
the internal space; in Leodonta zenobia and L. dysont, in 
the precostal and costal. In L. tellane, besides the bright 
red patches of the precostal and costal spaces, scales of 
the same colour occur in the apices of the subcostal, 
submedian, and internal spaces—of all, in fact, except 
the median. Pereute leucodrosime has the precostal and 
costal red patches; so also has P. swainsonii; but in 
P. charops these spaces are occupied by yellow streaks 
with no accompaniment of red. 

Even in the genus Dismorphia, profoundly altered as 
it has been by mimicry, the same marks occur. D. melia 
has a conspicuous red spot in the inner division of the 
precostal space; while D. astyocha, D. prawinoe, and 
others possess a very clear representative of the same 
spot, with another one in the median interspace. ‘The 
colour in these latter species is seldom so bright as in 
the first-named ; it displays, in fact, a tendency to become 
assimilated to the chestnut tinge of other portions of the 
wing ; the spots are nevertheless quite recognizable, and 
undoubtedly correspond with those in the former genera. 
In some species, such as D. nemesis, there is no red, but an 
indication of the yellow precostal streak is plainly visible. 


offering a very close resemblance to the females of Papilio poly- 
metus and some of its allies, in bringing about which resemblance 
the red areas of the hindwing take a considerable share. It is 
interesting to observe that good as is the general effect of the 
imitation by these Pierines of the Papilio pattern, the imitative 
process has its limitations, and is seen on a strict examination to 
fail in more than one particular. Thus, (1) the Pierine is able to 
produce an excellent representation of the Papilionine reds and 
yellows, but apparently it never reproduces the metallic blues and 
greens, of which touches are found in the males of several of the 
mimicked species; (2) the costal and submedian spaces in the 
Pierine invariably retain their yellow patches on the underside ; 
these are never yellow in the Papilio ; (3) the red basal patches 
on the underside of the Pierine give just the same general effect as 
similar patches on the Papilio ; but a close scrutiny will reveal the 
curious fact that the patches of the Pierine belong always to the 
wing, and those of the Papilio, in almost every instance, to the 
body. The wide distribution of the red basal patches among the 
Pierine forbids us to suppose that they were evolved for the pur- 
pose of mimicry in these few species ; but it is worth noting that 
their presence affords material ready to hand for a sufficiently 
deceptive though not absolutely exact copy of a conspicuous 
Papilionine feature. See also note on p. 298. 


286 Dr. Frederick A. Dixey on the 


In those species of Pieris (such as P. habra and 
P. locusta) which present on the underside of the hind- 
wing a pattern only slightly modified from that of 
Leodonta and Catasticta, the red basal marks are clearly 
to be distinguished. A bright red patch characterises 
the inner division of the precostal space, and in most 
specimens is also prolonged into the outer division, while 
the apices of the median and internal spaces are likewise 
provided with conspicuous red touches. Comparing 
the underside of P. locusta with that of any of the 
American mimetic forms of Mylothris, we cannot well 
resist the conclusion that the central horizontal red or 
chestnut band in the latter species represents the touches 
of red in the median and internal spaces of P. locusta. 
The relation is best traced in the case of M. lypera, in 
which insect the part played by the yellow streak of the 
costal interspace in the production of the mimetic 
pattern is also clearly to be seen; but other species, such 
as M. lorena and M. pyrrha, whose mimicry of Heli- 
conine forms is further advanced than that of M. lypera, 
still show the origin of the central red band almost 
equally well. In Hesperocharis hirlanda a somewhat 
similar effect is produced by slightly different means. 
The basal red is here confined to the precostal and 
internal spaces, not being found (as in Pieris) in the 
median. The horizontal red band is therefore not 
central but costal, its mnermost extremity being con- 
stituted not (as in Mylothris) by the red patch of the 
internal space, but by that of the mner division of the 
precostal. The whole of the precostal red, which is 
absent in the three species of Mylothris just referred to, 
is prominent in H. hirlanda. Although these differences 
involve a change in the relative position of the horizontal 
bands of red and pale yellow in the latter species, yet 
the general resemblance of the underside to that of 
M. pyrrha ¢ is considerable, and is probably quite 
enough to tell perceptibly in the insect’s favour. Other 
species of Hesperocharis, which are unaffected by 
mimicry, show the basal marks in the same position, 
but, as a rule, not very definitely, except at or near the 
root of the precostal space. Here there generally 
occurs an orange spot which partly represents the red 
precostal patch in H. hirlanda (see H. nereis, H. mar- 
chalii, and H.anguitia), but which in some species (as 


Phylogeny of the Pierine. 287 


H. erota) can be distinctly seen to belong to the apex of 
the median interspace as well as to the root of the pre- 
costal.* A similiar orange spot is visible in_ several 
species of Pieris, such as P. buniw and P. thaloe; and 
of Leptophobia, as L. elodia and L. elewsis. In the last- 
named insect again it can generally be made out that 
the orange spot does not belong entirely to the root of 
the precostal space, but is furnished with a constituent 
from the apex of the median interspace as well. ‘This 
double character of the spot is much more evident in 
DL. tovaria; and on referring once more to almost any 
species of Colias, the conviction forces itself strongly 
upon us that in these basal marks in Leptophobia, 
Hesperocharis, and Pieris, and so in Huterpe, Leodonta, 
Catasticta, and consequently even in JDelias and 
Prioneris, we see represented the chief material out of 
which the pink basal patch in Colias, Gonepteryx, Catop- 
silia, and their allies is constructed. It is probable that, 
as above suggested, another element may enter into the 
composition of the basal patch in these genera, namely 
one or more members of the series of yellow streaks; a 
conclusion which seems to be somewhat favoured by the 
condition in Belenois, Herpxiia, and Appias; but the 
two series are distinct enough from one another in such 
genera as Catasticta and Leodonta; and an unnamed 
species of Hesperocharis in the British Museum shows, still 
more plainly than those above mentioned, the co-existence 
of a relic of the basal red in the situation of the pink 
patch in Colias, with a series of yellow streaks in the 
interspaces like those of Catasticta and Belenois. 

Summary.—Witk regard to the whole assemblage of 
red and yellow marks on the underside of the hind- 
wing, the following appear to be the most probable 
conclusions :— 

The yellow precostal streak so commonly seen in 
butterflies of this group, including our common British 
species of Ganoris, as also the pinkish patch at the apex 


* In order to avoid a possible ambiguity, I may here mention 
that I regard the “apices” of the interspaces as pointing towards 
the “base” of the wing. When a precostal nervule is present, I 
speak of an “outer” and “ inner” division of the precostal space, 
the nervule forming the line of demarcation. By the “ raot” of 
the precostal space, I mean the part immediately adjoining the 
body. 


288 Dr. Frederick A. Dixey on the 


of the cell, of which our native species of Colias and 
Gonepteryx present good instances, are relics of a more 
elaborate system of marking seen in a high condition of 
development in various exotic genera. This system 
consists essentially of a series of red and yellow markings 
occupying more or less completely the pale spaces in- 
cluded between the radially disposed dark lines that 
coincide with the nervures on the one hand, and the 
concentrically arranged dark spots or bars of the marginal 
and submarginal series on the other. That this system 
of markings is a very primitive one is rendered probable 
by the fact (1) that it is represented by vestiges in so 
many genera of both hemispheres; and (2) that it occurs 
in a well-developed condition in genera so widely 
separated in a geographical sense as the South American 
Leodonta and Catasticta, and the Australian and Indian 
Delias and Prioneris. Asa general rule, the spaces at 
the base of the wing tend to be occupied with red, and 
those on the dise and at the periphery with yellow; but 
in some cases, as has been seen, the red may encroach 
upon the submarginal yellow (as in Delias ceneus, 
D. eucharis, D. argenthona, etc.), or may be prolonged 
from the base in various directions across the wing 
(Hesperocharis hirlanda, M. lypera, M. pyrrha, and other 
American species of Mylothris). In the first-named 
genera, which exhibit the system in its full state of 
development, the distinction between the red and yellow 
markings is a perfectly easy one; but in such genera as 
Pieris, Leptophobia, Hesperocharis, Belenois, Ganoris, 
Synchloe, and Colias, whose markings are no doubt 
ultimately derived from the Catasticta and Delias pattern, 
it is not always easy to say to which part of the original 
system the relics that are present belong. On an in- 
spection of all the forms, it would, however, seem to be 
the case that those members of the yellow part of the 
system most often persist which are capable of being 
reinforced, as it were, by a neighbouring red patch 
towards the base of the wing. For instance, the most 
persistent marking of all is, as we have seen, the yellow 
precostal streak. ‘This is in most species of Leodonta 
and Catasticta closely bordered on by a bright red spot 
in the inner division of the precostal space ; and on tracing 
the condition of that space from these more ancestral 
genera downwards, through Pieris, Leptophobia, and 


Phylogeny of the Pierinee. 289 


Hesperocharis, we find it become tolerably evident 
that the dark yellow vestige which at last alone 
remains is the result of a kind of convergence both in 
colour and position of the yellow and red markings that 
at first lie simply side by side and are clearly dis- 
tinguishable the one from the other.* A further instance 
of possible convergence is supplied by the basal pinkish 
patch, found in the group of genera headed by Colias, 
which we have seen reason to identify with original 
markings of both the red and yellow kinds.+ In other 
cases, the persistence of various members of the present 
system is determined by the exigencies of a mimetic 
pattern, as in the species of Mylothris and Hesperocharis 
above referred to (p. 286). 


III. Pasytogenetic ConcLusions. 


1. The evidence of the Wing-markings. 


We are now in a position to consider as a whole the 
various sets of facts that have been above recounted, 
and to endeavour to construct, from the evidence at 
hand, a theory of the phylogenetic history of the entire 
group. This, of course, is only attempted under the 


* This, it is true, applies only or mainly to the New World 
forms ; the similar basal red patch that partly occupies the pre- 
costal space in certain species of Delias and Prioneris not seeming 
to have left any traces in those Old World genera (Belenois and 
T'eracolus), which appear to stand in much the same relation to 
Delias and Prioneris as Pieris and Hesperocharis do to Leodontu 
and Catasticta. 'The deep orange, however, of the precostal streak 
in some Old World species of Mylothris (as M. chloris and M. 
agathina), and in some species of Belenots (as B. helcida) which 
resemble them, may perhaps be derived wholly or in part from the 
precostal red ; but if so, it does not bear in itself a plain record of 
its origin as does the like feature in the corresponding genera from 
America. 

+ Again, in Pieris pylotis the only marking to be seen on the 
under surface of the hindwing (besides the basal spot) is a short and 
narrow orange streak near the apex of the median interspace— 
an interspace that in more primitive forms, such as P. locusta, 
Mylothris lypera, and others, is provided with an apical red patch. 
It is also worthy of notice that in many species of Belenois the 
spots of series I are apt to retain or assume an unusually deep yellow 
or orange colouring like that of the precostal streak—this series 
being in some species of Delias, and at least one of Prioneris, a 
sharer in the bright red of the basal patches (see p. 284). 


290 Dr. Frederick A. Dixey on the 


limitations above referred to, and is not to be taken 
as more than a connected statement of the probable 
import of the facts derivable from one _ particular 
source—those facts, namely, which relate to the colours 
and markings. I need hardly say that I recognise to 
the full that any phylogenetic conclusions founded on 
these data must of necessity be open to checking and 
correction in the light of information arrived at in other 
ways. 

The evidence that has now been examined seems to 
indicate that the wings in the earliest form of Pierine 
were uniformly overspread with a dark neutral tint.* 
The first variation from this condition appears to have 
occurred by the paling of certain areas in the principal 
interspaces between the nervures; a modification that 
may be seen in probably its simplest extant form in the 
remarkable American Pierine Hucheira socialis (Fig. 1). 
Here each interspace, including the discoidal cel! in both 
wings, possesses a more or less definite pale patch, those 
of successive interspaces being so arranged as to form a 
somewhat indistinct and interrupted band crossing the 
disc of both wings from the costa to the inner margin. 
Besides this central series of pale patches, there is also 
a submarginal row of much smaller and fainter spots of 
the same greyish-white hue. The whole pattern is 
repeated with very little change on the under surface, 
but on the hindwing with even less distinctness than 
above. The underside of the hindwing in this insect, 
indeed, probably exhibits the very oldest kind of Pierine 
colouring to be seen in any existing species.t 

The primitive system of marking manifested by 
Hucheira socialis persists with little alteration on the 
upper surface of many species of Catasticta, as, for 


“ This accords generally with the opinion expressed by Mr. 
Wallace: “There are, in fact, many indications of a regular 
succession of tints in which colour development has occurred in 
the various groups of butterflies from an original grayish or 
brownish neutral tint.” Darwinism,” 2nd edition, 1889, p. 274. 

+ While entirely agreeing with Staudinger and Schatz that the 
remarkable forms Styx infernalis and Pseudopontia paradoxa are 
probably of great antiquity, I cannot but regard the special 
Pierine affinities claimed for them by these authors (and especially 
for the latter) as more than doubtful. See Staudinger, Schatz, 
and Rober, “ Exotische Schmetterlinge,” 1892, sub voe. 


Phylogeny of the Pierine. 291 


instance, C. bithys (Fig. 2), C. toca, and C. colla. In other 
species of the same genus, the next steps are shown by 
the enlargement of the pale spots (as in C. anaitis ¢), 
their assumption of a purer white colour (as in U. 
anattis ©), and the confluence of those of the inner row, 
as in CU. trezene and C. ctemene ¢. In the latter insect 
the enlargement and confluence of the white patches has 
proceeded so far that the white must now be considered 
the ground colour, the original dark neutral tint being 
confined to the base and apices of the forewings, a 
narrow and interrupted marginal band on the hind- 
wings, and the course of the nervures with their branches 
(Fig. 3). In C. corcyra, the same process is continued 
almost to its furthest limit, the new ground colour has 
supplanted the old in every place except the apex and a 
narrow slip along the costa of the forewing. C. corcyra 
is thus practically a ‘‘ white” butterfly; and so within the 
single genus Catasticta a complete transition is to be 
found, from a dark almost black ground colour, with 
small and indistinct pale greyish patches, to a ground 
colour of dead white, with hardly more than a trace 
remaining of the original dark surface tint. This 
passage from C. bithys to C. corcyra is really an epitome 
of the whole range of variation in pattern throughout 
the entire group of Pierinzx. 

In such species of Catasticta as C. semiramis, where 
the pale patches are tolerably distinct, and have not yet 
coalesced, the dark marginal and submarginal series of 
spots, so characteristic of the Pierines as a whole, are 
visible at an early stage of their emergence. As has 
been shown above, on p. 267, the marginal series (called 
M) is formed by the remains of the dark ground colour 
between the outer series of primitive pale patches 
and the outer border of the wing; while the sub- 
marginal series S comes into view between the outer and 
inner series of pale patches. Besides these, a relic of 
the dark ground colour is seen to remain about the 
disco-cellular nervules in the forewing, which forms the 
first beginning of a definite discoidal spot ; and another 
row of minute pale patches, each occupying the centre 
of an interspace on the extreme outer margin of the 
wing, begins to split up the marginal series in the 
manner already spoken of. The definite character of 
series M is assumed sooner on the forewing than on the 


292 Dr. Frederick A. Dixey on the 


hindwing, and in the latter situation the discoidal spot 
has at this stage scarcely become recognizable ; in some 
species, however, a patch of pale-coloured scales covers 
the lower disco-cellular nervule of the hindwing in the 
midst of a wide and not sharply-defined area of dark 
ground colour. 

So far little or no essential difference has been noted 
from the primitive pattern of Hucheira socialis; the 
variations produced having simply resulted from a 
greater or less extension of the intrusive pale tint along 
the lines originally marked out. But, on turning to the 
underside of any species of Catasticta, we find what at 
first sight appears to be a very considerable divergence. 
A careful examination, however, soon makes it plain 
that the pattern of the lower surface is throughout the 
genus essentially the same as that of the upper surface, 
though one or two new features are superadded. Con- 
fining our attention in the first place to the forewing, 
for instance in C. nimbice, we find the submarginal series 
S and the discoidal spot shown at least as clearly as on 
the upper side; while from the greater prominence of 
the pale spots at the extreme margin, M is brought still 
more plainly into view below than it is above. On the 
hindwing, the central pale band may be easily traced 
across the disc of the wing, leaving on the one side of it 
a basal, and on the other a broad marginal dark area. 
Traversing the latter can be seen a row of elongated, 
more or less wedge-shaped, yellow streaks, which repre- 
sent the more easily recognized series I of the forewing ; 
while a distinct row of yellow spots on the extreme 
margin continues the similar series from above. Though 
S and M are not yet sharply marked off from one 
another, the portions of dark ground colour to which 
they respectively belong are already separately recog- 
nizable. The undersides of C. nimbice, C. colla, C. toca, 
and C. corcyra will be found to make a very good 
transitional series, showing the gradual emergence of 5 
and M and the subsequent reduction of each to a mere 
festooned line.* The superadded features above alluded 


* An unnamed species of Catasticta in the British Museum 
presents a condition of the underside of the hindwing more exactly 
intermediate between C. corcyra and the usual type than any of 
the species mentioned. An individual of the same species is in 


Phylogeny of the Pierinze. 293 


to are (1) the streaks and touches of yellow that begin 
to occupy much of the area of the original pale patches, 
including, as has been seen, the spots of I on the hind- 
wing; and (2) the bright red patches visible in the 
basal region of the same wing close to the body. These 
have been already discussed at some length (see 
pp. 281, 285), and their importance has been shown in 
reference to the markings of more recent groups. In 
this place it is only necessary to note once more that 
their ancestral character seems to be proved not merely 
by their existence in a fully developed condition in 
genera so widely separated geographically as _ the 
Western Catasticta and Leodonta and the Hastern 
Prioneris and Delias, but also by the fact that there is 
scarcely a genus throughout the whole sub-family, 
whether in the Old or New World, that does not show 
some relic of their former presence. With regard to 
their origin, no clue appears now to exist. It seems 
impossible to trace them further back than to the 
Eastern and Western genera named; and the probably 
still more primitive form, Hucheira socialis, to which we 
should naturally turn for an indication of their develop- 
ment, affords us in this particular no information 
whatever. 

In the genus Leodonta we find some modification in 
the shape of the wings, and only three instead of four 
branches to the subcostal nervure of the primaries. But 
the wing pattern remains much the same as in those 
species of Catasticta where the white central band has 
grown into a well-defined area and the two series S and 
M have not yet become clearly distinguishable. The 
yellow streaks and red basal marks are still prominent 
on the hindwing beneath, and the separation between 
S and M is, in this genus also, more evident on the 
underside. 

In Pereute and Huterpe, which are undoubtedly very 
close allies of the genera just referred to, the development 
of the pattern has taken a somewhat different direction. 
The primitive dark ground colour is in the males of 
several species of Pereute, as P. charops, P. autodyca, and 


the Hewitson collection labelled as C. notha. This is undoubtedly 
erroneous, as the underside of the true C. notha is almost exactly 
like that of C. corcyra, 


294 Dr. Frederick A. Dixey on the 


P. swainsonii, relieved by dustings of pale pinkish or 
bluish grey, which do not seem to bear any particular 
relation to the lighter patches in Catasticta or Hucheira. 
These greyish areas are often, especially in the females 
(including those of the species named), brightened up 
into bars or broader expanses of vivid red (P. lewcodro- 
sime) or blue (P. telthusa), and the aspect thus produced 
is very unlike that of nearly all the members of the allied 
genera. But as has been shown above, at pp. 281 and 
285, an indication of affinity in coloration with those 
genera is preserved in the red and yellow patches of the 
underside of the hindwing ; and, what is very remarkable 
indeed, there are two species of Catasticta, namely C. 
teutamis and C. ctemene, the males of which are of the 
usual Catasticta type, while the females present, together 
with the ordinary Catasticta neuration and structure, the 
aspect characteristic of P. charops 9 and other highly 
coloured members of the genus Pereute. In the case of 
these two Catastictas there can, I think, be no doubt 
that it is the male rather than the female that represents 
the ancestral colouring of the group, and we are thus led 
to the conclusion that some common cause has brought 
about the divergence in the same direction of these 
female Catastictas and both male and female Pereutes 
from the ordinary character of their nearest relatives. 
This cause I believe to be mimicry, the models for which 
in the present instance are to be found in certain species 
of Heliconius, particularly H. melpomene, H. phyllis, and 
their allies.* ‘lhe resemblance is not perfect,t but in all 
probability is quite sufficient to be of much service to the 
Pierine mimics; and we find exemplified here, as in 
many other cases, the much greater readiness with which 
the female assumes the protective coloration. As we 


“ T also think it exceedingly likely that Papilio euterpinus, Godm. 
& Salv. (Ecuador), forms one of this mimetic group. It is, how- 
ever, remarkable that its range seems not to coincide with that of 
the Pierine, namely P. charops 9, which probably resembles it 
most closely in size as well as colour. 

+ The red colour of the species of Heliconius here mentioned (to 
which may be added H. vulcanus and H. hydara), as they occur in 
collections, is by no means so vivid as that of their supposed 
imitators. But Mr. A. G. Butler and Mr. F. A. Heron both 
inform me that the red in Heliconii has a special tendency to lose 
its brightness, even in specimens carefully preserved in the dark. 


Phylogeny of the Pierine. 295 


have seen, the male Catastictas are not affected at all; 
while in most species of Pereute the males have not 
advanced nearly so far along the mimetic path as have 
the females. Another point worthy of observation, which 
is paralleled elsewhere, is the fact that the males show a 
much closer approach to the complete mimetic pattern on 
the lower than on the upper surface.* 

A similar explanation will apply to the even more 
remarkable divergence shown by the genus Huterpe, 
though here the phenomena are so curious and complex 
as to require a more detailed treatment. This genus (as 
restricted by Butler, Cist. Entom. 1., pp. 34, 42) does not 
contain a single form that reproduces in general aspect 
the typical Pierine features ; every species indeed is the 
subject of a mimetic change, and such marks of their 
origin and relationships as they retain, are used up in a 
more or less modified condition to heip im the production 
of the deceptive pattern. Thus we have seen how in 
Huterpe critias, H. tereas, H. rosacea, and allied species, 
the area representing the white central band on the hind- 
wing, now tinged with rosy pink, has become an im- 
portant ingredient in the general imitation of Papilio 
zacynthus ¢ ; the red basal patches on the under surface 
also taking their part in the picture. There is, moreover, 
little doubt that the white spot on the forewings is 
identical in origin with the usual white central bar, while 
for the marginal yellow spots a counterpart is easily 
found in Catasticta nimbice and many of its allies. In 
Huterpe bellona we find the patch on the forewings 
bright yellow instead of white, while the rosy patch is 
intensified in colour and split up into three or four 
brilliant streaks radiating from the base towards the 
margin of the wing. The general effect is not very 
dissimilar from that of H. critias, and the identity of the 
markings is beyond question ; nevertheless the changes, 
slight as they are, are highly significant, for they import 
nothing less than the substitution of a member of the 
Heliconine for one of the Papilionine group as a model 
for imitation. Huterpe bellona undoubtedly mimics 
the Heliconine group of which H. thelxwpe and H. 
burneyi are conspicuous members; and it would be 
difficult to find a better instance to show how slight a 


* See Weismann, “Studies,” vol. i., pp. 7, 8 (English Edition). 


296 Dr. Frederick A. Dixey on the 


modification of an existing pattern may suffice to produce 
an effective copy of an entirely different insect. Facts of 
this kind seem to go a long way towards answering such 
objections to the theory of mimicry as those alluded to 
by Mr. Beddard in his “ Animal Coloration,” 1892, pp. 
214, 215. 

It is very remarkable that red basal patches, like those 
of the mimicking Pierines, occur on the underside of 
the hindwing in most, if not all, of the Heliconii and 
Papilionine mimicked by members of the genera Pereute 
and Huterpe ; and in some, e.g., H. melpomene, there is 
even a well-defined yellow precostal streak, Now 
although there is no difficulty in ascribing most of the 
features in the coloration of these Pierine genera to the 
operation of mimicry, it would seem, for the reasons 
given above (p. 285, note), that the origin of these par- 
ticular marks, which are so wide-spread and so persistent 
throughout the whole Pierine subfamily, must be 
excepted. Nor, im the absence of any evidence of a 
direct causal relation, does it seem to my mind more 
satisfactory to consider the occurrence of these patches 
in the three diverse groups, Papilionines, Heliconines, and 
Pierines, as the effect of similar external conditions. 
If we assert them to be purely ‘ accidental,” we are met 
by the fact that although they are found in some species 
of both Papilio and Heliconius that are apparently not 
the subject of mimicry, yet they are most distinct and 
most prevalent in those species that are copied by 
Pierine imitators ; and the same considerations (amongst 
others) seem to show that the supposition that they are 
simply due to inheritance from a common ancestor of all 
the groups must be inadequate. Before the phenomenon 
is dismissed as inexplicable, it may be worth while to 
consider the following suggestion. 

According to the well-known principle laid down by 
Fritz Miiller and expanded by Meldola, by which these 
authors have been enabled to account for cases of parallel 
resemblance between the species of protected genera, 
and also of the grouping of allied inedible species into 
distinct sets, each with its own scheme of coloration,* it 


* F. Miller in “ Kosmos,” 1879, p. 100 ; Meldola in “ Proc. Ent. 
Soc. Lond.,” 1879, p. xx., and “ Ann. and Mag. Nat. Hist.,” Dec., 
1882. See also Wallace, “ Darwinism,” 2nd ed., 1889, pp. 249-257, 
and Poulton, “Colours of Animals,” 1890, pp. 192-195. 


Phylogeny of the Pierinee. 297 


is advantageous for any two or more protected species 
to join their forces, in order to share the toll levied upon 
them by inexperienced enemies. In such cases it no doubt 
often happens that one species serves as the standard to 
which the others conform, whether by way of convergence 
or of arrested divergence; but there seems no reason 
why, especially if there is no conspicuous inequality of 
numbers, there should not be akind of ‘‘ give and take ” 
arrangement between mimicker and mimicked, the latter 
advancing some way to meet the former for their mutual 
benefit. In other words, when two species, A and B, 
form an association of this kind, it need not be supposed 
that the form of A remains fixed, while B assimilates 
itself to it, or vice versa ; but the association may really 
be formed by both A and B converging towards a point 
between them, or, in short, mimicking each other. 
The acceleration of the process which in many cases 
would result, must of itself be an advantage. Now if 
the grouping of kindred inedible forms in associations of 
this kind is beneficial, it would seem that the benefit 
might extend to members of families far removed from 
one another, like the three now under discussion, as well 
as of the same family or genus;* the only requisites 
being (1) that all should be more or less inedible, and (2) 
that all should have patterns capable of assimilation to 
one another. Nor does there seem to be any reason why a 
mutual convergence of the kind indicated should not 
take place under these conditions also. With respect to 
(1), it has, I think, nearly always been assumed that the 
Pierine which resemble the members of other families 
are edible; and this is no doubt the case with the 
numerous members of the genus Dismorphia + (Leptalis 
of Dalman), which were among the first to attract notice 
to the subject of Mimicry in general. But no direct 
proof seems to exist that the species of Huterpe and 
Pereute now under discussion are edible; and indeed 
Mr. Wallace considers (Trans. Ent. Soc. Lond., 3rd 
series, iv., pp. 309, 310; Darwinism, 2nd ed., 1889, 


** Such an association is already well known to occur between 


Heliconide and Danaide. 

This, however, has been questioned by Mr. Beddard, who cites 
some observations of Scudder as tending to show the contrary. 
See “‘ Animal Coloration,” 1892, p. 215. 


TRANS. ENT. SOC. LOND. 1694.—PART Il. (JUNE.) U 


298 Dr. Frederick A. Dixey on the 


p- 244) that the genus Delias (Thyca of Wallengren), 
which is probably nearly related to Huterpe and Pereute, 
possesses a disagreeable flavour, or some other special 
means of protection. If this is shared by its American 
relatives, the first of the above requirements would be 
satisfied, while the second (2) has been shown to be met, 
in the special instance of the red basal patches, by the 
occasional presence of somewhat similar spots or 
patches (whose origin is not now under discussion) in 
other species of both the Papilionine and Heliconine 
groups, which afford quite sufficient material for the 
assimilative process to work upon. Until, therefore, 
direct proof is forthcoming of the edible qualities of the 
Pierines in question, it will not be unreasonable to sup- 
pose that they have joined an “inedible association,” 
and to attribute this curious agreement in a detail of 
pattern between members of such diverse groups to the 
operation of what may perhaps be called “ reciprocal 
mimicry.”’* The Pierines have no doubt moved furthest 
from their original form, having in most respects clothed 
themselves in Papilionine and Heliconine colours, but 
the latter have themselves advanced some way to meet 
the Pierines, in adopting from them a more distinct 
and characteristic employment of the red basal patches.+ 


* This, it will be seen, is distinct from “convergence” in the 
usual sense, inasmuch as that term has hitherto been employed 
(as by Poulton, ‘ Colours of Animals,” 1890, p. 195) to signify the 
assimilation of one form to another, rather than the mutual 
approach by two forms toa mean between them. 

t+ Fritz Miller notes (“ Kosmos,” 1879, p. 100, and translation 
by Meldola, Proc. Ent. Soc. Lond., 1879, p. xxiv.) that “in the 
province of Santa Catharina, Huterpe tereas is common in the 
forest-paths almost throughout the entire year, while its model, 
Papilio nephalion, is, on the other hand, a rare butterfly.” This is 
so far in favour of the supposition that the resemblance is not due 
to mimicry by the Pierine, at least in the strict sense. 

There igs one more piece of evidence that seems to favour the 
conclusion arrived at above. So far as I am aware no explanation 
has yet been offered of the fact that it is the females and not the 
males of Papilio polymetus, P. zacynthus, etc., that are resembled 
by Huterpe tereas and E. critias ; whereas the males, which display 
brighter colours, afford at least as good if not better models for 
imitation. I would suggest that this is really due to “reciprocal 
mimicry.” The protection gained by the resemblance between the 
Pierines and Papilios is not all on the side of the Pierines, but 
mutual ; and the female Papilios have, as is usual, felt the need 
of it more urgently than the males. For this reason the female 


Phylogeny of the Pierinee. 299 


Huterpe theano and EH. ewrytele present us with another 
kind of departure from the ordinary Pierine condition, 
the latter being an excellent copy of Lycorea atergatis 
and Ceratinia (Ithomia) dionea; while EH. theano ex- 
hibits the form of H. ewrytele with a coloration much like 
that of Pereute charops ¢. 

We find then that, thus far back in the growth of the 
Pierine stock, a large section has been diverted under 
the influence of mimicry from the regular course of 
development of the Pierine pattern; and in order to 
continue the history of the latter, we must retrace our 
steps and take up the thread where we left it before 
beginning to discuss the mimetic forms of Pereute and 
Huterpe. 

Although it does not appear that the Old World can 
show any truly Pierine form so ancient as Huchevra 
socialis, there does exist a genus which seems to repre- 
sent in the Hast nearly the same stage of development as 
Catasticta, Leodonta, Pereute, and Huterpe in the West. 
This is the genus Delias, the close relationship of which 
with the western forms mentioned has been indicated by 
Doubleday and Westwood (Genera of Diurnal Lepi- 
doptera, vol. i., p. 33), and more distinctly suggested by 
Wallace (Trans. Ent. Soc. Lond., 3rd series, iv., p. 344).* 
The pattern of Delias belladonna is essentially similar to 
that of the early Catastictas, consisting as it does of a 
dark ground-colour relieved by paler touches in the 
interspaces, between which touches the ground-colour is 
already beginning to show a rudimentary division into a 
marginal and submarginal chain with a vaguely indicated 
discoidal patch in each wing. On the underside we have 
the same pattern in a slightly more distinct form; while, 
as in Catasticta, the pale areas of the hindwing and the 
apical region of the forewing are furnished with yellow 
streaks. ‘The red basal patches, so characteristic of 


Papilios have been led to meet the Pierines by discarding, or at 
any rate by not adopting, the bright metallic blues and greens that 
ornament the other sex. This was no doubt a shorter and easier 
way to the formation of an “inedible association,” than would 
kave been the acquisition by the Pierines of colours more nearly 
resembling those of the male Papilios. 

* Delias agrees closely in neuration with Pereute and Leodonta. 
See Butler, Cist. Entom., i., p. 40. Staudinger and Schatz, “ Exot, 
Schmetterl.,” 1892, Theil. ii., p. 63. 


300 Dr. Frederick A. Dixey on the 


Catasticta and its allies, are not represented in D. bella- 
donna, but in all the members of the pasithoe group they 
are not only present, but have become a very conspicuous 
feature in the coloration. It is not difficult to bring the 
other markings of D. pasithoe, D. thisbe and their allies 
into relation with those of D. belladonna, though the 
dark series S and M are as a rule less clearly indicated. 
The pasithoe and belladonna groups of Delias may thus 
be considered as on the whole the most ancient.* Inthe 
well-known D. eucharis we have the series S and M 
complete; the female, as is usual, showing less departure 
than the male from the more generalised form. As in 
Catasticta, no sooner have the dark series emerged into 
distinct existence than they begin to disappear ; they are 
much less recognizable in D. hyparete and D. agostina 
than in D. eucharis, and in D. hierte they reach the 
vanishing point. D. eucharis and its immediate allies 
follow the early condition of Delias in having the inter- 
spaces on the underside of the hindwing more or less 
filled in with yellow, but they depart from it in having 
lost the red basal patches, and in possessing a conspicuous 
submarginal band of red spots (series I) between the 
dark series S and M; these are best seen in D, eucharis 
itself. An early stage of this red series is probably 
present in D. ceeneus and the closely allied D. philotis, 
which seem to be linked with the still more primitive 
butterflies of the pasithoe group through such forms as 
D. belisama and D. descombesi (see p. 284). Regarding 
then D. coeneus as a new starting-point, we may trace 
from it in one direction the Indian group headed by 
D. eucharis, and in another the Australian forms 
D. aganippe, D. harpalyce, and D. nigrina. It is to be 
observed that both D. ewcharis and D. aganippe, though 
probably more recent than D. ceneus so far as regards 
the underside of the hindwing, nevertheless retain in 
other respects more of the primitive pattern ; from which 
we may probably conclude that some form now lost, 
rather than D. ceneus itself, supplies the true link with 
the belisama and pasithoe groaps, while D. ceeneus and 
D. philotis survive to show how “ series 1” acquired in 


* T follow, in nearly every particular, Mr. Wallace’s division of 
this genus into groups. See Trans. Ent. Soc. Lond., 3rd series, iv., 
p. 344, ete. 


Phylogeny of the Pierine. 301 


the eucharis and aganippe forms its conspicuous character. 
I have little doubt that the curious form D. aganippe is 
one of the oldest existing species of Delias in the 
Australian region, inferring this from its retention of the 
primitive series S and M with the discoidal patch in the 
forewing in fairly distinct form, from the presence of 
yellow streaks in the interspaces of the under surface, 
and from the occurrence of a basal red spot in the pre- 
costal space of the underside of the hindwing. Many of 
these points assimilate it to the pasithoe group, and even 
more closely to D. belladonna.*  D. harpalyce and 
D. nigrina seem to follow naturally on from D. aganippe, 
though in respect of the costal red they are perhaps a little 
nearer the still more primitive D. belisama. Another 
Australian offshoot of the D. ceneus stock is probably 
represented by D. argenthona, D. mysis, D. isse, and 
D. dorimene. The Indo-Malayan and Australian nysa 
group is probably derived from the pasithoe and bella- 
donna groups through D. orphne and D. momea. It 
would be most interesting to attempt to trace in detail 
the phylogenetic history of the whole of this extensive 
genus, but it must here suffice to have indicated what ap- 
pear to be the principal lines of derivation. ‘Two points 
of analogy with the kindred neotropical genera may be 
noted before we pass on. The first is that although the 
invasion of a paler tint does in Delias tend to spht the 
original ground-colour into marginal and submarginal 
series quite similar to those of Catasticta, and though 
these series in various stages of development and sup- 
pression, as also in Catasticta, meet us here and there 
throughout the entire genus, yet in many cases the series 
never emerge in any recognizable form, and there seems 
in several species a tendency for the white mvasion to 
begin near the bases of the wings, and simply sweep the 
dark ground-colour away towards the margins. Even in 
these cases, however (as in D. belisama and D. descom- 
besi), the females, on the underside, will usually show 
some indication of the primitive series. The second 
point to be noted is the way in which the resources of 
Pierine coloration are taxed to give the insects of this 
genus a brilliant appearance. As in the case of some of 
their neotropical congeners, this is effected not so much 


* See Wallace in Trans. Ent. Soc. Lond., 3rd series, iv., p. 349. 


302 Dr. Frederick A. Dixey on the 


by the importation of any new feature as by the 
brightening up and extension of features already present, 
and indeed often common to the whole subfamily. The 
reason for this ornamentation is doubtless the same in 
both cases—namely, to call attention to inedible qualities. 
The Hastern genus, however, seems to have itself become 
a model for mimicry (Wallace in Trans. Ent. Soc. Lond., 
3rd series, iv., pp. 309, 344), while the Western genera 
have modified their ancestral form in order to join an 
alien company of inedible insects. 

The genus Prioneris, though differing slightly from 
Delias in neuration and other structural characters, is 
probably nearly akin to it, and represents, so far as the 
colouring is concerned, a similar degree of antiquity in 
the Pierine stock. ~The close resemblance of pattern 
between different species of Delias and Prioneris, to 
which reference has already been made (pp. 258, 284, 
etc.), is no doubt a true case of mimicry, but represents 
probably the result rather of arrested divergence than 
of the acquisition by Prioneris of new imitative features. 

The two groups that have just been discussed, namely, 
those formed by Delias and Prioneris in the Old World, 
and by Catasticta, Leodonta, Huterpe, and Pereute in the 
New, thus constitute together a second grade, as it were, 
in Pierine development. The only earlier species yet 
mentioned is Hucheira socialis, but there exist certain 
other forms which appear to be but little inferior to that 
insect in antiquity ; and to these, with Hucheira, the 
name of “ Pierines of the first grade” may be applied. 
One of the forms now referred to is Metaporia agathon. 
This remarkable insect is probably the representative of 
an ancient group of Pierines, among which were to be 
found the common ancestors of the two second-grade 
assemblages already mentioned, and which, no doubt, 
supplied the link at present wanting between Hucheira 
and Catasticta. The relation of Metaporia with the Hastern 
assemblage is more direct than with the Western, for 
although it offers points of structural difference from 
Delias, it shows, nevertheless, a condition of the primi- 
tive Pierine pattern which is in all essentials identical 
with that of the earlier kinds of that genus. The cor- 
respondence of its markings on the upper surface with 
those of Delias belladonna and D. eucharis is at once 
evident, and, like D. belladonna, it has a patch of bright 


Phylogeny of the Pierinee. 303 


yellow in the precostal space on the under surface of the 
lower wings. This, however, is usually confined to the 
iner division of the space.* Mr. Wallacet considers 
that Metaporia agathon forms with Pontia nabellica the 
fragments of an extensive natural group. He also 
points out the readiness with which the pattern of 
M. agathon can be traced through P. nabellica, P. soracta, 
and P. hippia, to its final disappearance in P. crateqi. 
In this range of species, to which may be added 
P. oberthiiri, P. leucodice, P. belucha, and others, we 
recognise the now familiar phenomena of the emergence, 
establishment, and evanescence of a marginal and sub- 
marginal series of spots, together with a discoidal patch, 
formed out of relics of an original dark ground-colour. 
Another primitive Pierine feature, namely.the per- 
sistence of dark ground-colour along the course of the 
nervures and nervules, is found throughout the group; 
and in P. crategi survives the disappearance of almost 
every other indication of the original coloration. 

The line thus marked out ends with P. cratagr; but 
in another direction there is little difficulty in linking 
Pontia with Hucheira, and so demonstrating the intimate 
connection with one another of the three last-named 
genera. Thus Pontia recalls Hucheira by the outline of 
its wings, especially the concave costal margin,{ and in 
pattern P. nabellica shows no very great departure from 
the same genus. Buta more striking sign of affinity 
exists in the common larval habitation, which, though 
merely rudimentary in P. crategi, and belonging only to 
the early larval stages, is no doubt a degenerate or 
undeveloped form of the elaborate silken nest con- 
structed by H. socialis. Webs of a structure apparently 
comparable with the latter are made by Neophasia ter- 
lootit and an allied species mentioned (though not 
described) by Behr.§ The affinity of the two latter 
insects with the genus Pontia appears certain; and 
though Behr is perhaps wrong in making them con- 


* One specimen of J/. agathon in Coll. Brit. Mus. has a few 
orange scales in the outer division of the precostal space as well. 
+ Trans. Ent. Soc. Lond., 3rd series, iv., p. 313. 
+ Noticed by Staudinger and Schatz, who compare it to that of 
Parnassius, Exot. Schmetterl., Th. ii, p. 62. 
Trans. Amer. Ent. Soc., 1869, p. 303; Proc. Calif. Acad. 
Sci., second series, vol. ii., 1890, p. 91. 


304 Dr. Frederick A. Dixey on the 


generic with H. socialis (which he does not seem to have 
seen.),* his description, nevertheless, renders it extremely 
probable that they are more closely related to that 
species than any other known butterfly. The Pierines of 
the first grade may accordingly be said to consist of 
Eucheira, with Behr’s two “ Neophasias,” Pontia and 
Metaporia. The prevalent larval habit of spinning is no 
doubt an indication of the high antiquity of the group, 
and suggests, as do other features, a relationship with 
the Parnassids. This suggestion is further borne out by 
the aspect of Mesapia jpeloria, which is no doubt an off- 
shoot of the genus Pontia. But such relationship is 
certainly of no close or direct kind, and is probably only 
referable to an extremely remote ancestor of the two 
subfamilies.+ 

Leaving the genus Pontia fora time, we may make 
a fresh start with Metaporia agathon, from which point it 
is not difficult to trace another principal stem, with 
numerous and important ramifications. ‘The females of 
several species of Mr. Moore’s genus Huphina, for 
instance H. phryne ?, show what is to all intents and 
purposes the same pattern as M. agathon (Figs. 7, 8) ; 
and the males differ only or chiefly by the more ready 
admission of the white invasion, at the expense of the 
remains of dark ground-colour. The pattern of such a 
form as H. nama 9, seems at first sight to belong to 
a different category, but a very good transition from 
that of M. agathon is afforded by H. eperia 9, which 
shows how the arrangement of white patches in H. nama 
? has grown out of the discal and cellular white, while 
the more marginal series I sinks gradually into the back- 
ground, and with its disappearance the two dark series 
S and M lose of course their distinctive character. Iu 
one ancestral point, namely the fillimg up to a greater 
or less extent of the interspaces of the hindwing under- 
side with yellow, Huphina shows a closer resemblance to 
Delias and Prioneris than to M. agathon, in which insect 
this feature is only slightly indicated. In many species 


* N. menapia, included by Behr in the same genus, differs from 
Eucheira is neuration and other points. 

t+ Davidina armandi has been spoken of as a transitional form. 
But, as remarked by Staudinger and Schatz (op. cit., Th. ii., p. 57), 
it is shown by Oberthiir’s figure (Etudes, iv., pl. 1i, fig. 1) to be 
without any of the points characteristic of the Papilionide. 


Phylogeny of the Pierinee. 305 


of Huphina the yellow is very vivid, in some it is 
warmed into a rich orange, and it not infrequently, as in 
H. naomi, H. lea, and H. judith, appears on the upper 
surface as well. 

From the pattern of Huphina to that of Oatophaga the 
passage is easy through such species as H. cassida, H. nabis, 
C. paulina (Fig. 9), and C. ega. In the latter genus, as 
in the former, the dark ground-colour is retained in 
much larger measure by the females than by the males; 
the latter indeed (as in most specimens of C. galena 4) 
have often lost it altogether. Other marks of speciali- 
zation shown by the male Catophagas are the sharply- 
pointed shape of the wings, and, above all, the presence 
of a tuft of long hairs springing from the base of each 
of the anal valves. All these characters belong equally 
to the next genus Appias (which indeed is not easily to 
be distinguished from Catophaga), though here the 
specialization of the males has in many forms been 
carried to a much greater extent, and is occasionally in 
some respects shared by the females, as in A. nero. In 
this insect the female shows the remains of the usual 
dark marginal and submarginal series standing out upon a 
ground colour of rich red almost as brilliant as that of the 
male, and altogether different from the ordinary Pierine 
white or yellow. The females, however, of A. celestina, A. 
clementina (Fig. 10), and others, do not depart, like the 
males, from the ordinary facies of the group, * and are 
indeed, barely distinguishable from the females of C. 
jacquinotii, C. alope, and other species of Catophaga. 
The assemblage of species united by Mr. Distant as 
Saletara (including S. panda, S. cycinna, ete.) 1s un- 
doubtedly an offshoot from the celestina group of Appvas. 

In the three last-mentioned genera, although the two 
series S and M are generally more or less traceable, and 
although in most cases a decided remnant of ancestral 
black persists along the costa of the forewing, there is 
as a rule no relic of the original ground-colour in the 
region of the disco-cellular nervules; that is to say, 
there is no discoidal spot or patch, In Hiposcritia, how- 
ever, which is apparently an early and purely Indian off- 
shoot of Catophaga, some species (as H. pandione) 
exhibit an incipient discoidal patch, still inconnection with 


* See Wallace, Trans. Ent. Soc. Lond., 3rd series, iv., p. 301. 


306 Dr. Frederick A. Dixey on the 


the dark-coloured costa ; in others (as H. lalage) this has 
become a distinct discoidal spot (Fig. 11). H¢poseritia, 
by its anal tufts, belongs decidedly to the Catophaga and 
Appias group; but the shape of its wings and the occa- 
sional persistence of dark ground-colour about the disco- 
cellular nervules indicate that its origin is to be placed 
somewhat far back towards Huphina and Metaporia. 

Starting again from Metaporia and Pontta, we find in the 
small Siberian P. leweodice an unmistakable link between 
these genera and Synchloe. In 8S. callidice, which per- 
haps come nearest to Pontia, the female shows S and M 
in a well-defined condition, together with a large distinct 
discoidal patch; the nervures and nervules are also in 
many instances clothed with black scales. From S. 
callidice we can advance in one direction through 
S. chloridice to 8. glauconome, S. johnstonii, and VN. 
hellica ; while S. daplidice and its immediate allies form 
another slightly divergent branch from 8S. chloridice. 
S. callidice again in all probability marks the point at 
or near which the assemblage of species grouped as 
Ganoris leave the main Synchloe stem; G. napi being no 
doubt the species which is closest to the original stock, 
as is shown by the large persistence of S and M, the 
tendency towards blackening of the nervures, the 
occasional indication of a discoidal spot, and the peculiar 
coloration of the underside of the hindwings. It is 
noticeable that Ganoris generally, including G. napi, 
retains the precostal yellow streak, which, though 
present in most species of Synchloe, happens to be 
absent in S. callidice. 

Most species of the genus Tutochila strongly resemble 
Pontia in pattern (the resemblance between T'atochila 
and P. lewcodice has been pointed out by Mr. Butler, 
Proc. Zool. Soc., 1872, p. 62), while 7’. autodice 9 comes 
nearer to M. agathon than does any species of Synchloe. 
The whole neuration of Tatochila is very similar to that 
of Pontia; and even the peculiar arching of the Ist 
subcostal of the hindwing (well seen in P. cratzgi) is a 
noticeable feature in 7’. theodice and other members of 
the same genus.* Tutochila may, therefore, be considered 
to be a derivative from the Pontia stem at a point some- 
what further back than the existing species of Synchloe. 


* See Mr. Butler's figures in Cist. Entom.,vol.i,, pl. iii., figs. 7 and 9. 


Phylogeny of the Pierine. 307 


A comparison of Tatochila (especially of T. theodice) 
with Phulia nymphula will show an almost exact identity 
of pattern, the correspondence extending even to the 
chevron-shaped spots of S in the hindwing, and to the 
peculiar triple striation of the nervules on the under 
surface. The neuration of Phulia is distinct, but not 
really far removed from that of Tatochila, some species 
of which show a near approach to its most remarkable 
feature ; viz., the emission of the second radial from the 
subcostal, and the consequent obliteration of the upper 
disco-cellular. Phulia again is closely allied in structure 
with Mr. Moore’s genus Baltia,* which, however, retains 
the short 8rd subcostal nervule lost by Phulia and 
several species of Synchloe. Phulia and Baltia are thus, 
in all probability, the terminal twigs of another branch 
which issued from the Pontia stem between the departure 
of Tatochila and Synchloe. Neophasia menapia appears 
to me to be an offshoot of the same stem at an earlier 
stage than Tutochila. It may possibly belong to the 
Huchetra group, but it is not known to make a social 
web,t and Behr gives no real grounds for making it 
congeneric with ‘‘ N.” terlootii. As above noted, its 
neuration is very distinct from that of Hucheira. 

The African species of the genus Mylothris{ appear 
to hold a somewhat isolated position. ‘Their neuration 
shows points of likeness with the equally isolated 
Australian genus Hlodine and the widely-spread and 
probably ancient Nychitona, || which genera they also 
somewhat resemble in texture of wing. In pattern 
they are chiefly remarkable for the strong and distinct 
‘development of series M, and of the precostal orange-red 


* Moore’s type is “ Mesapia” shawii of Bates, which is certainly 

generically distinct from MM. peloria, and much nearer Synchloe. 

. The butterfly described by Moore in Proc. Zool. Soc., 1882, p. 234, 
as Synchloe butleri is undoubtedly a Baltia. 

+ Stretch, however (Papilio, ii, pp. 106, 107), describes the 
larvee of N. menapia as suspending themselves from great heights 
at the end of long silken threads. Hven the pup were, in some 
cases, suspended. It is probable that the numerous threads cover- 
ing the bark of the trees seen by Stretch (2bid.), were also made 
by these larvae. 

~ I cannot but agree with Mr. Trimen (South African Butter- 
flies, 1887, vol. iii, p. 29) that the American P. pyrrha, Fabr., 
and ‘allies, are not generically akin to the African J/. agathina, M. 
chloris, etc, with which Mr. Butler unites them on account of 
their correspondence in neuration (Proc. Zool. Soc., 1892, pp. 37, 38). 

|| Distant, Rhopalocera Malayana, 1882—1886, p. 287. 


308 Dr. Frederick A. Dixey on the 


on the underside of the hindwing. We may, perhaps, 
regard all three genera as relics of an ancient fauna of 
the Eastern Hemisphere coeval with the earliest forms 
represented by the present Delias stock, and anterior to 
the various branches which have diverged from that 
genus or from Metaporia. The precostal orange suggests 
a kinship with Delias on the part of Mylothris, though 
no such link exists in the case of the other two genera, 
and, as pointed out by Mr. Trimen, who considers 
Mylothris and Thyca (Delias) to be allied genera, the 
former genus, like the latter, undoubtedly contains 
subjects of mimicry by Pierine of other groups. Thus, 
‘The imitation of M. agathina by P. thysa, Hopff., is 
deceptively close in both sexes, and M. poppxa, Cram., 
is similarly copied by P. rhodope, Fabr., on the West 
Coast. MM. agathina is also mimicked by the female 
Hronia argia, Fabr.’’* 

We now have to deal with the genus Belenois, which 
presents some special difficulties. In the first place, we 
find an assemblage of species (B. lasti, B. nagare, B. 
majungana, and B. isokart) from Hast Africa and Mada- 
gascar, with respect to which Messrs. Grose Smith and 
Kirby, after noting that they are now placed with Phris- 
sura in the Collection of the British Museum, go on to 
observe that they “ differ entirely in neuration from the 
types of Phrisswra, agreeing in this particular with 
Belenois, in which genus we prefer to include them.’’+ 
The type of Mr. Butler’s Phrissura is apparently 
P. illana, Feld.,t and in 1872 (Proc. Zool. Soc., 1872, 
p- 51) the only other species of Phrisswra admitted by Mr. 
Butler was P. palisma, which is undoubtedly a very near 
relative of P. illana. Both of these species offer differ- 
ences in neuration from B. lasti and its allies.|| Now, 


* South African Butterflies, vol. iii., p. 39 (1887). 

+ Rhopalocera Exotica, Oct. 1892. 

~ Mr. Butler first gave it as P. cynis (Cist. Entom., iii., 1870, 
p. 49), but afterwards corrected it to P. illana (Trans. Ent. Soc. 
Lond., 1871, p.171). Mr. Distant has since made cynis the type 
of his new genus Udaiana. (Rhopal. Malayan., 1882—1886, p. 286 
and note). 

|| In P. illana (forewing) the discoidal cell is unusually short, 
the second subcostal starts from the end of the cell, and the 
second and third median nervules come off near together; in 
B. lasti the cell is of the ordinary length, the second subcostal is 
emitted before the end of the cell, and the second and third 
median nervules do not start particularly near together. 


Phylogeny of the Pierine. 309 


however, certain forms are included under Phrissura in 
the Collection of the British Museum (P. phaola, P. 
sylvia, P. eudowia, etc.) which appear to me to agree in 
structural characters with B. lasti, and to differ from 
P. illana. If, therefore, Messrs. Grose Smith and 
Kirby are right (which I do not doubt) in separating 
B. lasti, ete., from Phrissura as represented by P. illana, 
I am strongly of opinion that P. phaola, P. sylvia, 
P. eudoma,* and P. coniata should go with them; 
and to this assemblage I would add every so-called 
Belenois that is furnished with anal tufts, together 
with ‘ Belenois”? or * Glutophrissa” saba. We should 
then have a natural group of African and Malagasy 
Pierines, between which there would exist no assignable 
difference in structure, while they would be all alike 
characterised by the possession of anal tufts like those 
of Catophaga and Appias. They may, perhaps, be 
considered as a section of Belenois, but to my mind they 
seem by their neuration, no less than by the obvious 
character of the anal tufts, to come much nearer to 
Appias, of which genus I am disposed to regard them 
as an African offshoot. The South American species 
Glutophrissa poeyi, G. margarita, G. castalia, etc., are 
very closely related to the present section.+ 

Although the structural affinity of the forms that have 
just been discussed with each other and with Appias 
seems undeniable, there is no doubt that in pattern they 
mostly show a marked divergence from that genus. 
This appears to be due to mimicry, the models for which 
are usually supplied by the genus Mylothris. Thus B. 
lasti is said by Messrs. Grose Smith and Kirby to be 
“nearest to B. trimenia, Butl.” ; the latter, however, is a 
true Mylothris with the characteristic neuration of that 
genus. Considerable resemblance also obtains (as men- 
tioned above, p. 308) between P. rhodope, Fabr., and M, 


: P. eudowia, Cram., is apparently identical with P. sylvia ?, 
Fabr. 

+ Mr. Wallace (loc. cit., p. 312) includes these American and 
African forms with Appias, Catophaga, and Hiposcritia in his 
genus Tachyris, For the purposes of the present paper it will 
perhaps be sufficient if I refer to the American and African 
species with Appias-like structure and anal tufts collectively as 
: Ben B,” while designating polisma and illana as “ Phris- 
sura A.” 


310 Dr. Frederick A. Dixey one the 


poppea, Cram.* The peculiar facies of G. saba 2 seems 
to be most likely due to mimicry of Nyctemera apicalis, a 
protected moth. Those members of the group that have 
not been affected by mimicry retain few traces of the 
original ground-colour, and present very much the 
appearance of an ordinary male Appias (G. saba 6, G. 
castalia, etc.). 

The other species usually included under Belenois, 
besides differing in points of structuret from those just 
discussed, show as a rule a greater persistence of dark 
ground-colour. Both S and M are usually present in 
fair development, either separate or fused, and the 
discoidal patch on the forewings is generally well 
marked. ‘The-markings in several species show a strong 
resemblance to those of Synchloe hellica and S. johnstonii ; 
but notwithstanding this, I am disposed to think that 
Belenois proper really represents an offshoot of another 
part of the Old World Pierine stem, that, namely, re- 
presented by Delias and Prioneris. The distribution of 
dark and light ground-colour in such species as B 
mesentina might easily be derived from those of D. 
belladonna and D. eucharis, and the underside of the 
hindwing in this and other forms of Belenois offers only 
slight modifications from that of D. belladonna. OB. 
peristhene exhibits a curious resemblance on both sur- 
faces to D. nysa which may perhaps be due to mimicry, 
although the ranges of the two insects only coincide for a 
small part of their extent ; it is not improbably in any case 
an indication of real affinity. In neuration, those species of 


* T adopt Mr. Trimen’s unravelling of the strange confusion 
that surrounds the synonymy of rhodope, Fabr., and poppea, Cram. 
(South African Butterflies, vol. i., p. 35, and note). The upshot 
no doubt is that a true Mylothris (Papilio poppea of Cramer 
according to Mr. Trimen) is closely copied by at least one Pierine 
of the “B” group of Phrissura (Papilio rhodope of Fabricius 
according to the same authority). I do not propose to embark 
upon the question further than to observe that the “ Synonymic 
Catalogue” identifies the two, and that Mr. Trimen himself 
unfortunately speaks of rhodope in the text, when he must mean 
poppea (ibid. p. 35). 

+ Negatively by the absence of anal tufts, positively by the 
presence (in many) of anal hooks, also by the straight direction and 
greater relative length of tke upper disco-cellular nervule in the 
forewing. In many species of this group, which we may designate 
Belenois proper, the first subcostal branch anastomoses with the 
costal. 


Phylogeny of the Pierine. 311 


Belenois in which the first subcostal is distinct are very 
near Prioneris, and the remainder seem to approximate to 
Delias by the partial loss of a subcostal branch, as well 
as by the contour and direction of the disco-cellular 
nervules. It is to be observed that some species of 
Belenois proper, like those of the “‘ Phrissura B” group, 
are mimics of various species of Mylothris, as B. thysa of 
M. agathina. Pinacopteryx is probably a collateral or 
derived branch of Belenois; the female of P. capricornus 
retains S and M on both surfaces in a well-marked 
condition. 

We now come to a large and important group of 
genera which appears to take origin from the older 
Pierine stock at a point nearest to the genus Synchloe. 
An early stage of divergence is reached by Teracolus ;* 
many species of which show the primitive series S and 
M, together with the discoidal spot and the precostal 
yellow of the underside of the hindwing. The curious 
genus Herpxnia is probably a survival of the links that 
once united Teracolus with Synchloe ; in neuration it is 
intermediate between the two, and in pattern, together 
with aberrations peculiar to itself, it shows points of 
contact with both. The pattern of the upper surface in 
Teracolus is easily derivable from that of Synchloe, the 
nearest approach to Synchloe in this respect being made 
by the Jdmais group ; compare, for instance, 7. dynamene 
with S. hellica ?. But the passage from the underside 
of the hindwing in Synchloe to that of Teracolus is more 
difficult ; Herpznia, however, enables us to bridge over 
the interval very fairly, for if H. tritogenia and H. 
lacteipennis be interposed between S. glauconome and 
T. puellaris ?, the abruptness of the transition is 
removed. The pupa of Yeracolus tends to be boat- 
shaped, with a sharply-pointed rostrum and large 
prominent wingst—which characteristics we shall find 


* T follow Mr. Butler (Cist. Entom. vol. i., p. 36), with whom Mr. 
Trimen is in agreement (South African Butterflies, vol. iii., p. 82), 
in uniting Teracolus, Idmais, and Callosune under the single head 
of Teracolus. 

+ These features are not equally well developed in all species of 
Teracolus. They are very distinctly shown in pupe of 7. evarne 
(Coll. Hope) ; pupz of 7. calais (Coll. B. Mus.), on the other hand, 
are slender, less recurved, and nearer the Ganoris or Synchloe form, 
The pupa of 7. pleione (Coll. B. Mus.) is stout, and not unlike 
that of Gonepteryx rhamni; compare Mr. Trimen’s description, op. 


312 Dr. Frederick A. Dixey on the 


in a greater or less state of development in all the 
genera of the particular group of Pierines now under 
discussion. Other features, which appear first in Teracolus 
and will be met with again later on in the history of the 
er ae are the pinkish fringe of the wings in some species 
(as 7. wallengrenii, and often in T. protractus) and the 
faint pinkish Colias-like spot or patch at the root of the 
cell in the hindwing underside of others (as J. tri- 
punctatus, T. celestis 2, T. eupompe, and T. theogone). 

The genus Jaias follows easily upon Teracolus, the 
females of the two genera showing, as usual, a greater 
affinity in colouring than the males (compare, for instance, 
the males and females of 7. tone and J. marianne *). 
Series § is often well preserved in Jvias, especially on the 
underside and in the female; the discoidal spots are also 
prominent, and begin on the lower surface to assume an 
ocellate character. In neuration, Jxias differs only 
slightly from Teracolus; while the pupa, as shown in 
drawings by Captain Boys of I. marianne and I. evippe 
(Coll. Hope), has the same characters as Teracolus in a 
more strongly-marked condition. 

From the usual system of colours and markings in 
Ivias, there can be little doubt that that genus represents 
the transition from T'eracolus to Colias. The underside 
of several species of the latter genus (for instance, C. 
subaurata) is strongly recalled by that of I. marianne, 
in which the Colias-like condition of the discoidal spots 
is one of the most noticeable features. The affinity 
suggested by the colour-pattern is borne out by the 
structure of the antennz, which in Jvias show a distinct 
approximation to the gradually thickened form so 
characteristic of Colias and its immediate allies. The 
pupa in Colias, as in Teracolus, seems to vary somewhat 
in form; but in all known cases it shows a tendency, 
sometimes strongly pronounced, towards the acuminate 
and boat-shaped outline characteristic of this group of 
genera. 


cit., vol. iil., p. 82. I refrain from using the terms “ wing-covers” 
or « wing-cases,” the inapplicability of which has been shown by 
Prof. Poulton (Trans. Linn. Soc., 1890, series ii., vol. v., part 5, 
. 188). 
* Mr. Butler points out the intermediate character of the two 
African species, Jzias eulimene and I. venatus. Proc. Zool. Soc., 
1871, p. 254. 


Phylogeny of the Pierinee. 3138 


The genus Xanthidia is transitional between Colias 
and Terias, in neuration coming nearer the latter. ‘The 
pattern on the upper side is scarcely modified from that of 
Colias; on the under surface, however, 8 has lost much 
of the distinctness it possesses in so many species of the 
latter genus, and its relics take part in a general 
mottling which is very probably protective in object. 

In Terias the resolution of series Sand M, which is still 
visible in most of the females and many of the males of 
Colias, has usually disappeared from the upper surface ; 
the underside of the hindwings, however, generally shows 
S in a somewhat modified condition. The pink edging 
to the wings, so characteristic of Colias, and occasionally 
visible in Xanthidia nicippe, is indicated in several 
species of Terias, as T’. messalina, T. delia, and 1’. rhodia, 
The pupa of Terias is more sharply acuminate and re- 
curved than that of Colias (see pupa of 7. mandarina and 
T. eecavata in Coll. B. Mus.). 

Sphenogona presents in pattern no marked differences 
from Yerias, with which genus it is so closely allied. 
S. gratiosa, like T. agave, X. nicippe, and some other 
species of these genera, shows a relic of the pinkish Colias- 
patch at the root of the cell on the underside of the 
hindwing. In neuration, Sphenogona, while generally 
resembling Terias, is peculiar in emitting the first and 
second subcostal branches of the hindwing from a short 
footstalk beyond the end of the cell. In this respect it is 
intermediate between Verias and Leucidea, which latter 
genus presents no markings to guide us, but has no 
doubt taken its origin directly or indirectly from Terias.* 
Another probable descendant of Terias is the genus 
Nathalis, which agrees very fairly with Terias in neuration, 
except that it wants one nervule in the forewing, probably 
a branch of the subcostal. There isa striking resemblance 
of pattern between N. iola and TJ. elathea 3. In the 
absence of paronychia Nathalis reverts to the condition of 
Colias. 

Pyrisitia seems to represent an early offshoot of the 
stem leading from Colias to Terias. In most points it 


* Staudinger and Schatz (op. cit., Theil. ii, p. 66, etc.) consider 
Leucidea to be more closely related to Pontia (Nychitona) than to 
Eurema (Terias). The balance of evidence seems to me to be 
against this view. 


TRANS. ENT. 800, LOND. 1894,—PaRT II. (JUNE.) X 


314 Dr. Frederick A. Dixey on the 


agrees with the latter genus, but in the position of the 
subcostal nervules of the hindwing it retains the condition 
of the former. 

A fresh departure from Colias is headed by the genus 
Meganostoma, which offers a close resemblance to Colias 
in both structure and pattern.* The series S and M, 
fairly distinct in the female of M. philippa, are also 
indicated on the upper surface in the males of some of 
the species. In the males of some other species they 
are fused above, but in all they are distinguishable 
beneath. In the outline of the wings Meganostoma 
shows a close approach to Gonepterya, from which genus 
indeed the females of some of the species are hardly to 
be distinguished. 

From Gonepteryx the line passes on to Amynthia and 
Rhodocera. All these’ genera retain many Colias-like 
points, particularly the pinkish patch at the apex of the 
cell on the hindwing undersurface. The males of the two 
latter, like those of Meganostoma and many species of 
Colias, also possess the well-known patches of raised 
. scales above the subcostal nervure of the hindwing ; and 
in the same two genera is seen for the first time the 
contrast between areas of flat and raised scales over the 
whole upper surface of the wings which is so conspicuous 
a feature in Catopsilia, Phebis, and Callidryas. 

Dercas appears to be an Indian offshoot of Gonepteryz. 
A good transition from the latter to the former is 
afforded by D. wallichii, as pointed out by Mr. Wallace, 
Trans. Ent. Soc. lLond., ord series, iy., ‘p. 398: 
Kricogonia is probably an analogous derived form in the 
Neotropical Region. 

The Eastern genus Catopsilia forms with the Western 
Callidryas, Aphrissa, Metwra, and Pheebis, a group which 
clearly belongs to the present section. ‘Their structure 
and coloration relate them closely to Gonepteryx, though 
in some respects they show signs of a more ancient 


* Mr. Butler (Cist. Entom., iii., p. 46) describes the neuration in 
the forewing of Meganostoma as being like that of Gonepteryz, 
“excepting that the second subcostal is emitted just before the end 
of the cell.” In some individuals, at any rate, the second subcostal 
is emitted at the end of the cell, and in M. cesonia it may even be 
thrown off after the end of the cell, as often in Colias. This indeed 
is the condition represented in Cist. Entom., vol. 1., pl. ii., fig. 4. 


Phylogeny of the Pierine. 315 


ancestry. The retention of the pink patch on the under- 
side of the hindwing, and (in several species) of the pink 
fringe, brings them near to Colias; to which they also 
approximate by the form of the antennz, by the very 
general occurrence of special patches of raised scales on 
the hindwing of the males, by the frequent presence, 
especially in the females, of the series S and M and the 
discoidal spots in a more developed condition than that 
characteristic of Gonepteryz, and by the common 
tendency of the latter spots on the lower surface towards 
ocellation. The oldest forms of the whole group appear 
to be those contained in the genus Catopsilia. C. florella, 
for example, is very closely related to Gonepteryx, and 
still more nearly to Amynthia, with which it corresponds 
in colour and in texture of wings (compare especially 
the undersides of C. florella ¢ and A. clorinde). The 
peculiar thickening of the nervures, especially the sub- 
costal and median, on the under surface of the hindwing, 
which in Gonepterya and Amynthia give almost the 
effect of a folding of the surface, is plainly visible in 
C. florella. In neuration, C. jflorella agrees minutely 
with A. clorinde. Another indication of the superior 
antiquity of the Hastern Catopsilia as compared with the 
Western Callidryas is found in the shape of the pupa. 
This in Catopsilia (as remarked by Mr. Trimen, op. cit., 
vol. ii1., p. 184) is only moderately acuminate and slightly 
recurved, thus showing no very great departure from 
the Colias and Gonepteryx form ; in Callidryas, however, 
so far as is known, it is always very sharply acuminate 
and strongly recurved, showing an exaggeration of the 
“poat-shaped”’ condition almost as marked, in a differ- 
ent way, as that of Huchloe.* The Colias-hke pink 
edging appears also to be found more frequently in 
Catopsilia than in Callidryas (though it occurs in Calli- 
dryas senne) ; and while the Colias-like spot at the root 
of the cell on the underside of the hindwing occurs in 
both the Eastern and the Western divisions of the 
group, it has in the latter lost the ancestral pink tinge 


*T agree with Mr. Butler that the pupa figured in Lep. 
Exot., pl. xlv., fig. 4a, as P. agarithe is very probably that of 
C. philea. It closely resembles a pupa of C. ewbule in the Hope 
Collection, which last exactly corresponds with Burmeister’s figure 
in the “ Atlas de la Description Physique de la Rép. Argent.,” 
1880, pl. v, fig. 2. 


316 Dr. Frederick A. Dixey on the 


which it usually retains in the former. It is not very 
easy to form an opinion as to the relative antiquity of 
the four Western genera; if we are to judge from the 
pupe, so far as they are known, it would seem that 
Aphrissa has undergone less modification than either 
Phebis or Callidryas. Another possible indication of 
the greater antiquity of Aphrissa is afforded by the 
shape and size of the palpi, which in this genus, 
especially in the females, show less departure than is the 
case in either Phebis, Metura, or Callidryas, from the 
Colias and Gonepteryx type. -Phebis with its offshoot 
Metura, from which it differs only in the contour of the 
wings, is probably the most highly specialised genus of 
all; this being shown by the possession of a tuft of 
long hairs on the hindwing of the male, which is not 
found in Aphrissa and Callidryas, and by the absence in 
most species of nearly all indication of dark ground- 
colour from the upper surface. It is remarkable that 
the Western genera just treated of are less closely akin 
to the South American Amynthia than is the Hastern 
genus Catopsilia, and especially the African C. florella. 
The genus Hronia seems to represent a branch of the 
present section which leaves the main stem somewhere 
between the points marked by Teracolus and Imias. 
With the exception of the presence of a fifth subcostal 
in the forewing (which, however, is sometimes wanting), 
the neuration of Hronia agrees fairiy with that of 
Teracolus. The antennee in Hronia differ in different 
species, but are all intermediate between the Teracolus 
and the Colias type; and the pupa of H.cleodora, with 
its very convex and prominent keel, is ike that of one 
of the stout forms of Teracolus, e.g., T. evarne. The 
assemblage of species distinguished by Mr. Butler as 
Nepheronia is probably older than Hronia proper, as is 
shown by the tendency of its antennz to terminate in a 
definite club, like the more ancient genus Teracolus. 
N. thalassina also retains a primitive Pierie feature in 
the yellow precostal streak of the hindwing underside, 
which again is found in some species of Teracolus. 
Eronia proper, on the other hand, is slightly nearer to 
Teracolus in neuration. The curious Hronia (?) lucas 
from Madagascar, which has only four subcostal nervules 
in the forewing, and whose antenne are slender, with a 
definite club, is probably one of the oldest and most 


Phylogeny of the Pierinee. Beil, 


generalised members of the Hronia and Nepheronia 
group now extant.* 

The last-named insect presents points of resemblance 
with Hebomoia, which again seems to be an offshoot of 
the same stem, that, namely, leading from T'eracolus 
towards Ixias and Obolias. The pupa of Hebomoia 
glaucippe, as fioured by Horsfield (H. I. C. Catalogue, 
see Distant, Rhopal. Malayana, 1882—6, p. 283) and 
Moore (Lep. Ceylon, 1880—1, pl. 49, fig. 1b), is stout, 
moderately acuminate, and much recurved, in which 
particulars it agrees well with the pupa of Iaias. 

Most of the insects of the genera Hronia, Nepheronia, 
and Hebomoia, retain in greater or less measure some por- 
tions of the primitive marginal and submarginal series. 
These, as usual, are most often to be met with in the 
females, and in several species of Nepheronia are utilised 
in the formation of mimetic patterns modelled on those 
of various Danaids and sometimes of other Pierines. A 
noticeable feature in some members of this group, per- 
taining chiefly to the males, is the brightening of the 
pale ground colour of the apex of the forewing into a 
brilliant yellow or orange patch. This character is first 
seen in Teracolus, where the orange of the apex may 
further deepen into crimson or violet; it passes on to 
Hebomoia, to Ivias and to Rhodocera ; in Colias, however, 
it gives place to a general yellow or orange suffusion of 
the pale ground colour, still strongest in the males.t 
It is found in Hronia (?) lucasti and H. leda, but not in 
other members of that genus; it is also absent in 
Nepheronia. 


* EF, (?) lucasii was originally described (as Callidryas lucasi) by 
Grandidier (Rév. et Magas. de Zool., Aug. 1867, p. 273). He, 
however, took the female for the male, and his supposed female 
C. lucasi is really the female of Catopsilia thawruma. Mabille in 
the Hist. Phys. Nat. et Pol. de Madagascar, vol. xviii, 1887, 
p. 281, gives a correct description of both sexes under the name of 
Eronia lucasii, but makes no mention of Grandidier’s mistakes. 
This species will not come into the genus Hronia as at present 
defined. By Brauer and others it has been called a Ptychopteryz ; 
the latter genus, however, was characterised by Wallengren from 
a species of Zeracolus (T. subfasciatus, Swains., Vid. Trimen, 
South African Butterflies, vol. iii., 1889, p. 92), and has no real 
claim to stand. 

+ In many species of the latter genus, indeed, the females may, 
as is well known, revert almost entirely to the ancestral white. 


318 Dr. Frederick A. Dixey on the 


This last feature in coloration is again met with in 
Eroessa and Huchloe, the neuration of which genera 
corresponds pretty closely with that of Hronia, not only 
in the presence of five subcostal branches in the forewing, 
but also in other particulars. The general likeness 
between Hronia leda and Huchloe belia, Linn., is very 
striking, and strongly suggestive of a near affinity 
between them. The well-known form of the pupa in 
Huchloe is an exaggeration of that seen in Hroma 
cleodora.* In some other points, however, Huchloe is 
nearer to Colias, as in the strong development of the 
discoidal spot in the primaries, and in the possession by 
some species of pink legs and a pink edging to the wings. 
This last feature is characteristic of the charlonia group 
of Huchloe, and is best seen in LH. lucilla. The underside 
of the hindwing in the same group has very much the 
character of the corresponding region in Colias paleno, 
including the pale undeveloped discoidal spot. The 
anteunee, however, of Huchloe are very distinct in form 
from those of Colias, and indeed are not much nearer 
those of Hronia. On the whole it seems most likely that 
Huchloe is a somewhat aberrant branch which takes its 
rise from the Pierine stem at a point near the divergence 
from the same stem of Hebomoia on the one hand, and 
H. (2) lucastt with the rest of the Hromia and Nepheronia 
group on the other. The isolated form Hroessa chilensis, 
which in neuration approaches the older Hronias (as 
E. (?) lucasti) and Hebomova, is perhaps a survival of a 
once more widely-spread and numerous assemblage, 
among which were to be found the immediate ancestors 
of the present-day Huchloes. Zegris seems to be a some- 
what highly modified offshoot of the Huchloe branch. 

We must now retrace our steps as far as to the group 
which I have called “ Pierines of the second grade,” those, 
namely, that are typified by Delias in the Hastern and 
Catasticta in the Western hemisphere. The latter genus 
with its near ally Leodonta forms a starting-point for a 
New-World division of the Pierine stem, which, if not 
equal in magnitude to the great division headed in the 


* The pupa in Ewchloe is not always recurved. See Edwards’s 
figures of Anthocharis (Euchloe) genutia and A. ausonides in 
“ Butterflies of North America.” See also Schatz, Exotische 
Schmett., Theil 11., 1892, p. 71. 


Phylogeny of the Pierine. 319 


Old World by Delias, Prioneris, and Metaporia, is never- 
theless of high importance and very great interest. 

The first departure from the condition of Catasticta 
seems to be that taken by the butterflies of the genus 
Pieris as restricted by Butler, closely associated with 
which must come the American species of Mylothris. The 
underside of Pieris locusta exhibits a pattern which is but 
little removed from that of Catasticta, the yellow streaks 
and red basal patches being still apparent on the under- 
side of the hindwing; while the upper surface of both 
wings is shared between the original dark and intrusive 
light ground colour in much the same manner as in C. 
ctemene, C. corcyra, and other species of Catasticta. In 
Pieris as a whole, the dark ground-colour has to a great 
extent disappeared from the upper surface; several 
species however (as P. pylotis and P. bunie) retain a 
discoidal spot in the forewing, which in some, as P. 
thaloe 2, is connected with a dark streak along the costa, 
and in others, as P. demophile 2, is included in a larger | 
remnant of ground-colour which passes as a dark fascia 
obliquely across the wing from the costa to the outer 
border. The underside of the hindwing in this genus 
tends to lose the Catasticta-like character preserved in 
P. locusta, in consequence of a general paling which takes 
effect first in the basal half and spreads outwards (as in 
P. demophile), until in such species as P. bunice it reduces 
the wing to very much the same condition as that seen 
in Ganoris brassice or G. rapi. The ground-colour in 
P. buniew and P. pylotis is indeed even paler than in 
these species of Ganoris, being without the powdering of 
black scales which the latter possess; the hindwing 
however of both P. bunie and P. pylotis retains a dis- 
coidal spot, and, as has been already noted (pp. 287, 289, 
note), the predominance of the paler colouring is never 
so great as not to leave unmistakable relics of the yellow 
streaks and red basal patches. 

The genus Leptophobia appears to be a kind of con- 
tinuation of Pieris. The arrangement of light and dark 
ground-colour preserves a parallel course im the two 
genera, and the inclusion of the discoidal spot of the 
forewings in a dark fascia which passes from the costa 
for a greater or less distance obliquely towards the outer 
margin is a common feature in Leptophobia as well as in 
Pieris. In the present genus the underside of the hind- 


320 Dr. Frederick A. Dixey on the 


wings is almost uniformly pale, but distinct indications 
of the primitive red patches are nevertheless still to be 
met with (see p. 287). 

There would also seem to be little doubt that the 
American species allied to M. pyrrha, which are included 
by Mr. Butler* under Mylothris, with the neuration of 
which they agree, are closely related to Pieris. The pattern 
of every one has been more or less altered by mimicry, 
but in those males whose upper surface is unaffected by 
this kind of modification, the arrangement of the dark 
and light ground-colour is seen to present the ordinary 
features of Pieris or Leptophobia (compare, for instance, 
the upper side of M. pyrrha 3 with that of P. thaloe). In 
M. lorena is found an oblique dark bar crossing the 
forewing just as in P. demophile and L. stamnata, this 
mark being utilised, both in M. lorena and the female of 
M. pyrrha, for the formation of the mimetic pattern. 
It has already been shown+ how the primitive yellow 
streaks and red basal patches have been turned to account 
in the same direction on the underside of the hindwing, 
which, from the additional necessity for protection ex- 
perienced by the insect when at rest with its wings 
closed, becomes the most important field for the mimetic 
process. It may be here remarked that the antennz of 
these American forms, while agreeing in character with 
those of Leptophobia, Pieris, Catasticta, Leodonta, and the - 
genera allied to these last, differ conspicuously from those 
of M. agathina, M. poppea, and the other African species 
of Mylothris. 

Hesperocharis diverges somewhat widely from the 
genera last discussed, both in neuration and in the 
character of its antenne; it retains, however, in many 
cases indications of the primitive marginal and sub- 
marginal dark series in a more recognizable form than 
any (see, for instance, the chevrons on the underside of 
H. erota, which represent series 8). The yellow streaks 


* Proc. Zool. Soc., 1872, pp. 36-38. It appears to me that the 
neuration of the American forms of Mylothris is simply that of 
Pieris, minus the third subcostal nervule in the forewing, which 
branch is already almost obsolete in the latter genus. 

t See above, p. 286. The transitional series which is there made 
to begin with M. lypera and M. lorena might easily be carried back 
to P. thaloe, which shows tbe true Pierine pattern all ready, as it 
were, to take on the mimetic condition of the other species. 


Phylogeny of the Pierine. 321 


and red basal patches, present in all species of Hespero- 
charis, are in some (as H. nereis) marked with special 
distinctness. The curious manner in which these have 
been made use of in H. hirlanda for the production of 
a mimetic pattern, has already been fully discussed 
(p. 286). 

ot cannot but think that U. monuste shows by its 
pattern that it stands on a level with Pieris as a 
derivative from the Catasticta group. Its neuration 
hardly differs from that of Pieris, and it would probably 
be more appropriately placed (together with its imme- 
diate allies, U. joppe, U. swasa, etc.) in or near that 
genus than with U. cynis in Mr. Distant’s genus Udaiana, 
as at present in the National Collection. 

The position of the genus Dismorphia is not easy to 
determine. The pattern of those species that appear to 
have undergone least modification may, however, be 
derived without much difficulty from Pieris or Lepto- 
phobia ;* and the persistence in some cases of the red 
basal spots has already been remarked (see p. 285). The 
structure of the antenne points to the same line of 
ancestry. On the other hand, the very remarkable 
neuration is quite unlike that of Pieris; a certain 
approach to it, however, is made by Hesperocharis, which 
is almost without doubt a close ally of that genus. 
Moschoneurat is very nearly akin to Dismorphia, from 
which it differs only slightly in neuration; while the 
Palearctic genus Leptosia has characters which link it 
with both.t We may probably regard the three last- 
named genera as terminal twigs of a branch now lost, 
which left the main stem at or near the genus Pieris, 
and of which Hesperocharis is a still earlier offshoot. In 
Dismorphia and a few species of Moschonewra much of 
the original colouring has been retained and modified for 
purposes of mimicry. In Leptosia and the remainder§ of 
Moschoneura this colouring has given way to the usual 
Pierine invasion of white. 


_ * Asomewhat different and, as it seems to me, less probable view 
is advanced by F. Miiller, Jenaische Zeitschr., x., pp. 1-12. 

+ The figure in Cistula Entomologica, vol. i., pl. iv., fig. 9, omits 
the second discoidal of the forewing. 

t The forewing is nearer to Dismorphia and the hindwing to 
Moschoneura. 

§ Pseudopieris of Godman and Salvin, 


322 Dr. Frederick A. Dixey on the 


2. The evidence of distribution. 


It now remains to briefly indicate the bearing of the 
geographical distribution of the various forms that have 
been mentioned upon the question of their kinship and 
relative antiquity. 

As we have already seen, the oldest form of Pierine 
now extant is probably Huchewra socialis. This insect 
appears to be as limited in its geographical range as it is 
isolated in its zoological position, for itis found only in the 
mountain-ranges of Mexico, which may be considered as 
a southern extension of the ‘ Rocky Mountain” division 
of the Nearctic Region.* Its nearest allies appear to be 
Behr’s two species of Neophasia (see p. 303), which inhabit 
the same region with itself, and the Pontias and Metaporias 
of the high lands of Central Asia, most of which forms 
are known to retain the ancient larval habit of spinning. 
These facts seem to point to the conclusion that Huchewra 
is the relic of an archaic group of Pierines which once 
occupied the great mountain regions of both the Palz- 
arctic and Nearctic continents, and whose immediate 
descendants, still represented in the Hast by Metaporia 
and Pontia, have in the West become extinct (unless 
Behr’s Neophasia be a survival) after giving origin to 
the group of genera headed by Catasticta. 

From one or other of these two primary stems, the 
Kastern or the Western, nearly the whole of the existing 
genera of Pierines may be derived. There are, however, 
a few exceptions, which, perhaps, constitute relics of an 
ancient Pierine fauna coeval with the groups above 
mentioned, but’ not, like them, the progenitors of a 
numerous and varied offspring. ‘The chief of these are 
the genera Hlodina and Nychitona, the former of which 
is entirely confined to the Australian Region, while the 
latter has a very wide distribution throughout the 
Kthiopian, Oriental, and Australian. The African 


* T here follow Mr. Sclater’s division of the earth’s surface into 
six Zoological Regions, which arrangement, adopted by Mr. Wallace 
in his ‘ Geographical Distribution of Animals,” 1876, has stood the 
test of time and experience better, in my opinion, than any alterna- 
tive distribution that has been proposed. I also adopt, for conve- 
nience, the smaller divisions, or “sub-regions,” as determined by 
Mr. Wallace in the above-named work, 


Phylogeny of the Pierinee. 325 


species of Mylothris belong possibly to the same 
category. 

The genera derived from the Catasticta group remain, 
for the most part, within the confines of the Neotropical 
Region. This is the case with Pieris, Leptophobia, 
Hesperocharis, the American species of Mylothris,* Dis- 
morphia, Moschoneura, and those species allied to 
monuste at present included in the genus Udaiana.t 
It is remarkable that the Nearctic Region does not 
furnish a single species that can be supposed to be 
derived from the present stem.{ Inasmuch, however, 
as the Palearctic genus Leptosia seems to belong 
rather to this than to any Eastern branch, the conjecture 
may be hazarded that connecting forms now extinct 
once occupied the Nearctic Region, from which the 
Palearctic continent received the forerunners of its 
present Leptosias, probably by way of Behring Strait. 

Turning now to the Eastern Metaporia, which inhabits 
the borderland between the Palearctic and Oriental 
Regions, we find it emitting one clearly-defined branch 
in the Palearctic direction. This is the branch to 
which belong the various species of Pontia, as P. nabel- 
lica, P. soracta, P. hippia, P. belucha, P. leucodice, and 
P. crategi. In the Chilian or Andesian division of the 
Neotropical Region we find the genus Tatochila, which 
appears not to belong to the regular Neotropical Pierine 
stock, but to be closely related to the Palearctic 
Pontias. It is conceivable that the latter stem may 
have spread from Asia into the western portion of the 
Nearctic continent, and thence down the mountain- 
chains to the south. Neophasia menapia, at present 
inhabiting the Californian and Rocky Mountain sub- 
regions, seems to me to be more nearly allied to both 
Pontia and Tatochila than (as Behr thinks) to Huchetra 
socialis, and may very possibly be a relic of the original 
invasion, Another indication of the same invasion is 
afforded by the genus Phulia, now found with the 
nearly-allied Tatochila only in the Andesian or Chilian 


* See p. 320, note. 

+ See p. 321. 

+ The instance of U. monuste, which straggles into the southern 
districts of the Nearctic Region, hardly forms an exception to the 
above statement. 


324 Dr. Frederick A. Dixey on the 


sub-region, to which it no doubt made its way along the 
great mountain-chains in a similar manner. Its close 
ally, Baltia, remains in the high lands of Central Asia, 
where it bears much the same relation to Synchloe as 
Phulia to Tatochila. Another early offshoot from the 
Kastern Pontia stem is Mesapia peloria,* which has no 
representative in the Western Hemisphere. 

The above descendants of the mountain Metaporias 
belong, as has been seen, in the first place, to the 
Paleearctic and western portion of the Nearctic Region, 
only reaching the Neotropical by extension along the 
chain of the Andes. Other derivatives of Metaporia, 
however, took their course directly southwards. The 
first of these is the Delias and Prioneris group, the more 
ancient members of which are, speaking generally, to be 
found in the northern portion of the Indian peninsula, 
while the Australian and other southern forms represent, 
as a rule, a somewhat later stage of development. 
Another is the important branch headed by Hwphina, 
which genus, like Delias, has spread downwards through- 
out the Oriental Region, and by way of the Indo- 
Malayan and Austro-Malayan islands to the Australian 
continent. The Australian species of Hwphina are 
clearly derived from the Oriental, and those forms (such 
as H. phryne ?) which are nearest to M. agathon in 
colouring are also its closest neighbours geographically. 
Of the two genera (Hiposcritia and Catophaga) which 
appear to be immediately derived from Hwphina, the 
former is confined to the Oriental Region; while the 
latter, hke Huphina itself, has spread along the Austro- 
Malayan Islands to the Australian mainland. This is 
also the case with Appias,t the origin of which genus 
from the Catophaga stock is no doubt to be assigned to 
the Oriental’ Region. But, unlike the other genera, 
Appias seems to have extended its borders westwards, 
and to have given rise to the “ Phrissura B”{ group in 
the Ethiopian Region, and even to Glutophrissa in the 
Neotropical. If this be the real origin of these two 
latter genera, we have to enquire how they reached the 
African and South American continents respectively. 


* See p. 304, 

T It would seem, however, that no true Appias actually reaches 
the Australian continent. 

{ See p. 309, note. 


Phylogeny of the Pierine. O20 


With regard to the first, there is little difficulty in 
supposing the passage to have been effected by land 
either now or formerly existing, the Ethiopian Region 
having been in this, as in so many other instances, first 
entered from the north. But it is not easy with our 
present knowledge to imagine an overland passage for 
these butterflies from the Oriental or Ethiopian Region 
to the Neotropical. The northward route, which we saw 
to be the one probably adopted by the ancestors of the 
Chilian Phulias and Tatochilas in spreading from Central 
Asia, is excluded in the present instance by the entire 
absence of any trace of such a passage from both 
the Palearctic and Nearctic Regions; and although a 
transit by way of a formerly existing “ Antarctica” is 
conceivable, it would seem more likely that the crossing 
from east to west was effected in the region of the tropics. 
After all, however, the difficulty of supposing an Atlantic 
sea-passage is not overwhelmingly great. The unusual 
facilities possessed by insects for crossing large extents of 
sea have been remarked by many writers,* and among 
insects the butterflies of the Pierine group are especially 
given to migration for great distances in countless hordes.t+ 
It is worth noting that in the case of each of the other 
three chief Pierine genera whose present distribution 
seems to have involved one or more long sea-passages, ?.€., 
Terias, Colias, and Callidryas, special observations exist 
of their migratory propensities. In 1874 a large swarm 
of Terias lisa reached the Bermudas from the American 
continent ;{ the swarm of butterflies described in a well- 
known passage|| by Mr. Darwin consisted chiefly of a 
species of Colias; while descriptions of the migratory 
fight of Catopsilia and Callidryas are numerous, 
among the most striking pieces of testimony being that 


* See especially Wallace’s “Geographical Distribution,” 1876, 
vol. i, p. 32; and the same author’s “ Darwinism,” 1889, p. 359, etc. 

+ Trimen, Trans, Ent. Soc. Lond., 1870, p. 382 ; “ South-African 
Butterflies,” 1887, vol. iii, p. 32. Moore, “ Lepidoptera of 
Ceylon,” 1880, 81, p. 116. Distant, “ Rhopalocera Malayana,” 
1882—86, p. 285, etc. Mr. Trimen suggests that there is “an 
evident connection or relation between these wonderful migrations 
of certain species of Pierine and the well-known habit of nearly 
all the members of the Sub-family of flying straight onward in 
one direction.” 

+ “ Psyche,” Dec. 1875, p. 121. 

|| “ Voyage of the Beagle,” ed. 1860, p. 158. 


326 Dr. Frederick A. Dixey on the 


of Mr. Spruce, who saw Callidryas “launching boldly 
out over the Pacific Ocean.’’* 

The earliest species of Synchloe were undoubtedly 
differentiated from Pontia or Baltia in the Palearctic 
Region, from which the genus spread (probably east- 
wards) into the Nearctic. Synchloe proper can hardly 
be said to enter the Indian Region,+ but in its progress 
westwards it has sent an offshoot downwards into the 
Ethiopian, consisting of S. johnston and S. hellica. 
S. glauconome of Arabia and Egypt remains to mark the 
course of the invasion. Ganoris, a further Palearctic 
development of Synchloe, has accompanied that genus 
into the Nearctic Region and has also spread into the 
Oriental. A curious extension of the range of the 
Palearctic G. rapx into the Nearctic Region has been in 
progress during the last thirty-three years, the first 
transatlantic specimens having been seen at Quebec in 
the year 1860.{ ‘This introduction was undoubtedly 
effected by human agency, and differs from the natural 
passage of species between the two Regions in having 
taken place by the Atlantic instead of the Pacific route. 

Though Synchloe itself is far more characteristic of 
temperate than of tropical districts, it has given rise to a 
large and important Pierine branch which has spread far 
and wide through tropical and temperate parts alike. 
The birthplace of Herpexnia and Teracolus, the two 
earliest members of this extensive section, is apparently 
the eastern portion of the Mediterranean division of the 
Palearctic Region; from which locality the former has 
spread through Arabia and Abyssinia into the African 
continent, while the latter has not only followed Herpenia 
into Africa, but has also largely occupied the two western 
Oriental sub-regions. Those forms of the genus Izias 
that show least divergence from Teracolus are found in 
the Nile provinces of Eastern Africa, but the bulk of this 
genus has moved eastwards, its distribution being 
characteristically Indian. A few species, however, are 
found in some of the Indo-Malayan islands, and in 
Austro-Malaya as far east as Timor. 


* Journal Linn. Soc., Zool., ix., pp. 355—357. 

+ See Wallace, Trans. Ent. Soc. Lond., 3rd series, iv., pp. 242, 3. 

+ Scudder, “Butterflies of the Eastern United States,” 1889, 
vol. i1., pp. 1175-1190 ; Edwards, “ Butterflies of North America,” 
vol. 1., 1868-72, sub. voc. P. virginiensis. 


Phylogeny of the Pierines. 327 


To the same borderland of Western Asia and Eastern 
Africa may be assigned the place of origin of Nepheronia, 
which has sent a western branch into Africa, and an 
eastern into the Indian peninsula and Malayan islands, 
one species reaching the Australian continent. The 
African branch has given rise to Hronia proper. Attention 
has already been drawn to the curious fact that the form 
which links the Hronia group most closely with Teracolus, 
viz., H. (?) lucasw, survives in Madagascar. Hebomoia, 
another offshoot of this part of the Pierine stock, is now 
almost entirely Malayan ; its place of origin was, however, 
in all probability further west. The birthplace of Huchloe 
is problematical, but the present distribution of the 
charlonia group, which seems to contain the oldest 
members of the genus, would appear to make it probable 
that the Mediterranean sub-region witnessed the rise of 
this, as of so many other more or less direct descendants 
of Synchloe, from which central area it successively over- 
spread the Palearctic and Nearctic continents. The 
isolated geographical position of Hroessa chilensis is very 
remarkable, its affinities being apparently with Hastern 
rather than Western forms. It is probably, as before 
suggested, a solitary survival of a once more widely- 
spread group, among which were to be found the 
immediate ancestors of the present-day Huchloes. 

No other genus in the whole sub-family has so extensive 
a range as Colias, species of which are found in every one 
of the six great Zoological Regions. Here again, I have 
little doubt that the site of original divergence is Asiatic, 
and is most nearly represented in the present condition of 
‘the earth’s surface by the borderland between the 
Palearctic and Oriental Regions on the north-west 
frontier of India. From this centre one or two species 
have ranged into South Africa and the Indian peninsula ; 
but the greater number have turned northwards, and 
after populating the Palearctic and Nearctic continents 
with numerous species, have penetrated to the circum- 
polar lands within the Arctic Circle, have passed down 
the great mountain chains of Central and South America 
to Chili and Patagonia,and have even established outposts 
in Venezuela and the Sandwich Islands.* 

The powers of dispersal possessed by the genus Terias 


* The occurrence of Colias in the last-named locality is, how- 
ever, not entirely free from doubt. 


328 Dr, Frederick A. Dixey on the 


are almost as remarkable as those of Colias ; perhaps 
even more so when we take into account their weak 
fight, and the fact that their migrations must have been 
intertropical Mr. Wallace, however, has drawn 
attention to their habit of frequenting “ gardens and 
plantations and skirts of forests rather than their deeper 
recesses,” and also of “assembling on the margins of 
streams and on the sea beach,”? and has remarked that 
“these habits lead to their being frequently carried olf 
by winds,” and that ‘‘it is thus perhaps that some of the 
species have so wide a range and offer such perplexing 
variations.”* Whatever may have been their means of 
dispersal, there can, I think, be no doubt that they took 
their rise from the Colias stock in the Western Hemi- 
sphere, the line of descent passing through Xanthidia 
to Terias and Sphenogona; Pyrisitia, Nathalis, and 
Lewcidea being given off by the way. All these genera 
are mainly Neotropical with Nearctic extensions. Terias 
itself, however, as is well known, so far from remaining 
within these limits, has overspread the warmer portions 
of the Hthiopian, Oriental, and Australian Regions, and is 
even found in the Manchurian sub-region of the Pale- 
arctic. 

It seems on the whole most probable that the origin of 
Gonepterya is also to be referred to the Western Hemi- 
sphere, where Meganostoma marks the transition from 
Colias. Gonepterya itself seems to have passed to the 
north by way of California and so across into the 
Palearctic Region, while Rhodocera and Amynthia re- 
present a Neotropical development of the same stock, the 
Central American genus Kricogonia perhaps remaining ° 
near the original seat of divergence. Gonepterya having 
reached the Palearctic Region has extended to its 
westernmost extremity. Its only offshoot appears to be 
Dercas, which probably arose in the debateable Man- 
churian area, where the Palearctic and Oriental faunas 
are much mixed, and thence spread southwards through 
the Indo-Chinese sub-region to Sumatra and Borneo. 

Catopsilia and Callidryas, like Terias, must, it would 
seem, have undergone intertropical migration. Their 
oldest forms appear to be Catopsilia florella, C. hybleza, 


* Trans. Ent. Soc. Lond., 3rd series, iv., p. 320. See also 
above, p. 325. 


Phylogeny of the Pierine. 329 


C. thauruma, etc., which are probably derived from the 
Neotropical genus Amynthia, coming nearest to A. 
clorinde. Inasmuch as all these are African insects, and 
the New World Callidryas, Pheebis, and Metwra are less 
closely allied to Amynthia, it seems necessary to suppose 
that the earliest forms of this group in the Neotropical 
Region have become extinct, the present Callidryas 
group surviving as their modified descendants; while an 
early dispersal of these ancestral forms took place across 
the Atlantic to Africa, of which invasion C. florella, etc., 
remain as comparatively unmodified relics. The Oriental 
and Australian Catopsilias are the ultimate developments 
of this invasion. 

The distribution of Belenots is remarkable, the bulk of 
the species belonging to the Ethiopian and Australian 
Regions. .The Oriental Region is poorly supplied, except 
for the abundant B. mesentina, which is found in all 
parts of the Indian peninsula, and even enters the 
Mediterranean district of the Paleearctic Region as far as 
Asia Minor. Notwithstanding the present poverty of 
the Oriental Region in species of Belenois, it seems 
probable that this area is really the birthplace of the 
genus, which, as we have seen, appears to be derived 
from that primitive part of the Pierine stem now repre- 
sented by Delias and Prioneris. B. mesentina and 
B. taprobana, of India and Ceylon, may probably be 
regarded as survivors of the original race of Belenois, 
whose descendants have spread south-eastwards to 
Australia, and south-westwards to Madagascar and the 
African continent. Pinacoptery« is in all probability a 
local modification of elenois within the Ethiopian 
Region, while Daptonwra, whose history is otherwise 
hard.to. account for, may perhaps have originated from a 
branch of the African Belenois which at some remote 
period found its way westwards across the Atlantic. 


In concluding this paper I wish to express my great 
indebtedness to several friends who have given me 
valuable help during its progress. It was by the kind- 
ness of the late Professor Westwood that I was enabled 
to begin the study of the Pierine group in the Hope 
Collection at Oxford, and the facilities for work afforded 
me by him have been continued and increased by his 
successors in the charge of the department, first by 


TRANS, ENT. SOC. LOND. 1894.—part 11. (JUNE.) Y 


330 Dr. Frederick A. Dixey on the 

Mr. W. Hatchett Jackson, and now, in a special manner, 
by Professor Poulton, F.R.S., the present occupant of 
the Hope Chair, at whose request I have undertaken the 
arrangement of this part of the Hope Collection. I am 
also under great obligations to Colonel Swinhoe, who 
has very kindly helped me in various ways, and 
especially in the determination of the EHastern species, of 
which he has so intimate a knowledge. Lastly, my 
thanks are due to the members of the staff of the 
Natural History Department of the British Museum, 
particularly to Messrs. A. G. Butler, W. F. Kirby, and 
F, A. Heron, who have given me every assistance in 
examining the fine series of Puerimxe preserved in 
the National Collection. 


ITV.—Inpex or Species MENTIONED. 


The Order is that of the National Collection in the 
British Musewm. 


PIERINA. 


Prioneris thestylis, Doubl., 258, 267, | Delias harpalyce, Don., 257, 284, 


281 
clemanthe, Doubl., 258,281,284 
vollenhovii, Wall. . . 284 
autothisbe, Hubn., 258,281,284 
sita, Feld. oe ee (BA 


Delias eucharis, Drury, 2D1, 204, 


268, 280, 284, 288, 300, 310 
hyparete, Linn., "57, 280), 284, 


300 
hierte, Hiibn. . . 284, 300 
agostina, /Tew. . 268, 300 
mysis, abr. . . 284, 301 
isse, Cram. . 284, 301 
echo, Wall. 284 


argenthona, Fabr., 284, 288 
301 
philotis, Wall. . . . 300 
ceeneus, Linn., 268, 284, 288, 
300 


duris, Hew. 284 
dorimene, Cram. 301 
belisama, Cram., 277, 284, 
300, 301 

descombesi, Boisd., 268, 277, 
284, 300, 301 
257, 284, 300, 
301 


nigrina, Fabr., 


” 


300, 301 

aganippe, Don., 257, 277, 
284, 300, 301 

thisbe, Cram., 257, 258, 280, 
281, 283, 300 

belladonna, Fabr., "257, 258, 
267, 280, 299, 300 

nysa, Fabr. . 280, 301, 310 


momea, Boisd., 268, 280, 301 
egialea, Cram. 284 
pasithoe, Linn., 257, 258, 267, 

281, 283, 300 
crithoe, Boisd., 277, 281, 284 
orphne, Wall. . 301 


Pereute autodyca, Boisd., 281, 293 


” 
PP) 
” 


” 


swainsonil, Gray . 285, 294 
charops, Boisd., 281, 285, 293 
leucodrosime, Koll., 281, 285, 

294 


telthusa, Hew. 294 


Euterpe tereas, Godt., 258, 281, 284, 


295, 298 
critias, Feld., 284, 295, 298 


rosacea, Butl. . 295 
bellona, Cram. 295 
theano, Boisd. 299 
eurytele, Hew. 299 


Phylogeny of the Pierinee. 


Mylothris lorena, Hew. 
pyrrha, Fabr., 286 288, 307, 320 
lypera, Koll. 285, 288, 289, "390 
chloris, Fabr. . 279, 289, 307 
agathina, Cram., 268, 279, 289, 

307, 308, 320 
, poppea, Cram., 268, 308, 310, 


320 

,, vhodope, Fabr.* . 308, 310 
trimenia, Buti, wre oe MOOG 
Blodina egnatia, Godé.. .307; 362 
Nychitona xiphia, Fabr., 507, 322 


Hesperocharis hirlanda, Stoll, 286, 


. 286, 320| Terias lisa, Boisd. & Lec., 276, 325 


dol 


* deva, Doubl. 276 
» mandarina, De POre. 013 
»,  floricola, Boisd. . 276 
» hecabe, Linn. . 276 
»,  excavata, Moore . 313 
» Mmessalina, Fabr. . 313 
» agave, Cram. . , 283, Bias) 
»  delia, Cram. 313 
» Thodia, Feld. 313 

elathea, Cram. 313 


Sphznogona gratiosa, Doubl. & 
Hew. . 276,3313 


288, 321 mexicana, Boisd., 283 

marchalii, Guwér. 286 Pyrisitia proterpia, Fabr., 276, 313 

‘ erota, Luc. . 287, 320| Gonepteryx cleopatra, Linn, 259, 

3 anguitia, Godt. . 286 272 
nereis, Feld.. .286, 321 _ rhamni, Linn., 259, 264, 

Mpeheiva socialis, Westw., 25 995 290; 272, eli 328 
308, 322 Amynthia meerula, Fabr., "260, 272, 

Leodonta zenobia, Feld. . 281, 285 398 
» tellane, Hew. . 259, 285 clorinde, Godt., 260, 272, 
» dysoni, Doubl. 259, 281, 285 315, 329 
Catasticta nimbice, Boisd., 259, 281,| Dercas wallichii, Doubl., 260, "973, 
292, 295 314, 328 

» colla, Doubl. Zo 202 », lycorias, Doubl., 260, "273, 
,, bithys, Hiibn., . 259, 285, 291 275 

» anaitis, Hew. 259, 281, ae Meganostoma philippa, Fabr., 265, 
291 — 314 

» Semiramis, Duc. . .281, 291 Bs cesonia, Stoll., 265, 314 

», toca, Doubl. . ae oe 292 | Teracolus calais, Cram. . soe 
» notha, Luc. . 293| _,, dynamene, Kiug.. . . ‘311 
,, teutamis, Hew. . 294} ,, amata, Fabr. . 261, 266 
. ctemene, Hew., 291, "294, 319| ,, protractus, Buti. . wa owe 
» corcyra, Feld. . 291, 292, 319| ,, phisadia, Godé. hk aAeeOo 
troezene, Feld. . sue 201)|. |, puellaris) But, 215, SLL 
Nathalis iola, Boisd. . S1D)| (,; tenis, Ainge. Adds 203 
Colias cunninghamii, Butl. 7A Us vesta, Reiche : Sy Va) 
» Meadii, Edw... . 270| ,, ‘hewitsoni, Kirb. , 261, 266, 275 
» fieldii, Ménétr. ZO 5, tripunctatus, Bul. 312 

) edusina, Meld. . . . 270) ,, pleione, King. . 311 

, edusa, Mabr.. 259, 264,271] ,, coelestis, Swinh. 312 

» phicomone, Esp.. .259, 264) ,, subfasciatus, Swains. . 317 

» yale, Linn. . 259,264,270] ,, regina, Trim., 261, 266, 275, 

» subaurata, Butl. . reroks 279 
», barbara, H. Edw. 270| ,, ione, Godé., 261, 266, 275, 279 

oe pelidne, Boisd. . <. » 270) ,, danae, Habr. . 261, 275 

3, paleno, Linn., 259, 264, 270,| ,, eupompe, Klug. . .283, 312 
212, 216, 318| ,,° wallengrenu, Butl. . 312 

Xanthidia nicippe, Cram., 276, 283,| ,, etrida, Boisd. 261 
313| ,, evanthe, Boisd. 275 


* Probably not a Mylothris ; see Trimen, 


flies,” vol. iii., 1889, p. 35, note. 


“ South-African Butter- 


332 


Teracolus eucharis, Fabr. 261 
» evarne, Klug. . . d11, 316 

5, theogone, Boisd. 312 
», antevippe, Boisd. 283 
,, omphale, Godt. 275 
» evippe, Linn. 275 
Txias eulimene, Klug. . 312 
»  venatus, Butl. . 312 
5 evippe, Drury . 312 

» Marianne, Cram., 261, 266, 
275, 312 

Hebomoia glaucippe, Linn., 261, 
275, 317 

Ptychopteryx lucasii, Grandid., 316, 
ali, 318, 327 

Rhodocera leachiana, Godt., 260, 


265, 272, 273, 328 
Callidryas thalestris, Hiibn., "254, 


274 

‘ philea, Linn., 254, 260, 
273, 315, 325 

a senne, Linn. ~ old 

: eubule, Linn. .274, 315 
Metura rurina, Feld. . .316, 329 
Phoebis trite, Linn.. 260, 274, 329 
»  agarithe, Boisd. 315 


Aphrissa godartiana, Swains., 260, 
273, 316 

Catopsilia flava, Butl. . 280, 266 
catilla, Cram. . 260, 273, 274 
phlegeus, Wall. 260 
hyblea, Boisd., 328 
thauruma, Reak., 273, 317, 329 
» florella, Fabr. . 273, 315, 328 
Kricogonia lyside, Hiibn., 314, 328 


3) 


Leptophobia eleusis, Lue. . 287 
rf elodia, Boisd. 287 

A stamnata, Luc. . 320 
tovaria, Feld. 287 


Pieris "habra, Doubl. . . 281, 286 


5 locusta, Feld., 281, 286, 289, 
319 
., thaloe, Godé., 287, 319, 320 
» demophile, Linn., 282, 319, 
320 


pylotis, Godt., 277, 289, 319 
bunie, Hiibn., 277, 282, 287, 


319 

Hiposcritia pandione, Hiibn., 277, 
305 

» lalage, Doubl., 256, 277, 306 
Catophaga ega, Borsd. . . 805 
Ks zamboanga, Feld., 256 

Bs paulina, Cram. 305 


- 


Dr. Frederick A. Dixey on the 


Catophaga lankapura, Moore . 


256 

A galena, Feld. . . 305 

ye jacquinotii, Wall., 305 

33 alope, Wall. 305 
Appias celestina, Boisd. . 269, 305 
»  Clementina, Feld. . 305 


nero, Fabr. . . . 256, 305 


hombronii, Tue. - 283 

»  lyncida, Cram. 269, 283 
zelmira, Cram. . a) ee 
Glutophrissa poeyl, Butl. 309 


” 


margarita, Hiibn., 279, 309 
castalia, Fabr. .  . 309, 310 
» Saba, Mabrs . 309, 310 
Saletara panda, Godt. . 805 

» cyclmna, ee: 305 
Phrissura phaola, Dowbl. . ae 
sylvia, Fabr. . 279, 309 
eudoxia, Cram.. . . 309 
coniata, Butl. «ale 
illana, Feld.. . .308, 309 
lasti, Grose Smith, 308, 309 
nagare, Grose Smith, 308 
majungana,Grose Smith, 308 
isokari, Grose Smith . 308 
polisma, Hew.'. .308, 309 
Daptonura lycimnia, Cram. 279, 329 


” 


Belenois helcida, Boisd. . 289 
»  thysa, Hopf. . 308 

»  peristhene, Boisd., 256, 268, 

280, 310 

»,  taprobana, Moore 329 

»  auriginea, Butl. . 279 


mesentina, Cram., 255, 268, 
276, 279, 283, 310, 329 
calypso, Drury, 268, 276 
creona, Cram. . - 216 
teutonia, Fabr., 256, 268, 
276, 279, 280 

coronea, Cram., 256, 268, 


276, 280 
»  gidica, Godt. 280 
severina, Cram. 256 


Synchloe daplidice, Linn., 253, 254, 
259, 263, 267, 270, 278, 306 


a glauconome, Klug., 306, 
311, 326 
» johnstonii, Crowley, 268, 


306, 310, 326 

hellica, Linn., 255, 264, 
268, 270, 278, 279, 306, 
310, 311, 326 

callidice, Hisp., 253, 254, 
264, 268, 270, 306 


Phylogeny of the Pierine. 


Synchloe protodice, Boisd. & Lec., 
264, 268, 271 

chloridice, Hitbn. 306 
Tatochila autodice, ’ Hitbn. , 254, 264, 
270, 272, 283, 306 

a theodice, Boisd., 254, 264, 

210, 210, 306 

Phulia nymphula, Blanch., 307, 323 
Neophasia menapia, Feld., 304, 307, 
323 

» terlootii, Behr., 303, 307, 322 

) Sp. Behr. 303, 307, 322 
Pontia crategi, Linn., 257, 267, 277, 
278, 303, 306, 323 

»  hippia, Brem., 278, 303, 323 

5 soracta, Moore, 257, 267, 277, 
279, 303, 323 

» belucha, Marshall, 303, 323 

» leucodice, Hversm., 303, 306, 
323 

»  nabellica, Boisd., 323 
oberthiiri, Theech-. .. 303 
Mesapia peloria, Hew., 304, 307, 324 
Baltia shawii, Bates . 307, 324 
a putleri, Moore 307, 324 
Metaporia agathon, Gray, 257, 267, 
277, 279, 302, 304, 324 

i phryxe, Boisd., 277, 279 
Ganoris napi, Linn., 252, 253, 263, 
»  bryoniz, Ochs. . 270 

» brassice, Linn., 252, 253, 263 


303, 


»  gliciria, Cram., 253, 264, 
278 

» rape, Linn, 252, 253, 263, 
326 

Pinacopteryx pigea, Boisd. 279 
S larima, Boisd. . 256 

0 capricornus, Ward, 
268, oho) 311 


Huphina cassida, Fabr., 305 
»  phryne, Fabr., 257, 277, 


309 
Huphina eperia, Boisd. 304 
» naomi, Wall. . 305 
Me lea, Doubl. . 305 
5 aspasia, Stoll, . . 258 
» judith, Fabr. . 258, 305 
i nabis, Lucas. . . 305 
nama, Moore. . 258, 304 
Udaiana cynis, "Hew. . 808, 321 
»  monuste, Linn. . 321, 323 

5 ‘Suasa, Boisd. . a2L * 
»  joppe, Boisd. . Pee 4! 
Herpeenia eriphia, Godt. 279, 283 
“ tritogenia, Klug. dll 
i lacteipennis, Buil. . 311 
Nepheronia valeria, Cram. 262 
” gea, Feld. . 267, 277 
5 iobea, Boisd. . 262 
” thalassina, Boisd., 266, 
210, 316 
argia, Fabr., 279, 308 
Eronia cleodora, Hiibn. . 316, 318 
», leda, Doubl., 262, 266, ice 
318 
Euchloe belia, Linn. . . 318 
»  eupheno, Esp. 261 


»  cardamines, Linn., 261,276 
»  reakirtil, Edw. . 261 
»  genutia, ’ Fabr. ~ 206,018 
»  belemia, Esp. . 276 


5»  ausonia, Hiibn., 261, 266 
= ausonides, I BT IE MNS) 
»  hyantis, Hdw. . 261 
»  tagis, Hiibn. . 261, 266 
»  charlonia, Donz., 276, oe 


lucilla, Butl., 261, 276, 318 
: 318 


Zegris eupheme, Esp. 
Leptosia sinapis, Linn... 321, 323 
Eroessa chilensis, Gueér. _ 318, 527 


Moschoneura methymna, Godt., 321 
Dismorphia melia, Godt. . . 285 


304, 324 = astyocha, Hiibn.. 285 
»  coronis, Cram. = 200 i praxinoe, Doubl, 285 
»  timnatha, Hew., 258, 277 ce nemesis, Latr. 285 
DANAIN&. 
Lycoreaatergatis, Doubl.& Hew.299 | Ceratinia (Ithomia) dionxa, Hew.299 
HELICONIN&. 
Heliconius melpomene, Linn., 294, | Heliconius hydara, Hew. . 294 
296 : phyllis, Fabr. . 294 
‘4 thelxiope, Hiibn, . 295 5 burneyi, Hiibn. 295 
7 vulcanus, Butl, . 294 


304 On the Phylogeny of the Pierine. 


PAPILIONINA. 


Papilio nephalion, Godé. . . . . 298 
zacynthus, Fabr. . 295,298 

a2 | polemmeren, Godt. . 285,298 
euterpinus, Godm. & Salv., 294 


” 
INCERT#A SEDIS. 

Styx infernalis, Staudgr. . . . 290 

Pseudopontia paradoxa, Meld. . 290 

Davidina armandi, Oberthr. . . 304 


Exeianation or Puates III., IV., & V. 


PATH Air 


Fic. 1. Eucheira socialis. 
2. Catasticta bithys. 

3. C. ctemene. 

4, Delias belladonna. 
5. D. eucharis 9. 

6. D.nysa o. 

7. Metaporia agathon. 
8. Huphina phryne ¢. 


PLATE IV. 


Fic. 9. Catophaga paulina 9°. 
10. Appias clementina 9. 
11. Hiposcritia lalage ¢. 
12. Mylothris agathina. 
13. Belenois mesentina ¢. 
14. B. peristhene. 
15. Synchloe daplidice 9. 
16. Ganoris napi ¢. 


PLATE V. 


Fic. 17. Teracolus hewitsoni. 
18. Ixvias marianne °. 
19. Hebomoia glaucippe 9. 
20. Colias hyale @. 
21. Catopsilia crocale, var. flava 9. 
22. Callidryas philea 9 .* 
23. Eronia leda 9. 
24. Euchloe charlonia. 


These figures are only designed to illustrate the actual points 
mentioned in the text. They are not intended to show specific 
characters. The same letters and numbers stand for the corre- 
sponding markings throughout all the figures. See explanations 
in the text, pp. 254, 264, 269, 273, 274. 

Norr.—In Fig. 15, S 8 is placed one space too high up. 


_* This figure was drawn from a specimen in the Hope Collec- 
tion, which had been erroneously labelled C. thalestris. The error 
has unfortunately found its way into the plate. 


(oa5ni 


X. Some notes on the Micro-Lepidoptera whose larve are 
external feeders, and chiefly on the early stages of 
Eriocephala_ calthella (Zygenidx, Limacodide, 
Hriocephalide). By Tomas ALGEeRNon CHAPMAN, 
M.D., F.E.S, 


[Read Feb. 7th, 1894.] 
Puates VI. & VIL. 


I present these notes now, as there seems little pro- 
bability that I shall, at an early date, be able to carry 
further my observations on WHriocephala, and it is 
desirable to put them, so far as they go, at the disposal 
of anyone who may wish to continue them; and I pre- 
sent along with them sundry notes that tend to confirm 
the view, that the Zygxenidxe, Limacodidx, and Hrioce- 
phalide form a group which, though the last member is 
as low as the lowest 7imex, and the first as high as Butter- 
flies or Noctuze, has nevertheless been evolved on its own 
lines, from a common source, as a separate branch of 
Heterocera. 

The broad facts of the pupal structure, and the slug- 
like form and habit of external feeding of the larva, pro- 
bably suffice to support this view; but further points of 
alliance are no doubt most useful in confirmation. 


\ 


It becomes necessary, in the first place, to relate my 
observations on Hriocephala calthella, since it is largely 
to these observations that 1 owe the confidence I feel 
that this grouping of families is justified by fact. 

The genus Micropterya was divided by Stainton (“‘ A 
Monograph of the British Species of the Genus Micro- 
pteryx of Zeller,’ Trans. Ent. Soc. Lond., new series, 
vol. i1., pp. 26-40, 1850-1) into two sections—A. Hrioce- 
phala, of Curtis, and B, Micropteryx, Hiibner. At that 
time the larvee were unknown, and the division was made 
by the neuration. Shortly after, the larvaof Micropteryx, 
Hiibner, was discovered, and very soon material was accu- 
mulated that would have justified what must now be 
done, viz., separating the Hriocephale from the Micro- 
pteryges, not only as a distinct genus but asa separate 


TRANS. ENT. Soc. LOND. 1894.—part it. (JUNE.) 


336 Dr. T. A. Chapman’s notes on Micro-Lepidoptera 


family ; and the following might be a brief outline of the 
characters, distinguishing them from each other and from 
other forms. 


Micropterygidx. Ovum: Ovoid in shape, delicate, trans- 
parent, and nearly colourless; laid in the parenchyma of 
a leaf. 

Larva: Leaf-mining, without legs. 

Pupa: In a dense subterranean cocoon, escaping there- 
from for emergence; segments of abdomen all “ free; ” 
has large jaws, used for assisting the pupa out of cocoon, 
like those of Trichoptera, but larger and more elaborate. 

Imago: Large six-jointed maxillary palpi, no trace of 
jaws, 9th and 10th abdominal segments of ¢ form a knife 
arrangement for piercing leaves for oviposition; 8th is 
an external ordinary (but terminal) segment. (In Adelidx 
the 8th is absorbed in the piercing apparatus, and the 
7th is the external terminal segment.) 


Hriocephalide. Ovum: Spherical, opaque, covered with 
a snow-like coating, laid externally. 

Larva: Short, square, and angular, with 8 rows of 
globular appendages, and 8 pairs of abdominal legs, an 
anal sucker, long antenne, feeding exposed. 

Pupa: (Probably not unlike a Nepticula, and in a 
cocoon above ground.) I have only seen the head and 
antenna piece, and cannot build up the whole pupa from 
that with any confidence. 

Imago: Six-jomted maxillary palpi, used as feeding 
hands: well-developed, serviceable jaws; ovipositor simple, 
tubular, of three pieces, last abdominal segment the 7th. 
‘There is also the neurational difference noted by Stainton. 


I propose, then, to accept and accentuate this division, 
leaving the name Micropteryx for the “higher” group, 
whose larve have been known for so long, and as to 
whose oviposition and corresponding structure of the 
imago I had the pleasure of assisting my friend, Dr. 
Wood, in his researches, though in view of my observa- 
tions on their pupx, I think it must be very doubtful 
whether they can be called the “higher” group any 
longer. 

The LHriocephalide attracted my renewed attention 
when engaged in working out the oviposition of the 
Micropteryges. In the far-off days, when the life-histories 


a 


whose larve are external feeders. 337 


of Micropterye were being recorded in the ‘¢ Annuals,” 
I was much impressed with the resistance of the Hrioce- 
phalide to investigation; but looking, as I then did, 
on the Tineina generally as subjects fit only for the work 
of past masters, I had little idea that they would con- 
tinue this resistance for so many years, and that it would 
be left for me to make the first impression against it. 

It was evident, on comparing the structure of calthella 
with that of purpurella, or other Micropterye—as indeed 
it was to a great extent without that knowledge—that 
the habits of Hriocephala must be very different from 
those of Micropteryw. 

I believe my observations have been made on both 
calthella and seppeila, and I have not always been very 
careful to determine which species I had in hand, and 
may have had others. But in the observations I have 
made I have not determined that any decisive differences 
were observable. 

In calthella, then, the ovipositor is quite short, simple, 
and tubular; no knives, or rods, or other complex appa- 
ratus, as in purpurella. The egg, therefore, must be 
laid in a very easy, simple way. This is confirmed by 
noticing that the egg is large and spherical, not, there- 
fore, to be slipped into any narrow chink or crevice. It 
seemed also certain that it was not laid on the leaves 
of any plant, so I assumed it must be laid in moss or 
rubbish, on the ground. By providing a number of 
moths with such materials in May, 1891, I succeeded in 
obtaining a large number of eggs, and in hatching the 
young larve; but I altogether failed to induce them to 
eat. The egg and young larve are, however, so very 
remarkable—so unlike all our ideas of a Lepidopterous 
insect—that even so meagre a measure of success was 
worthy of note. I delayed doing more than mention the» 
result for two reasons. Firstly, because one of our lead- 
ing—perhaps I ought to say our leading lepidopterist— 
was pleased to suggest that the beast I had under 
observation was an Julus, or a Crustacean, or some un- 
known monster, and that repetition was very desirable, 
with many precautions, to make sure I had Micropteryz, 
i.e. Hriocephala, and nothing else. This advice I was 
very willing to follow, because not only was it obviously 
a great compliment to the extraordinary nature of the 
observation, but as a second reason for delay I hoped to 


338 Dr. T. A. Chapman’s notes on Micro-Lepidoptera 


be able to make my observations more full, and even 
perhaps to rear the larvee. 

I accordingly, in June, 1892, secured a good supply of 
moths and placed them in many different jars. In three 
of these I gave them nothing but moss, which I had 
obtained in winter, then cleaned and boiled it, and kept 
it in a dry condition until it was wetted and given to the 
moths. Hgos were freely laid in this sterilised moss as 
in the other jars. I may say that I myself considered this 
precaution to be absolutely unnecessary, as the eggs laid 
agreed precisely in size and form with those dissected 
from the moth—they were seen against the deceased moth 
that had laid them, they were laid in several different 
jars, and finally Dr. Wood had, by repeating my procedure, 
induced moths to lay, and had in fact confirmed all 
my observations. However, the sterilised moss experi- 
ment has been made successfully, and several dozen 
batches of eggs have been laid, so that there is no room 
for the most severe scepticism to suggest a doubt. 

In 1892 I supplied the moths with flowers of Ranun- 
culus repens (common buttercup), and though I believe 
observations have been made on the mouth-structures 
and habits of these moths, in which they are in truth very 
anomalous, as in so many other ways, I may mention 
my own observations. They use their great claw-like 
maxillary palpi with sharp knife points to scrape and 
tear at both the pollen of the stamens and the surface of 
the petals, in the latter case perhaps collecting fallen 
pollen. They certainly do something very like eating as 
regards this pollen, and digest and use it, as would appear 
from two circumstances: firstly, that very slender moths 
get very fat and lay many eggs, and, secondly, their 
dejecta are very abundant. There is obviously room for 
much detailed observation in this matter, which J did 
not make, as I wished all my material to be devoted to 
making sure of eggs, and I had not too much time to 
give to the subject. 

Moths will live in confinement for three weeks if fed 
in this way and kept damp enough. 

They pair readily, and apparently do so more than 
once. When laying, the female moth creeps down 
among the moss to a depth of an inch or more, and 
seems anxious to get to the bottom and lay her eggs on 
the bottom of the jar, sand or earth. If the stratum is 


whose larve are external feeders. 339 


too deep for this she will lay them on a spray of the moss, 
always in little groups, rarely as few as two or three, 
usually six to ten and even more. I have counted as 
many as twenty-five. She will often remain and die 
beside her last batch of eggs. I prepared various jars 
with mosses of different sorts, and especially several with 
growing Mniwm, which I thought might be the proper 
food, but in this I proved to be mistaken. 

About the middle of August, 1892, I found several larve 
of calthella about 1°5 mm. in length in ajar in which some 
moss was growing, but I could see none in my specially 
prepared pots of Mniwm, nor indeed in any of the other 
jars; these larva, though so much larger than the newly- 
hatched ones, were precisely the same in all respects, so 
far as I could see without disturbing them. They were 
kept very wet, the moss dripping and the sand below 
under water, in consequence of my directions to keep them 
moist during a temporary absence being misunderstood. 
The result, however, was decidedly satisfactory, and at 
the end of September the moss was still growing, and 
there were two larve usuaily to be seen of a length of 
about 2mm. On October 21 I found there were three 
larvee in this jar, the two largest measuring with the head 
retracted 3°5 mm. in length, and therefore probably full 
grown. 

I also found several about 1:3 mm. in length in another 
jar in which moss was growing, but which had been kept 
much too dry. In one of the Mniwm jars there was also 
one about 1:5 mm. long, seen for the first time close by 
where a bit of moss (not Mniwm) was growing. I was 
on this date successful in making two other observations. 
One was in seeing one of the larvae in my best jar actually 
eating the delicate stem of growing moss, of which a con- 
siderable piece of one side had disappeared. The other 
was due to one of the larvee having got on to the glass, 
when not only was it easy to see that it resembled the 
newly-hatched larvee as to processes, abdominal legs, etc., 
but that it also possessed the trefoil sucker. These 
larvee seemed to be constantly on the move, at times 
not to be seen, and when seen always in a fresh place ; 
they walk slowly but steadily, using the abdominal 
appendages as legs, and often raising the fore part of the 
body and stretching about as if in search of something. 

They are not cannibals, as one walked straight along 


340 Dr. T. A. Chapman’s notes on Micro-Lepidoptera 


another and neither attempted to injure the other in any 
way. As judged by the sizes of their heads, there were 
at least three sizes of larvee on October 21st, which with 
the newly-hatched one makes four sizes ; but there is little 
doubt that there would be one or two intermediate sizes 
between the young larve and the smallest seen at this 
date. The intestinal contents, so far as they are visible 
through the larva, were in one instance green, in two 
others brownish. Occasionally a good end view of the 
larva is seen, and then its angularity (on cross section) is 
very evident, the spaces between the double rows of 
processes being hollow and the processes placed on the 
angles of a flat, raised surface. None of the drawings I 
have bring out this peculiarity quite satisfactorily. 

These larve preserve also the long antennz; these 
have an elegant curvature, and are placed on the head so 
as to look, as it moves from side to side, ridiculously 
like the horns on a Hereford ox—the proportionate 
length of horn to head being not very different in the 
two cases. 

The moss on which these larvee thus happened to be 
reared was named for me by the Rev. A. Ley as 
Hypnum prelongwm, a common species. ‘There were 
also present, though it was doubtful if these were 
growing, two other common species, Hypnum tamari- 
scinum and swartzii, and possibly others. 

I must admit that I was nearly as much astonished at 
finding the full-grown larva retain the peculiar structure 
of the young larva, as I was when I first saw the newly- 
hatched larva. Having so few and wishing very much to 
obtain the pupa if possible, I did not sacrifice any of 
these, for closer examination; but one that died, of 
2°5 mm. long, though not quite satisfactory, gave me a 
good view of the abdominal legs and ball appendages ; 
the head was unfortunately retracted and could not be 
satisfactorily made out. ‘All these larvae perished during 
the winter, and left no trace, except some remains that 
make a rather poor slide. 

In 1893 I prepared eight jars with various mosses and 
succeeded in having plenty of eggs laid in them, but for 
some reason or other the young larve, which hatched 
abundantly, appeared to have all perished, until I was 
pleased and surprised early in November to observe two 
full-grown larvee in one of the jars. 


whose larve are external feeders. 341 


In January, 1894, I observed some threads of Isaria 
in this jar, and found they proceeded from what turned 
out to be a cocoon of calthella, with a dead full-grown 
larva curled up inside; the cocoon was fairly tough, of 
yellowish silk and with scraps of moss coating it, ovoid, 
3mm. long and 1°5 broad. ‘The contained larva was 
somewhat damaged by the fungus and in removing it 
from the cocoon. The abdominal legs are all present, 
but the structure seems a little more modified, either 
really or by the Isaria, than in the two-third grown larva 
already referred to. The antennz are very long, the 
first long joint being very long, the second reduced as 
compared with younger larve. The ball appendages are 
proportionally rather smaller. The dots in the sulci 
between are now large, round, smooth, disc-like plates, 
comparing markedly with the rough surrounding skin, 
and having a central pigment spot. 


The ova are spherical, 0°46 mm. in diameter, of 
calthella almost white; of seppella 0-41 mm., a very 
little smaller and distinctly yellow. They have a snowy, 
mealy look, owing to a provision of a close coating of 
minute rods standing vertically on the surface of the ege 
and often tipped with a small bulb (of fluid?) ; whether 
these are adpressed to the surface of the egg when laid, 
or whether they afterwards develop in some way, I 
do not know, but I think the latter. Their function 
would appear to be to protect the egg from too close 
contact with the possibly very wet surface on which 
it lies. 

The young larva is difficult to examine owing to its 
delicacy, to its retracting its head when disturbed, and to 
its rapid shrivelling by desiccation, when removed from 
its natural habitat in damp moss. 

The peculiarities of its form and structure may be 
stated to be its angular outline, the possession of a 
number of remarkable appendages to each segment, of 
eight pairs of abdominal legs of unusual structure, and of 
an oval sucker; that the antennz are remarkably long 
for a lepidopterous larva, and that the head is retractile, 
so far, that it may occupy the interior of the 2nd thoracic 
segment. 

The larva does not appear to alter these characters 
during its growth to maturity. The antenna of the adult 


342 Dr. T. A. Chapman’s notes on Micro-Lepidoptera 


larva are not perhaps proportionately so long, and the 
abdominal legs have shorter and thicker bases. 

The description, therefore, and figures of the larve, 
though chiefly taken from preserved specimens of the 
young larva, are not probably far out, if applied even to 
the full-grown one. 

~The larva is thick and short and fairly cylindrical, 
apart from its angular section, and tapers very little, ter- 
minating rather abruptly at either end, especially when 
sulky and with retracted head. The angular outline is 
due to eight rows of peculiar appendages, so disposed 
as to form two subdorsal rows and two lateral rows 
on either side, each double row arising from the 
angles of a raised ridge, and the intervening spaces 
being rather hollowed. The general surface is raised in 
ridges, or rather marked by sulci that are chiefly trans- 
verse in direction, but communicate with each other to 
form a network, and in places forming a beautiful rosetted 
pattern. 

The general result is a division of each segment into 
five subsegments, the balls or appendages are on the third 
of these. The fourth and fifth, in the centre of the 
dorsum, and again in the centre of space between the 
subdorsal and lateral pairs of ridges, are united into 
one by a circular area, in the centre of which is a dot or 
spot. 

This description applies to the 2nd and 3rd thoracic 
and 1st to 7th abdominal segments. The Ist thoracic 
segment has two transverse rows of ball appendages, with 
six in the first row and four in the second. 

On the 8th abdominal segment the appendages are 
similarly in two rows, but deficient in number, there 
being only eight altogether ; whilst on the 9th segment 
are only six. 

These appendages on 8 and 9 abdominal are longer, 
larger, and more club-shaped, and project backwards from 
their points of attachment; those on the other segments 
shorter and more rounded, are directed forward—those 
on the Ist thoracic are, however, similarly a little larger 
and longer than the others. 

These appendages arise from special ball-like points, 
encircled by a special area, and are globular in form, or 
in some cases nearly pyriform, with a definite neck or 
stalk ; they are dotted as though with spiculz in a rect- 


whose larve are external feeders. 343 


angular pattern, and appear to have some internal ribbing 
or skeleton, which remains stiff in a mounted specimen, 
whilst the surface loses its plumpness and shrivels. The 
abdominal legs are eight pairs on the first eight abdominal 
segments, arranged so as to suggest that this double row 
is a modification of the double rows of appendages on the 
upper surface. 

These legs are, however, of a very different structure 
from the balls of the upper surface, and also from the true 
lees on the thoracic segments. 

They consist of a long stem or shaft with, both towards 
its base and apex, some doubtful spicular projections ; 
this shaft 1s apparently cylindrical, and contains a body 
which is either a vessel or tube, or a tendon to the ter- 
minal claw. ‘The shaft arises from a conical base, to 
which it is not distinctly jointed, but is rather continuous 
with it. 

The terminal piece is a very distinct and separate 
structure, in general outline much like the terminal joint 
or claw of a thoracic leg, but with the sharp apex rounded 
off; the convexity is directed forwards (not inwards). It 
is of homogeneous semi-transparent texture, but marked 
by oblique lines, which suggest, whether correctly or not 
I cannot say, that the surface between them is raised in 
rounded ridges, which pass round the anterior and 
posterior edges and make it look like a conventional 
cornucopia. In preserved specimens the shafts of these 
legs seem laterally compressed. 

Another structure is ‘a sucker on the under surface of 
the 9th and 10th abdominal segments, its form is trefoil 
with one leaf forwards, or possibly only on 10th, the 9th 
being very narrow in front of it. 

This sucker, on a lepidopterous larva, is of course very 
unusual, and is a further point of relationship to Limaco- 
dids. The 14th segment carries dorsally two hairs. 
The true legs, besides the base, which is a little full and 
raised, consist of three joints, and much resemble the 
thoracic legs of an ordinary lepidopterous larva. The 
first large joint has two spurs on its inner margin, about 
the middle, and two or three hairs on the same zone, 
laterally and dorsally ; the second more slender joint is 
rather longer, and narrows about the middle, where there 
is a spur or bristle on its inner margin; on the outer 
side, at its extremity, is an indication of a bristle or two, 


344 Dr. T. A. Chapman’s notes on Micro-Lepidoptera 


but no definite appendage. The last joint is again rather 
shorter, and terminates in a sharp point. 

The head is rather longer than broad, and narrows a 
little forwards ; there are two strong mandibles, with four 
brown teeth. The antennz are very long, about equal in 
length to the transverse diameter of the head; there are 
two short thick basal segments, as to the first of which I 
am not very sure whether it is a true segment or a basal 
projection; there are two long segments about equal in 
length, and a nearly as long terminal joint, which is little 
more than a seta in thickness. ‘T'wo pairs of palpi are 
also visible—two and three-jointed, apparently those 
usual in lepidopterous larvee, but I have not defined their 
relations. There is also a central point (spinneret ?). 

I have also one observation bearing on the pupa. A 
moth that I placed on a slide was found to have a defec- 
tive antenna, and was accompanied by the head-piece of 
the pupa case, which was of the “ Incomplete” or 
“Micro” type, that is, consisted of the covering of 
antennee, head, and head-appendages in one piece. This 
observation renders tolerably certain what was antece- 
dently probable, that the pupa is of “micro” type, with 
3rd and following abdominal segments free. 


The pupal structure of Zygena and of Limacodes 
showed them to be micros, of a rather early type, whilst 
their ova also presented peculiarities nowhere to be met 
with among macros, and though not at all resembling 
closely those of riocephala, not at all unlike some 
Adelids. 

The larve of these two groups, however, present very 
wide differences from other micros. 

The only other micro-larva having similar form, and 
the habit of not mining or feeding internally, or under a 
web, was curiously that of Hriocephala. Unfortunately, 
though the pupa of Zygzena and Limacodes are of nearly 
the same micro-type, and that a low one, and Hriocephala 
must also have a pupa of low micro-type, I have, after 
trying to obtain it for three years, failed to do so. It 
cannot be taken for granted that it is of the same type as 
they are. We are therefore deprived, for the present, of 
the light that would throw on these relationships. It 
occurred to me, however, that if this relationship was 
real, and not a mere resemblance, some other points 


whose larve are external feeders. 345 


of similarity of structure and habits would be dis- 
coverable. 

The first point that occurred to me was that traces of 
abdominal legs, like those ef Hriocephala, might perhaps 
occur in newly-hatched larvee of some species of Zygxena 
or Limacodes. So: far, I have failed to detect such a 
structure, but find it recorded that the larva of Lagoa 
crispata possesses additional abdominal legs. I have not 
yet succeeded in obtaining eggs of this species. The 
pupa is unquestionably Limacodid, though Packard calls 
it a Liparid. Though many systematists consider Liparids 
and Limacodids to inosculate, the pupze prove them to be 
about as far apart, phylogenetically, as they well can be. I 
succeeded in obtaining eggs and young larve of Parasa 
chloris, and in rearing one larva; but though this is a 
most curious and interesting larva, it did not present any 
extra abdominal legs. It confirmed, however, the obser- 
vations on Limacodes testudo, which afforded me some 
very curious facts, and some very unexpected confirmation 
of the suspected relationship to Hriocephala. 

Liimacodes has suckers to the first eight abdominal 
segments, though the first and last of these are poorly 
developed ; these suckers are probably homologous with 
prolegs, and also with the eight pairs of abdominal legs 
of Hriocephala. 


When the larva of Limacodes testudo has completed 
its development within the eggshell, which is very easy 
to watch, owing to its transparency, flatness, and the 
facility with which the moth can be induced to lay them 
on glass, it is free from any spines or processes, but at 
the period of hatching certain long spines are rapidly 
developed; of these there are at least four on each 
segment, arranged as a dorsal and lateral series on either 
side. The ordinary tubercles can be detected as faint 
dots, but these spines are quite independent of the ordi- 
nary tubercles, and differently placed, and correspond in 
position to no larval processes I am aware of, except 
those of Hriocephala. 

In testudo the dorsal series on one side, though con- 
sisting of one spine on each segment, has them placed as 
though they were the double rows of Hriocephala, with 
alternate spines omitted, 7.e., the inner spine is wanting 
on the first abdominal segment, the outer one on the 


TRANS. ENT. SOC. LOND. 1894,—PART 11. (JUNE.) Z 


346 Dr. T. A. Chapman’s notes on Micro-Lepidoptera 


second, and so on alternately ; laterally the spines appear 
to belong to the lower row of the lateral series of Hrioce- 
phala, but the upper row is represented on the thorax by 
two spines. 

A further point of resemblance in these spines to those 
of Hriocephala, is in their peculiar spiculate apex, which 
reminds one much of the peculiar stellate spiculz of the 
knobs of Hriocephala. 

The rapid development of these spines occurs in this 
way : In the unhatched larva a circular mark exists at 
the site of each spine, and is the summit of a cylindrical 
body deeply imbedded in the larval substance. This body 
is, in fact, the spine, of which the upper third appears to 
be already stiff and solid, with its apex at the surface 
ready to emerge, whilst the lower two-thirds form a soft 
invaginated sheath surrounding this upper part. As the 
spine emerges, when half of the soft portion has emerged 
there appears upon it a spur, and when the emergence is 
complete, and, in fact, at any time, a distinct joint is 
visible at the base of the upper portion. The soft por- 
tion appears very rapidly to become hard. 

I had the good fortune to observe this emergence take 
place in many instances, and have preserved specimens 
at all the stages. 

These spines are in length about equal in length to the 
diameter of the larva, and are divisible into three portions. 
The basal is rather thick and smooth, and terminates at 
the lateral spur. The middle portion is continuous with 
the basal, and is structurally the same—only narrower— 
andis also quitesmooth. ‘I'he terminal third is separated 
from the middle portion by a joint, or transverse line of 
union, and has a series of minute points or teeth, appa- 
rently arranged in a somewhat spiral manner along its 
shaft, and terminates at the apex ina slight enlargement 
and a coronet of angular points, six to nine in number. 
The lateral points are difficult to see except where taken in 
profile, though I have some preparations showing them 
to be really rather numerous and spirally arranged. 
Like the structural framework of the knobs of Hriocephala 
they are less evident during life. 

‘he inner structure of these spines has all the appear- 
ance of consisting of a separate included tube running 
the whole length, and having a branch to the lateral spur. 

On the first moult these spines disappear, and are 


o 


whose larve are external feeders. S47 


replaced by straight spines, that is, they have a smooth 
outline, and taper continuously from base to apex. They 
have, however, a joint about 3th of their length from the 
base, the apex looking harder, browner, and more solid, 
and they appear to have a central tube. I need not 
allude to their apparent origin from deep tissues, and the 
skin looking like a thick coating of glass, through which 
they come, as this is, I think, a well-known peculiarity 
of these larvee (as also of many Lycenids). 

But the dorsal set of spines are now double, that is, 
the double row, of which the alternate members were 
wanting in the larva of the first stage, is now complete, 
and they remain so even in the adult larva, though they 
are now merely prominences, and not spines. In the 
second and third stages there are, especially in the tho- 
racic regions round the bases of these spines, very minute 
spines, apparently of a structure very similar to the last 
joint of the spines in first stage. 

The spinneret in this larva is remarkable up till the 
penultimate stage, in being not a pointed organ, but 
flattened out like a fish’s tail, and the silk it disposes on 
the leaves for the larva to walk upon, is not a thread, 
but a very thin ribbon. 

This larva has other very interesting peculiarities, most 
of which are, I imagine, well known. These I need not 
touch on, indeed all I am at present interested to touch 
on is the remarkable disposition and structure of these 
spines in the newly-hatched larva, parallel with nothing I 
know of in any other family than the similar arrange- 
ments in Hriocephala. 

Prof. A. 8. Packard has some excellent observations on 
spines of Limacodids, but on none, so far as I know, that 
quite parallel these in structure; and he does not, I 
think, refer to their disposition in the newly-hatched larva 
as similar to that I find in testudo. 

He figures, however, the young larva of Lithodia 
fasciola, which seems to be very like that of testudo, 
though less well-developed, and for this reason, want of 
sufficient material, and insufficient amplification, appears 
not to have noted any of the points I have here drawn 
attention to. 

Among many figures he gives of Ceratocampid and 
other spines, and similar figures elsewhere, and amongst 
my own observations, there are abundant instances of an 


348 Dr. T. A. Chapman’s notes on Micro- Lepidoptera 


inflated or elongated base, carrying an appendage articu- 
lated to it, or several such; but these appendages are 
always simple hair-like or spinous, just as they are in 
testudo after the first skin. 

In the case of Zygena I have failed to detect any 
structures in the young larve I have examined at all 
parallel to these, and must still rely on the structure of 
the egg, the form and habit of the larva, and the very 
primitive form of the pupa, for its alliance with this 
section. 

I have examined the eges of Limacodes testudo, Hetero- 
genea asellus, and Parasa chloris. They are all flat, 
oval, colourless, transparent, with lozenge-shaped net- 
work of cell-structure of the shell, easily seen if examined 
in suitable light and with moderate magnifying power. 
Such eges occur amongst the Micros and in some Pyrales. 
Nothing like them is anywhere met with, so far as I 
know, amongst Macros. 

The pupa also is of evident Micro type. The wing 
and appendage cases are not attached to abdominal seg- 
ments beyond the second. They are not difficult to 
detach, in some species, from each other. The maaille 
are small, but are prolonged outwards, and after pass- 
ing through a narrow neck terminate in a (sometimes 
rather twisted) club between the eyes, antenne, and 
legs. This represents the maxillary palpus, which no- 
where in Macros has any such development. 

Then movement exists in the 38rd and 4th abdominal seg- 
ments and in the g¢ pupa, also in the 7th. Further, the 
larva hes unchanged in the cocoon all winter, and moults 
to pupa in the spring, and the pupa escapes from the 
cocoon for emergence. 

These characters apply to the following species which I 
had alive last spring: Limacodes testudo, Parasa chloris, 
Limacodes scapha, Heterogenea asellus, Hmpretia stimulea, 
and Lagoa crispata. 

I do not know that a detailed description of each 
would carry us much further. ‘They vary in the propor- 
tion of parts, the extent of toothed armature on the back 
of the abdominal segments, and other sculpture. Other 
features that do not so much interest us in the present 
connection are the possession of a beak between the eyes 
(for rupturing the cocoon); the projection backwards 
of the meso-scutellum, so that its sharp apex almost 


whose larvxe are external feeders. 349 


reaches the 2nd abdominal segment ; a very remarkable 
structure that exists in other families, but nowhere else so 
well developed as here, and which I have called an eye- 
flange. Where, in most pupe, the eye abuts against the 
antenne, it is here rather separate, and a flat tlange-like 
margin, with sharp edge, and in some marked with 
radiating lines, surrounds the eye without quite joining 
the antenna. 

In Lagoa crispata the antennz of the male are avery 
marked feature of the pupa. Unfortunately, I had only 
this one pupa, and so failed to obtain eggs. ‘The cocoon 
is very like the others except in one very important point, 
it has a practicable lid. 


In Zygena the egg is of an ovoid form, with a delicate 
colourless and apparently structureless shell. ‘The con- 
tents divide into two portions—a yellow at one end, and 
a nearly colourless at the other, and till one is familar 
with them one is persuaded they are addled at least. 

The pupa (I have examined filipendulx, lonicere, 
trifolui, eeulans) is very different from that of Limacodes 
in form and colour, but in the most essential points the 
number of free segments, the looseness of attachment of 
the appendages, and in the dehiscense, it is practically 
identical. The maxillary palpus is nearly or quite obso- 
lete, and in some other points the appendage-cases have 
a structure differing from Limacodes. The dorsal head- 
cover is still well-developed. 

I may note that Syntomis, placed in Zygenidex or left 
close by in Syntomidx, not only has, as has been often 
remarked, a very Arctioid larva, but, as is demonstrated 
by the pupa, is really an Arctia, with no affinities what- 
ever with Zygxna. The ovum is also an Arctia ovum, 
not indeed very far from Caja, but differing from Zygena 
toto ccelo, 

I am indebted to Dr. W. G. Clements for the privilege 
of examining a cocoon and pupa of Huchromia fulvida, 
a species also credited with Aygenid affinities. The 
pupa in no way resembles Zygena, but though very 
delicate and transparent, affords no characters that I can 
certainly seize to separate it, much more than generically, 
from Lubricipeda. 

It presented a curious Arctioid peculiarity, viz., the 
adherence of the cocoon to the pupa. Caja cocoon, for 


350 Dr. 'T. A. Chapman’s notes on Micro-Lepidoptera. 


example, can hardly be touched without the cocoon 
adhering to the pupa, as if they had been wetted. 


These two instances seem as sufficient as a thousand, 
to illustrate that Zygena has been placed among Arctiid 
families, on the strength of some analogies of the 
imagines, which cannot be homologies, as they do not 
affect the earlier stages. 


ExpLanation oF Pirates VI. & VII. 


PLATE VI. 
Larva of Eriocephala calthella. 


Fic. 1. Larva first skin, slightly grown, dorsal view, x 100 diam. 
2. Larva first skin, newly hatched, dorsal view, X 50 diam. 
3. 5 55 lateral view, X 50 diam. 
4, Antenna, X 600 diam. 
5. Abdominal leg, x 900 diam. 
6. S200 scram: 
Carel Ball appendages, X about 900 diam. 


PLATE VII. 


Eriocephala calthella. 


Fic. 9. Ball appendage and rosetted structure of skin, half-grown 
larva, X about 300 diam. 
10. Thoracic leg, X about 200 diam. 
11, Abdominal leg of larva, two-thirds grown, X about 300 diam. 
12. Ova in moss, X 30 diam., the upper ovum near hatching. 
13. Form of sucker, ventral aspect of 13th and 14th segments. 


Larva of Limacodes testudo. 


Fic. 14. Newly-hatched larva, dorsal view, before emergence of 
spines, X 100 diam. 
15. » lateral view, xX 100 diam. 
16,27, 18: Stages i in emergence of spines, x 100 diam. 
19. Terminal portion of spine, X 150 diam. 
20. Thoracic leg, X 150 diam. 


Fig. 1 from drawing by Mr. A. Hammond, F.L.S., from a living 
larva. 

Figs, 2 and 3 from drawings by Mr. H. Knight, from living larva. 

Figs. 5, 8, 9, 14, 15, 16, 17, 18, 20 from drawings by Mr. E. Wilson, 
from preserved specimens. 

The others from my sketches. 


( 351 


XI. On the Tenebrionide collected in Australia and 
Tasmania by Mr. James J. Walker, R.N., F.L.S., 
during the voyage of H.M.S. “Penguin,” with 
descriptions of new genera and species. By GeorcE 
C. Cuampion, F.Z.S. 


[Read Feb. 28th, 1894. ] 
Puatse VIII. 


Ir is proposed in this paper, which is in continuation of 
one contributed by Mr. Gahan on the Longicornia (Trans. 
Ent. Soc. Lond. 1893, pp. 165—197), to give a list of 
the Tenebrionide collected in Australia and Tasmania by 
Mr. J. J. Walker in the years 1890—1, during the 
voyage of H.M. Surveying-Ship “ Penguin,” reserving 
for the present the Cistelide, Melandryidze, Anthicide, 
and the remaining families of the Heteromerous series, 
and also the Tenebrionidz subsequently obtained by him 
in other places. The material examined has been for- 
warded by Mr. Walker partly to myself and partly to 
the British Museum, but the whole of the specimens 
collected have been examined. Some few species are 
represented in the Museum-set only (mostly single 
examples), and these are specially noted below. Five 
new genera and thirty-three new species are described 
out of a total of 110 species. All the new genera and 
thirteen of the new species are from Tasmania, the 
remaining species being from North-west or West 
Australia. Three genera have not hitherto been recorded 
from the Australian continent, viz., Crypticus, Corticeus, 
and Palorus. Lyphia (= Lindia, Blackb.) is new to the 
Tasmanian fauna. Judging from the collection made by 
Mr. Walker, there is still much to be done in Tasmania, 
not only in the Tenebrionide, but in the other families 
of the Coleoptera. I am indebted to the Rev. T. 
Blackburn for a good deal of assistance in the pre- 
paration of this paper, for the purposes of which I have 
carefully studied the collections of Pascoe and F. Bates— 
both very rich in Australian species, and containing a 
large number of Australian types,—and also the Hope 
Collection at Oxford. 


TRANS. ENT. SOC. LOND. 1894.—ParT 1. (JUNE.) 


302 Mr. G. C. Champion on the Tenebrionide 


CoruLADES. 
Cotulades, Pascoe, Journ. Ent., 1., p. 119 (1860). 


Cotulades fascicularis. 


Cotulades fascicularis, Pascoe, Journ. Ent., i., p. 119, 
t. 7, fig. 5. 
Hab. 'Tasmania—Hobart. 


This insect is closely allied to C. (Tagenia) leucospila, 
Hope, the type of which I have examined, but differs 
from it in having the punctures of the elytral series finer 
and more closely placed, the thorax depressed in the 
middle, etc. Under bark (Walker). 


Docatis. 


Docalis, Pascoe, Journ. Ent., 1., p. 121 (1860). 


Docalis funerosus. 


Tagenia funerosa, Hope, Trans. Ent. Soc. Lond. 1845, 
po Lom 
Docalis exoletus, Pascoe, Journ. Ent., i., p. 121, t. 8, 
fig. 7 (1860). 
Docalis degener, Pascoe, loc. cit., p. 122. 
Hab. Tasmania—New Norfolk, Launceston, Franklin, 
Hobart. 


Found in plenty by Mr. Walker, under loose bark of 
Eucalyptus. Fresh specimens have numerous fascicles 
of white or brownish-white scales on the elytra, these 
not being apparent in worn or dirty examples. The 
insect varies greatly in size. I have examined the type 
of Tagenia funerosa, Hope (which is completely abraded), 
and also that of D. ewxoletus, Pascoe. D. degener is, 
apparently, missing from the Pascoe collection. In the 
Catalogue of Gemminger and Harold, and also in that 
of Masters, 7’. funerosa, Hope, is incorrectly placed 
under Cotulades. 


ELAScUs. 


Elascus, Pascoe, Journ. Ent., i., p. 119 (1860). 
The species of this genus have wholly the facies of 


collected in Australia and Tasmania. 353 


certain Colydiide, i.c., of Corticus and Sarrotriwm. 
They are found under bark of felled Hucalyptus trees. 


Elascus crassicornis. 


Elascus crassicornis, Pascoe, Journ. Ent., i., p. 120, 
be 7 okt onde 


Hab. 'Tasmania—New Norfolk. 


Elascus lunatus. 
Hlascus lunatus, Pascoe, loc. cit., t. 7, fig. 8. 
Hab. Tasmania—lLaunceston and Hobart. 


Three specimens, agreeing with the type in the Pascoe 
Collection. 


EDYLIus, n. gen. 


Mentum strongly transverse, trapezoidal, carinate down the 
middle; mandibles bifid at the tip; last joint of the maxillary 
palpi oblong-ovate, that of the labial pair similarly shaped ; 
maxilla with the inner and outer lobes coarsely ciliated ; labrum 
moderately prominent, emarginate in front ; head large and broad, 
deeply sunk into the prothorax, arcuate-emarginate in front, the 
antennary orbits rather broadly expanded, parallel behind, and 
extending outwardly nearly as far as the eyes, the latter small, 
strongly transverse, and almost entire, the epistoma not clearly 
defined ; antenne not reaching the base of the prothorax, joint 3 
twice as long as 2, shorter than 4, 4 and 5 equal, 6 a little shorter 
and broader, 7—11 widened, 7 as broad as long, 8—10 strongly 
transverse ; prothorax large, transverse, convex, broadly, horizon- 
tally explanate at the sides, with very prominent angles, the 
anterior ones subangularly produced in front, and nearly meeting the 
antennary orbits, the base feebly bisinuate ; scutellum transversely 
triangular, moderately large ; elytra about twice as long, and of 
about the same width, as the prothorax, connate, broad oval, rounded 
at the sides in front, acutely margined, the humeri obtuse, declivous, 
not meeting the hind angles of the prothorax ; prosternum convex, 
rather broad, declivous behind, margined on either side between 
the coxz; middle coxai cavities widely open externally, the 
trochantin large; hind coxe widely separated, the intercoxal 
process broadly rounded at the apex ; epipleurz entire, broad at 
the base, gradually narrowing to the apex ; third and fourth ventral 
segments with coriaceous hind margin ; legs elongate ; anterior 
tibize flattened and dilated, strongly produced at their outer apical 


304 Mr. G. C. Champion on the Tenebrionide 


angle ; intermediate and hind tibie slender, asperate, setose, the 
hind pair slightly bowed inwards in both sexes; tarsi sparsely 
clothed with long bristly hairs beneath, the two hinder pairs rather 
elongate, the first joint of the hind pair nearly as long as 3 and 4 
united, the anterior pair with the basal joints a little stouter and 
furnished with a brush of Jong silky hairs beneath in the male ; 
body oblong-oval, rather broad, apterous, sparsely clothed with long, 
appressed, silky hairs, the prothorax and elytra sparsely ciliate at 
the sides. 


This genus is proposed for a single species which 
appears to be not uncommon in Tasmania. It belongs 
to the “ Pedinides” of Lacordaire. H. canescens has 
much the facies of a small Asda. 


Hdylius canescens, n. sp. (Plate VIII, figs. 5, 5a, b, ¢.) 


Oblong-oval, rather broad, convex, ferruginous or obscure fer- 
ruginous, slightly shining ; above and beneath sparsely clothed 
with long, appressed, whitish, silky hairs, the pubescence denser at 
the sides of the prothorax and on the elytra, and on the pro- and 
epipleuree forming ciliz, the four hinder tibiz also with long hairs 
on their inner face. Head densely, rather coarsely punctate ; pro- 
thorax strongly transverse, much broader at the base than at the 
apex, the sides parallel behind, arcuately converging from about 
the basal third, the apex (viewed from above) very broadly trun- 
cate-emarginate, the base feebly bisinuate, the hind angles obtusely 
rectangular and very prominent, the anterior angles strongly pro- 
duced in front, the surface densely, moderately finely punctate, the 
punctures more scattered on the middle of the disc ; elytra con- 
fusedly punctured throughout, the punctures more diffuse, and 
much finer and shallower than those on the prothorax ; beneath 
shining, closely and rather coarsely punctate. Length 74—8}, 
breadth 4—42 mm. (g @). 


Hab. Tasmania—Hobart. 


Many specimens, found buried in the sand at roots of 
maritime plants, a few feet above high-water mark. 


CESTRINUS. 


Isopteron, Hope, Col. Man., iii., p. 112 (1840). 

Cestrinus, Erichson, in Wiegmann’s Archiv, 1842, i., 
Dey kile: 

Mitua, Hope, Trans. Ent. Soc. Lond., v., p. 56. 


collected in Australia and Tasmania. 350 


I have examined the type of Isopteron opatroides, 
Hope, aud it is inseparable from Cestrinus, Hr. Opatrum 
piceitarse, Hope, also belongs to the same genus. His 
definition, “ Tibiee anteriores dentate,” is incorrect and 
misleading : it, perhaps, refers to the subangular dilata- 
tion of the anterior tibise beneath—a male character of 
some of the members of the genus. As Hope did not 
describe his typical species of Isopteron till 1842 (and 
then under a different name to the one mentioned in the 
“ Manual”), it is not advisable to change Hrichson’s 
name. This genus really belongs to the group 
“‘ Pedinides,” and it should be placed near Blapstinus. 


Cestrinus trivialis. 


Cestrinus trivialis, Hr., loc. cit., p. 173; Lacord. Gen. 
Col., Atlas, t. 53, fig. 6. 

Cestrinus longus, Blanch., in Dumont d’Urville’s 
Voyage au Pole Sud, iv., Ins., p. 156, t. 40, fig. 14. 


Hab. Tasmania—Hobart, Launceston, George’s Bay. 


Originally described from Tasmania. The male has 
the anterior tibiee abruptly widened on the inner side 
from about the middle to the apex, and the basal joints 
of the anterior tarsi a little thickened. Jsopteron 
opatroides, Hope, is closely allied to this imsect, but 
differs from it in having ferruginous antenne and the 
elytral interstices granular. Opatrum piceitarse, Hope, 
is also very near C. trivialis, but it has the sides of the 
thorax less sinuous behind. 


Cestrinus aversus. 
Cestrinus aversus, Pasc., Ann. and Mag. Nat. Hist. (4), 
li., p. 278 
Cestrinus posticus, Pasc., loc. cit. 
Hab. Tasmania—Hobart. 


Numerous specimens; some with fully-developed, 
others with rudimentary wings. The anterior tibiz are 
simple in the male. ‘This and the preceding species 
occurred under stones not far from the sea-shore 


(Walker). 


356 Mr. G. C. Champion on the Tenebrionide 


Cestrinus punctatissimus. 


Cestrinus punctatissimus, Pasc., Ann. and Mam Nat. 
Hist. (4), iii., p. 278. 


Hab. 'Tasmania—Hobart. 


One example. 


Cestrinus brevis, sp. n. 


?. Oblong ovate, moderately convex, opaque, pitchy-black ; the 
upper surface thickly clothed with rather coarse appressed brown 
hairs, amongst which are scattered. yellowish-cinereous hairs ; the 
antenne ferruginous ; the legs fusco-ferruginous, the tarsi ferru- 
ginous. Head short, deeply sunk into the prothorax, densely, 
rugosely punctured, the antennary orbits extending about half- 
way across the eyes ; the epistoma short and limited behind by a 
deep transverse groove, feebly arcuate-emarginate in front ; an- 
tenn about reaching the base of the prothorax ; prothorax trans- 
verse, arcuate-emarginate in front, truncate at the base, widest at 
the middle, the sides obliquely converging thence to the apex, and 
sinuously converging behind, the hind angles rectangular, the 
anterior angles sharp and prominent, the surface coarsely, rugosely 
punctured, the interstices very densely, minutely punctate ; elytra 
a little wider than, and about two-and-a-half times the length of, the 
prothorax, with rows of rather fine, deep, subapproximate punctures 
placed in shallow grooves, the interstices feebly raised, very densely 
minutely punctured, and with fine scattered granules; anterior 
tibize slightly curved inwards, rather slender, the outer apical angle 
obtuse ; anterior tarsi simple. . Length 64, breadth 22 mm. 


Hab. W. Australia—Fremantle and Darlington. 


Two specimens, apparently both females. Less elon- 
gate than C. trivialis, Hr.; the head much shorter, the 
epistoma especially ; the thorax more transverse, and more 
narrowed in front and behind; the seriate punctures on > 
the elytra less coarse, the interstices more sparsely and 
more finely granulated; the pubescence of the upper 
surface not unicolorous. C. brevis is less elongate than 
any of the other species of the genus known to me. I 
have received a specimen of a closely-allied undescribed 
form from the Rey. T. Blackburn. 


collected in Australia and Tasmania. Soir 


Cxmvivs. 


Cxdius, Lacordaire, Gen. Col., v., p. 261 (1859), (nec 
Blanchard). 

(?) Plesioderes, Mulsant et Rey, Mém. Acad. Lyon, x., 
p. 34 (1860). 

Blanchard (Hist. des Ins., ii., p. 13) gives as characters 
for Cedius :—“ Antenne with the last five joints broad 
and compressed ; tibiz slightly widened, not crenulated ; 
body oval.” It is clear, therefore, that he had some 
other genus in view. lLacordaire’s description was 
taken trom Opatrum spheroides, Hope. Plesioderes is 
probably distinct from Czdius, Lac. 


Cedius sphxroides. 


Opatrum spheroides, Hope, Proc. Hnt. Soc. Lond., 
1842, p. 77; Trans. Ent. Soc. Lond., 1845, p. 107. 
Oxdius sphxroides, Lacord., Gen. Col., v., p. 262. 
Hab. W. Australia—Fremantle, Condillac I., Trough- 
ton I., Jones I. 


In the typical examples (2) of Opatrum sphzxroides, 
Hope, in the Oxford Museum, the elytra each appear, at 
first sight, to have two rows of prominent rounded 
elevations, the inner one extending on to the base of the 
thorax : these elevations, however, are partly formed by 
dense fascicles of scaly hairs, and are not always distinct. 
Found rather commonly under stones, near the sea-shore 


(Walker). 


Psrupoc2D1vs. 


Pseudocedius, Blackburn, Trans. R. Soc. S. Austr., 
Ml. gip. 9 (1890). 


Pseudocedius squamosus. 
Pseudocedius squamosus, Blackb., loc. cit. 
Hab. N.W. Australia—Roebuck Bay. 


Found in plenty by Mr. Walker, at roots of grass on 
the sea-shore. Mr. Blackburn’s specimens were obtained 
from the same locality. 


358 Mr. G. C. Champion on the Tenebrionidx 


CDIOMORPHA. 


Cxdiomorpha, Blackburn, Trans. R. Soc. S. Austr , 
X., p. 272 (1888). 


Czxdiomorpha australis. 


Cediomorpha australis, Blackb., loc. cit. 

Hab. W. Australia—H. Wallaby I. in the Houtman’ S 
Abrolhos Group, Fremantle, and Cape Leeuwin. 

Many specimens, found by Mr. Walker on sandy sea- 
shores, at roots of bent grass. This species is said to 
be widely distributed in South Australia. 


Prronotvs. 


Prionotus, Mulsant et Rey, Mém. Acad. Lyon, ix., 
p- 150 (1859). 
Achora, Pascoe, Ann. and Mag. Nat. Hist. (4), iii., 
p. 279 (1869). 
Prionotus serricollis. 
Asida serricollis, Hope, Proc. Ent. Soc. Lond., 1842, 
p. 77; Trans. Ent. Soc. Lond., 1845, p. 108. 
Opatrum denticolle, Blanch., in Dumont d’Urville’s 
Voyage au Pole Sud, iv., Ins., p. 154, t. 10, 
fig. 13 (1853). 

Prionotus denticollis, Muls. et Rey, Mém. Acad. Lyon, 
Xp. Lol, 

Hab. Tasmania—Hobart and Launceston. 

I have examined the type of Asida serricollis, Hope, 
from Adelaide; it does not differ from the Tasmanian 
insect. ‘The lateral thoracic teeth are almost or quite 
obsolete in some specimens. Under stones and logs 
in dry places, common (Walker). 


OPATRUM. 


Opatrum, Fabricius, Syst. Ent., p. 76 (1775) (part.). 

This genus is apparently well-represented in the 
northern parts of Australia, whence very few species, 
however, have been described as yet. Of the five species 
received from Mr. Walker from that continent, four are 
apparently new. Opatrwm seems to be replaced in 
Tasmania by the genus Cestrinus. It may be noted here 
that the descriptions of Opatrum seriatum and O. aus- 
trale, Boisd., are perfectly unintelligible. 


collected in Australia and Tasmania. 359 


Opatrum villigerum. 


Opatrum villiger, Blanch., in Dumont d’Urville’s 
Voyage au Pole Sud, iv., Ins. p. 154, t. 10, 
fies £0; 


Hab. S. Australia—Port Adelaide. 


Two examples, perhaps belonging to this species, the 
original specimens of which were obtained at Raffles 
Bay. 


Opatrum torridum, n. sp. 


Rather elongate, subparallel, moderately convex, black, opaque, 
sparsely pubescent, and usually thickly covered with an adherent 
earthy coating. Head broad, thickly punctured, the epistoma very 
deeply emarginate, the sides of the front broadly, obliquely, sub- 
angularly dilated, and extending more than half way across the eyes, 
the latter rather large ; antenne almost extending to the base of the 
prothorax, the penultimate joints transverse ; prothorax strongly 
transverse, not very convex, moderately explanate at the sides, 
broadly and deeply emarginate in front, strongly bisinuate at the 
base, which is also feebly emarginate in the middle, the sides 
moderately rounded, obliquely converging in front, and slightly 
sinuous behind, the anterior angles sharp and prominent, the hind 
angles sharply rectangular, the surface thickly and finely punc- 
tured, the interspaces densely, very minutely punctate and also 
finely granulate ; elytra about four times as long as the prothorax, 
and a little wider than it, subparallel in their basal half, and with 
subrectangular somewhat prominent humeri, moderately deeply, 
rather finely punctate-striate, the interstices almost flat, densely, 
very minutely punctured and also finely granulate ; anterior tibize 
gradually widening outwardly, their outer apical angle sharp ; 
anterior tarsi sparsely clothed with rather coarse hairs beneath ; 
body fully winged. Length 8i—83, breadth 3}—4 mm. 

Hab. N.W. Australia—Adelaide River. 


Two examples, apparently male and female. This 
species (of which I have also received a specimen from 
the Kev. T’. Blackburn) closely resembles some of the 
common Kuropean forms, such as O. (Gonocephalum) 
rusticum, Oliv. As is frequently the case in this genus, 
the sculpture is almost completely hidden by an adherent 
earthy coating; when this is removed, the minute 
punctuation and the granular elevations of the inter- 
stices of the elytra are easily seen. 


360 Mr, G, C. Champion on the Tenebrionide: 


Opatrum walkeri, n. sp. 


Moderately broad, subparallel, not very convex, black or brown- 
ish-black, opaque, clothed with a short, fine, decumbent, greyish 
pubescence, which is usually hidden by an adherent earthy coating. 
Head rather broad, deeply sunk into the prothorax, the sides of the 
front broadly, obliquely, subangularly dilated (forming almost a 
continuous outline with the sides of the prothorax) and extending 
far beyond the eyes laterally, and about half-way across them 
posteriorly, the epistoma deeply triangularly emarginate in front 
and limited behind by a fine groove, the surface densely, roughly 
punctured ; antenne short, not reaching the base of the prothorax, 
joints 8-10 strongly transverse ; prothorax twice as broad as long, 
feebly convex, broadly explanate at the sides, widest at the middle, 
broadly and deeply emarginate in front and bisinuate at the base, 
the sides strongly rounded at the middle, rather obliquely converg- 
ing in front, and constricted behind, the hind angles acute, the 
anterior angles rather sharp, the surface densely, rugosely punc- 
tured, the interspaces finely granulate and very minutely punctate, 
the disc with traces of a fine median groove ; elytra about three 
and a half times the length of, and at the base a little wider than, 
the prothorax, subparallel in their basal half, trisinuate at the 
base, with prominent rectangular humeri, closely and moderately 
coarsely punctate-striate, the interstices densely, very minutely 
punctate, granulate, and rather convex, the third, fifth, and 
seventh a little more raised than the others ; the legs and under 
surface densely, roughly punctate and pubescent ; anterior tibiz 
gradually widened to the acute outer angles; anterior tarsi 
sparsely clothed with rather coarse hairs beneath; wings fully 
developed ; the ventral segments 1-3 depressed along the middle in 
the male. Length 73—91, breadth 33—43 mm. 


Hab. N. W. Australia—Adelaide River. 


Found in plenty by Mr. Walker. This species may 
chiefly be recognized by the explanate rounded margins 
of the prothorax and the rather convex elytral interstices, 
the third, fifth, and seventh a little more raised than the 
others. In dirty specimens the minute dense punctu- 
ation of the surface is completely hidden, and the granu- 
lations are more distinct. O. walkeri is closely allied to 
O. moluccanum, Blanch., numerous specimens of which 
were obtained by Mr. Walker at the Islands of Amboyna 
and Damma. 


collected in Australia and Tasmania. 361 


Opatrum dispersum, n. sp.’ 


Moderately elongate, subparallel, not very convex, rather 
narrow, pitchy-brown, opaque, somewhat sparsely clothed with 
moderately long, appressed yellowish-cinereous hairs, which on 
the elytra form a very irregular treble series on each of the 
interstices. Head somewhat exserted, densely punctured, the 
epistoma very deeply emarginate and confounded with the front, 
the eyes rather large and completely divided, the orbits narrow 
and rounded off behind ; antennez scarcely reaching the base of 
the prothorax, joints 8—10 transverse ; prothorax convex, twice as 
broad as long, rather feebly arcuate-emarginate in front (sub- 
truncate if viewed from above), strongly bisinuate at the base, 
moderately rounded at the sides, widest a little before the base, 
the hind angles subrectangular, the anterior angles rather obtuse, 
the surface densely, rather finely punctured ; elytra about four 
times the length of, and scarcely wider than, the prothorax, 
subparallel in their basal half, finely and lightly punctate-striate, 
the interstices almost flat, finely and closely punctured, the 
punctures slightly muricate ; tarsi thickly clothed with long silky 
hairs beneath, the anterior pair simple; anterior tibie slender, 
gradually widening outwardly, the outer apical angle sharp. 
Length 64—7, breadth 2?—3 mm. 

Hab. N. W. Australia-—Port Darwin. 


Two examples, apparently including both sexes; three 
others have also been sent to me by Mr. Walker from 
Damma Island. This insect is perhaps generically 
distinct from Opatrum, the tarsi being clothed with 
silky hairs beneath; but in the present imperfect state 
of our knowledge of the somewhat numerous Australian 
species of this group, it is inadvisable to separate it. 
The punctures of the striz are fine and very closely 
placed, not coarser than those of the interstices. It is 
not impossible that this insect may be referable to O. 
seriatum, Boisd., from Radack ; the description of that 
species, however, is wholly inadequate, consisting of 
seven words only. 


Opatrum vagabundum, n. sp. 


Moderately elongate, rather narrow, convex, subparallel, black 
or brownish black, opaque, sparsely clothed with short brownish 
hairs, which are subserially arranged on the elytral interstices. 
Head somewhat exserted, densely, rather coarsely punctured, the 
epistoma very deeply triangularly emarginate and confounded 

TRANS. ENT. 80C. LOND. 1894.—ParT 0. (JUNE) 2a 


362 Mr. G. C. Champion on the Tenebrionidex 


with the front, the eyes small and completely divided, the orbits 
moderately broad and rounded off behind ; antennz extending to 
the base of the prothorax, joints 9 and 10 transverse ; prothorax 
convex, nearly twice as broad as long, rather deeply emarginate in 
front, strongly bisinuate at the base, moderately rounded at the 
sides, widest a little before the base, the hind angles subrect- 
angular, the anterior angles rather sharp, the surface densely, 
somewhat coarsely punctate ; elytra about three and a half times 
the length of, and of the same width as, the prothorax, parallel in 
their basal half, finely and rather deeply punctate-striate, the 
interstices slightly convex, flat toward the suture, sparsely gran- 
ulate, and feebly transversely wrinkled , tarsi sparsely clothed 
with rather coarse hairs beneath ; anterior tibiz slender, gradually 
widening outwardly, the outer apical angle sharp. Length 64—63, 
breadth 2?—3 mm. 


Hab. N. W. Australia—Baudin Island and Roebuck 
Bay. 

Two specimens. LHasily separable from O. dispersum, 
which also has completely divided eyes, by the distinctly 
granulate, rather convex elytrai interstices, and the more 
convex, more coarsely punctured thorax, which is more 
deeply emarginate in front, the broader antennary orbits, 
and the shorter pubescence. 


CryPpticus. 


Orypticus, Latreille, Régne Anim., ed. 1, i1., p. 298 
(1817). 


Crypticus submaculatus, n. sp. 

Elliptic, narrow, moderately convex, glabrous, piceous or obscure 
ferruginous, slightly shining, the elytra each with one or two faint 
oblong spots on the disc at about one-third from the apex, the sides 
anteriorly, and in one specimen the apex also, rufous or rufo- 
testaceous, the legs and antenne rufo-testaceous. Head very densely, 
minutely punctate; antenne short, about reaching the base of the 
prothorax, joints 6—11 strongly transverse ; prothorax strongly 
transverse, the base feebly truncate-emarginate and with oblong, 
distinct fovez, the sides rounded and converging from the base, the 
entire surface densely, minutely punctate ; elytra about two and 
one-third times longer than the prothorax, and at the sides forming 
a continuous outline with it, finely and rather deeply punctate- 
striate, the interstices almost flat, and closely, minutely punctate ; 
prosternum produced, lanciform, and received by the narrow 


S 
[os] 


collected in Australia and Tasmania. 


v -shaped process of the mesosternum; legs moderately long, 
slender, the tibial spurs long; the first joint of the hind tarsi 
elongate, nearly as long as the other joints united. Length 21— 
23 mm. 


Hab. N. W. Australia—Roebuck Bay. 


Two examples, found on the sandy sea-shore. This 
minute species possesses all the chief characters of 
Crypticus. In one specimen the markings on the elytra 
are scarcely visible. ‘he genus has not hitherto been 
recorded from Australia. 


Hyocis. 
Hyocis, Pascoe, Journ. Ent., u., p. 457 (1866). 


The species of this genus live at the roots of maritime 
plants on sandy sea-beaches, according to Mr. Walker. 


Hyocis bakewelli. 
Hyocis bakewellii, Pasc., Journ. Ent. i., p. 457. 


Hab. W. Australia—Fremantle, Albany, EH. Wallaby 
J. in the Houtmann’s Abrolhos Group. 


Sent in plenty from both localities. Mr. Pascoe’s 
description was taken from a single specimen, and he 
gives the colour as “ dark ferruginous.”’ In most of the 
Fremantle specimens the elytra have a common, ir- 
regular, O- or U-shaped mark a little beyond the middle, 
and some spots before and behind it, black. Those from 
H. Wallaby I. are testaceous, the elytra usually with some 
small black spots.* The locality given by Mr. Pascoe is 
Victoria. ‘Two other species of the genus have been 
described by Macleay. 


Hyocis subparallela, n. sp. 

Oblong-oval, moderately convex, opaque; piceous or pitchy 
brown, sometimes with the sides of the head, the sides, base, and 
apex of the prothorax, and some ill-defined patches on the elytra, 
ferruginous ; the upper surface thickly clothed with yellowish- 
cinereous appressed scaly hairs ; the antennze and legs ferruginous, 
the apical joints of the antennz more or less piceous. Head densely, 
rugosely punctured ; antennz short, not Sees the base of the 


* Specimens similar to fee are labelled in Pascoe’s ese don 
HT, punctipennis, Pasc. ; but I am unable to find any published 
description of this insect, 


564 Mr. G. C. Champion on the Tenebrionidx 


prothorax, joints 8—11 much wider than those preceding, 9 and 10 
strongly transverse ; prothorax strongly transverse, widest before 
the middle, the sides moderately rounded and gradually converging 
to the rather obtuse hind angles, the anterior angles somewhat 
prominent, the base feebly sinuate on either side, the disc rather 
convex and witha shallow median groove which becomes deeper 
behind, the entire surface densely, rugulosely punctured ; elytra 
three times as long as the prothorax, subparallel to about the 
middle, deeply punctate-striate (the punctures moderately coarse, 
approximate, and transverse), the interstices narrow (not wider than 
the striz), slightly raised, and thickly, finely punctate, the humeri 
subrectangular ; beneath densely, rugulosely punctured, the punc- 
tures on the abdomen finer than those on the metasternum, Length 
3—3 mm, 


Hab. W. Australia—Fremantle. 


Six examples. Longer and more parallel than ZH, 
bakewelli, Pasc., the thorax not sinuate at the sides be- 
hind, the ser iate punctures on the elytra finer and closer, 
the antennee much shorter, with the ninth and tenth 
joints strongly transverse. In Pascoe’s collection there 
is a much more closely-allied form, labelled H. griseipilis, 
Pasc., type, but I am unable to find the description 
of it. 


[ PHYCOSECIS. 


Phycosecis, Pascoe, Ann. and Mag. Nat. Hist. (4), xvi., 
p. 213 (1875). 

This genus, of which four species were described by 
Pascoe (two from Australia and two from New Zealand), 
has five-jointed hind tarsi, the basal joint of all the tarsi 
being small and not easily seen. It cannot, therefore, 
be retained in the Heteromera. Pascoe refers it to the 
“ Phaleriides ” without hesitation, and does not even 
mention the form of the tarsi. Phycosecis should perhaps 
be placed in the Clavicorn-series, near ‘lrogositida or 
Cucujidee. 


Phycosecis litoralis. 
Phycosecis litoralis, Pascoe, loc, cit., p. 214, nota. 
Hab. W. Australia—Fremantle. 


Described from King George’s Sound. In sand, under 
tidal refuse (Walker). | 


collected in Australia and Tasmania. 365 


TRACHYSCELIS. 


Trachyscelis, Latreille, Gen. Crust. et Ins., iv., p. 379 
(1809). 
Trachyscelis ciliaris. 
Trachyscelis ciliaris, Champ., Ent. Monthly Mag., xxix., 
p. 204. 
Hab. W. Australia—E. Wallaby I.in the Houtmann’s 


Abrolhos Group, Fremantle, and Cape Leeuwin. Many 
specimens. 


Trachyscelis levis. 
Trachyscelis levis, Champ., loc. cit. 


Hab. W. Australia—Port Darwin, Cape Leeuwin, 
KE. Wallaby I., Cassini I., Baudin I., and Adeéle I. 

Found in plenty at Baudin and Adéle Islands, more 
sparingly elsewhere. 


ScYMENA. 
Scymena, Pascoe, Journ. Hnt., u., p. 455 (1866). 


The Rev. T. Blackburn remarks (Trans. R. Soc. 8. 
Austr., x., p. 270), that this genus (as represented by his 
S. australis) belongs to the “ Pedinides,” according to 
Lacordaire’s system; nevertheless, it seems to me, as 
stated by Mr. Pascoe, to be nearest allied to Phaleria, 
which Lacordaire places, with some hesitation, in the 
“ Trachyscelides.” 


Scymena amphibia. 


Scymena amphibia, Pascoe, Ann. and Mag. Nat. Hist. 
(4), v., p. 94. 

Scymena australis, Blackb., Trans. R. Soc. S. Austr., 
Mey pearus! |. 

Hab. W. Australia—Albany. 

Mr. Pascoe’s specimens of 9. amphibia, which I have 
examined, were collected by Mr. Masters at King George’s 
Sound. Under tidal refuse on sandy beaches, common 
(Walker). 


366 Mr. G. C. Champion on the Tenebrionide 


HETEROCHEIRA. 


ileterocheira, Lacordaire, Gen. Col., v., p. 335, nota 
(1859); F. Bates, Trans. Ent. Soc. Lond., 1872, 
p. 266. 
Heterocheira australis, 
Uloma australis, Boisd., Voyage del’Astrolabe, Ent. 11., 
p- 258 (1835). 
Heterocheira australis, Lacord., Gen. Col., v., p. 336, 
nota; F. Bates, Trans. Ent. Soc. Lond., 1872, p. 266. 
Var.: Smaller, duller, and less elongate, the prothorax and 
elytra not so finely punctured, the strize deeper and more coarsely 
punctate, the interstices towards the sides and apex convex ; 
anterior tarsi with the second and third joints considerably widened 
in the male. Length 5—6 mm. 


Hab. N.W. Australia—Baudin I., Adéle I. 


Found in plenty at roots of grass on the sea-shore 
(Walker). 


DieHYRRHYNCHUS. 


Diphyrhynchus, Fairmaire, Rev. et Mag. Zool., 1849, 
p. 445; F. Bates, Trans. Ent. Soc. Lond., 1872, 
p. 267. 

Acanthosternus, Montrouzier, Ann. Soc. Ent. Fr., 1860, 
p- 290. 

Mr. F. Bates refers this genus and Heterochetra to the 
‘‘Diaperides,” without hesitation. He seems to have 
completely overlooked the very close affinity of Diphyr- 
rhynchus and Phaleria. Lacordaire’s Groupe “ Phale- 
riides”? would probably be better removed altogether 
from the ‘‘Trachyscelides,’” and placed as a separate 
section between the “ 'Trachyscelides”’ and the “ Diape- 
rides.” 

Diphyrrhynchus, so far as at present known, has pre- 
cisely the same habits as Phaleria, its species being 
found on the sea shore. It has the epistoma deeply 
emarginate, as in Scymena, and the intermediate joints of 
the four front tarsi are similarly dilatate in the male. 
Heterocheira resembles the parallel forms of Phaleria 
(P. parallela, Woll., etc.), but has the anterior tibiae much 
less widened. Mr. F. Bates (op. cit.) states that in both 
Diphyrrhynchus and Heterocheira, the first four joints of 
the intermediate tarsi are strongly, and those of the 


collected in Australia and Tasmania. 367 


anterior more broadly, dilated in the male than in the 
female: this is not so in any of the species before me, 
the second and third joints only being dilated and the 
penultimate one small. Diphyrrhynchus chiefly differs 
from Phaleria in having a much more prominent tro- 
chantin to the middle coxe. 


Diphyrrhynchus ellipticus, n. sp. (Plate VIII., fig. 1, ¢.) 


Regularly elliptic, convex, pitchy-black with a brassy lustre, 
opaque or slightly shining. Head short, deeply sunk into the 
prothorax, rather convex, finely and sparsely punctured; the 
epistoma very deeply emarginate for the reception of the labrum, 
separated from the sides of the front only by a faint (sometimes 
quite obsolete) oblique groove ; eyes coarsely granulated, oblique, 
small, the lower portion slightly larger than the upper portion, the 
antennary orbits as seen from above) extending nearly half-way 
across them; antennz fusco-testaceous, short, scarcely reaching the 
base of the prothorax, the outer five joints gradually widened, 
8-10 transverse, 11 twice as long as 10, rounded at the apex ; pro- 
thorax short, at the base about two-and-a-half times as broad as 
long, convex, the sides rapidly converging from the base, a little 
rounded anteriorly, and sharply but finely margined, the base feebly 
trisinuate and with a very shallow triangular fovea on either side, 
the anterior angles rather prominent, the hind angles subrectangular, 
the apex broadly and somewhat deeply emarginate, the disc very 
minutely and sparsely punctured, the lateral portion more distinctly 
punctate; scutellum broadly triangular, short, large, minutely 
punctate in front ; elytra regularly convex, scarcely wider than the 
prothorax at the base and forming almost a continuous outline 
with it, very sharply margined.and somewhat rounded at the sides, 
obsoletely and minutely punctate-striate, the strize becoming deeper 
towards the apex, the interstices very minutely, sparsely punctate, 
flat at the base, slightly convex towards the apex, the punctures of 
the striz closely placed and very little coarser than those of the 
interstices ; beneath piceous or pitchy-brown, shining, sparsely 
pubescent, and sparsely, moderately finely punctured ; legs pitchy- 
brown or fusco-testaceous ; the tibiz coarsely roughened and setose, 
flattened and dilated, the anterior pair very broad ; prosternum 
horizontal, widened and produced behind, ovate, the mesosternur 
obliquely carinate on either side and excavate in the middle for its 
reception. ¢@. Anterior and intermediate tarsi with the second 
and third joints broadly dilated. Length 4:—6, breadth 2*—3 
mm. (¢ ®. 


368 Mr. G. C. Champion on the Terebrionidex 


Hab. N. W. Australia—Baudin T., Troughton I., Port 
Darwin. 


Many specimens. Closely allied to D. ovalis, F. Bates, 
from New Caledonia, but more convex, duller, and more 
distinctly punctured, the elytra very finely punctate- 
striate. The elytra, at first sight, appear to merely have 
a series of very shallow, fine, longitudinal grooves, which 
become deeper towards the apex, the punctures in them 
being very little coarser than those of the interstices. 
The punctuation of the head is distinctly coarser than 
that of the prothorax and elytra. The Port Darwin 
specimens are less opaque and rather narrower than the 
others. 


Diphyrrhynchus apicalis, n. sp. (Plate VIIL., fig. 4, ¢.) 
Oblong-ovate, subparallel, convex, castaneous with a faint brassy 
lustre, opaque ; the upper surface very sparsely and exceedingly 
minutely punctate, the punctures on the head a little more distinct. 
Head short, deeply sunk into the prothorax, rather convex ; the 
epistoma very deeply emarginate for the reception of the labrum, 
separated from the sides of the front bya very shallow oblique 
groove ; eyes coarsely granulated, small, almost hidden beneath 
the anterior margin of the prothorax, the antennary orbits not 
extending half-way across them; antennz testaceous, very short, 
about reaching the middle of the prothorax, thickening outwardly, 
joints 7—10 transverse ; prothorax at the base barely twice as broad 
as long, the sides parallel behind, rounded and converging 
anteriorly, and sharply margined, the base very feebly trisinuate 
and witha shallow fovea on either side just within the margin, 
a narrow longitudinal space down the middle impunctate ; 
elytra at the base not wider than the prothorax, parallel to about 
the middle, the sides rounded and converging thence to the apex, 
the surface with regular rows of exceedingly minute punctures, 
which are scarcely distinguishable from those of the interstices and 
on the apical declivity are placed in rather deep striz, the inter- 
stices quite flat to about one-fourth from the apex, slightly convex 
beyond ; beneath, obscure reddish-testaceous, slightly pubescent, 
sparsely, moderately finely punctured ; legs reddish-testaceous ; 
the tibie very broadly widened, coarsely roughened and setose, 
the inner face of the anterior pair smoother; prosternum hori- 
zontal, widened and produced behind, lanciform, the mesoster- 
num slightly excavate in the middle for its reception. g¢. Anterior 
and intermediate tarsi with the second and third joints moderately 
dilated. Length 5 mm. ( 2.) 


collected in Australia and Tasmania. 369 


Hab. W. Australia— Roebuck Bay. 


One example. Hasily distinguishable from D. ellip- 
ticus by its subparallel shape, the shorter antennv, the 
less transverse thorax, its wider tibiz, and the still more 
minute punctuation of the upper surface. The elytral 
interstices are perfectly flat to about one-fourth from the 
apex and thence to the apex abruptly, moderately, con- 
vex; the striz are scarcely visible, except under a 
strong lens, though they are sharply defined on the 
apical declivity. 


CorTICEUS. 


Corticeus, Piller et Mitterpacher, Iter per Pos. Sclav., 
p- 87 (1783). 

Hypophleus, Fabricius, in Schneider’s Neu Mag. Ent., 
eh Boos 24 CZ O1),, 


Corticeus australis, n. sp. 

Moderately elongate, subcylindrical, shining, varying in colour 
from castaneous to black, the head always paler in front ; the 
antenne and legs testaceous or fusco-testaceous. Head closely, 
finely punctate; the eyes very large, oblique, coarsely granulated, 
separated by a space about equalling the width of one of the eyes 
as seen from above; antennz not nearly reaching the base of the 
prothorax, joints 5—11 stout, 5—10 very strongly transverse ; pro- 
thorax convex, not longer than broad, a little narrowed in front and 
behind, the hind angles distinct, the entire surface closely, finely 
punctate ; elytra about two and a fourth times longer than the pro- 
thorax, and a little less closely punctured than it. Length 2} mm. 


Hab. N.W. Australia—Adelaide River. 


This minute species is the first of the genus recorded 
from Australia. It has unusually large eyes, the inter- 
ocular space not wider than the diameter of one of the 
eyes as seen from above. Found under the bark of 
Hicus sp. Many specimens. 


PALORUS. 
Palorus, Mulsant, Col. Fr., Latigénes, p. 250 (1854). 


This cosmopolitan genus is not included in Masters’s 
Catalogue. Its species are sometimes found out of doors, 
under bark. Palorus is probably of Hastern origin. 


370 Mr. G. C. Champion on the Tenebrionide 


Palorus melinus. 


Hypophloeus melinus, Herbst, in Fuessly’s Archiv, v., 
p. 3/7, t. 21, figs. B, b (1784). 

Hypophloeus depressus, Fabr., in Schneider’s Neu Mag. 
Hnt., i:,°1, p. 20. 

Ins unicolor, Oliv., Mnt., my No. 18, gp. 22;98..2; 
figs. 8 a, b. 


Hab. N.W. Australia—Port Darwin, Adelaide River, 
Roebuck Bay. 


Numerous specimens, found under bark, away from 
habitations. ‘These examples are smaller, and have the 
head and prothorax more finely punctured than usual in 
P. melinus. A cosmopolitan insect, occurring all over the 
world; I have similar specimens from such widely- 
separated localities as Damma Island (Walker), Marocco, 
and Mexico. It is probable that two species are confused 
in collections, but I hesitate to separate them at present. 
P. delicatulus, Reitter, from the East Indies, the frag- 
mentary type of which has been lent me by Mr. René 
Oberthtr, is a sherter and broader insect than P. melinus. 
A third species, from Damma Island, at present un- 
described, has been sent to me by Mr. Walker. 


LyYpHIA. 


Lyphia, Mulsant, Opusc. Ent., ix; p. 166 (1859) ; 
Jacq.-Duval, Gen. Col. Hurop., ii., p. 305. 

Lindia, Blackburn, Trans. R. Soc. 8. Austr., x., p. 27 
(1888). 


Lyphia tasmanica, nu. sp. (Plate VIII, fig. 6.) 

Elongate, narrow, subcylindrical, subopaque, obscure ferrugin- 
ous, the head and prothorax infuscate. Head densely, finely 
punctate ; antenne ferruginous, very short, the apical four joints 
abruptly widened and strongly transverse ; prothorax convex, a 
little longer than broad, the sides parallel behind and feebly 
rounded in front, the hind angles acutely rectangular, the disc 
slightly depressed in the middle before the base, the entire surface 
very densely, finely punctate, the punctures showing a tendency 
to become longitudinally confluent ; elytra nearly two and a half 
times as long as, and slightly broader than, the prothorax, densely, 


collected in Australia and Tasmania. OWL 


very finely, confusedly punctate, here and there very distinctly 
transversely wrinkled, and with numerous interrupted darker lines 
resembling faint strize ; legs rufo-testaceous. Length 3;—34 mm. 


Hab. Tasmania—Hobart. 


Two specimens, found under bark. .This insect agrees 
with the description (so far as it goes) of Lindia 
angusta, Blackb., from Port Lincoln, except as regards 
the form of the antenne: as long as the head and thorax 
together in LZ. angusta, very little longer than the middle 
of the head in J. tasmanica. Narrower and smaller than 
the European LF. tetraphylla, Fairm. (= ficicola, Muls.), 
the upper surface much more finely punctured. ‘The 
elytral punctuation is confused, but in certain positions 
indistinct rows of punctures are visible. The dark 
lines apparently show through from beneath. The size 
of L. angusta is not mentioned by its describer.* 


ARRHENOPLITA. 


Oplocephala, Laporte et Brullé, Ann. Sciences Nat., 

Xxill., p. 338 (1831) (nomen przeocc.). 
Arrhenoplita, Kirby, Faun. Am.-Bor., iv., p. 235. 
Hvoplus, Leconte, New Sp. Col., p. 128. 


Two species of this widely-distributed genus have 
already been described from Australia. 


Arrhenoplita pygmea, un. sp. 

¢. Oblong, very convex, castaneous or rufo-testaceous, shin- 
ing, finely pubescent. Head short, very shining, smooth and 
depressed between the eyes, armed on either side above the point 
of insertion of the antennz with a very long, erect, nearly straight 
horn, the epistoma very short, limited behind by a deep groove ; 
the eyes black, large, very coarsely granulated, almost entire ; 
antennz short, not nearly reaching the base of the prothorax, 
joints 3—6 slender, very short, 7—11 greatly widened and forming 
a large 5-jointed club, 7—10 perfoliate, strongly transverse, equal in 
width, 11 very short, much narrower than 10 and closely articu- 
lated to it ; prothorax tranversely convex, nearly twice as broad as 
long, rounded at the sides, densely, finely punctate and with a 
smooth central line ; elytra about two and a half times longer than 
the prothorax, confusedly punctured, the punctures a little coarser 


* The specific name is preoccupied, Hypophleus angustus, Luce., 
from Algeria, being a Lyphia (cf. Bedel, Ann. Soc. Ent. Fr., 1887, 
pid), 


Sie Mr. G. C. Champion ow the Tenebrionidx 


and more scattered than those on the prothorax ; legs short, the 
tibia slender. 9. Head unarmed, with the interocular space less 
shining and somewhat thickly punctured, the transverse groove 
behind the epistoma apparently deeper ; the prothorax narrower, 
less convex, less rounded at the sides, more parallel behind. 
Length 2—2! mm. 


Hab. N.W. Australia—Port Darwin, Adelaide River. 


Numerous examples. This very small species ap- 
proaches the European [phicorynus chrysomeloides, Rossi 
(melanophthalmus, Muls.), but it is best placed in 
Arrhenoplita. It resembles a Cis. The tenth and 
eleventh joints of the antennz are so closely articulated 
as to appear subconnate. Found in dry fungus on old 
posts in company with A. exilis (Walker). 


Arrhenoplita exilis, n. sp. 

d. Oblong, very convex, castaneous, shining, glabrous. Head 
very shining, smooth and depressed between the eyes, and armed 
on either side between them with a long, erect, straight horn, the 
epistoma rather large and limited posteriorly by a shallow groove, 
the anterior margin sharply, triangularly raised on either side in 
front ; the eyes large, coarsely granulated, almost entire ; antenn 
nearly reaching the base of the prothorax, joints 3—6 slender, 3 
much longer than 4, 4—6 short, 7—11 perfoliate, widened, and form- 
ing an elongate gradually widening club, 7—10 transverse, 8—10 
strongly so and much wider than 7, 11 twice as Jong as 10, constricted 
at the middle, and truncate at the apex; prothorax transversely 
convex, nearly twice as broad as long, rounded at the sides, very 
finely, densely punctate ; elytra about two and a half times longer 
than the prothorax, closely, confusedly, very finely punctured. 
?. Head unarmed, densely, minutely punctured, with the trans- 
verse groove behind the epistoma deep ; the prothorax less convex 
and more parallel. Length 2} mm. 


Hab. N. W. Australia—Port Darwin. 


One pair. This species resembles A. pygmexa, but it is 
a little larger and more finely punctured, and destitute of 
pubescence. The antennz are more elongate, and have 
their apical joint fully as wide, and twice as long, as the 
tenth; in A. pygmexa the apical joint is extremely short 
and much narrower than the tenth. At first sight, the 
antennee appear to be 12-jointed, the apical joint being 
constricted at the middle and shaped like the united 
apical two joints in A. pygmea. 


collected in Australia and Tasmania. ane 


PLATYDEMA. 


Platydema, Laporte et Brullé, Ann. Sciences Nat., 
xxi, p. 390 (1831). 


Platydema limacoides. 


Platydema limacoides, Pasc., Ann. and Mag. Nat. 
Hist. (4), i., p. 280. 


Hab. Tasmania—New Norfolk, Franklin, and Hobart. 


A few specimens, agreeing perfectly with the type. 
The locality given by Pascoe is Victoria (?). Under 
Eucalyptus bark (Walker). 


Platydema tetraspilotum. 


Neomida tetraspilota, Hope, Proc. Ent. Soc. Lond., 
1842, p. 78; Trans. Ent. Soc. Lond., 1845, p. 108. 

Alphitophagus tasmanus, Mars., Ann. Soc. Ent. Fr., 
(a), vu, p. 110. 

Platydema pascoei, Macl., Trans. Ent. Soc. N.S.W., 
il., p. 280. 

Hab. Tasmania—Hobart, Launceston, New Norfolk, 
Franklin. 


I have examined Hope’s type in the Oxford Museum. 
Common, under bark (Walker). 


Platydema bicinctum, n. sp. 

Elongate-oval, narrow, very depressed, shining; piceous or 
pitchy-brown ; the front of the head, the sides, base, and apex of 
the prothorax narrowly, and a broad transverse fascia on the elytra 
a little below the base, and another just before the apex, connected 
along the suture, reddish-testaceous ; the antenn, legs, and under 
surface rufo-testaceous. Head densely, minutely punctate; the 
eyes large, coarsely. faceted, oblique, and deeply emarginate ; 
antennz about reaching the base of the prothorax, joints 4—11 
widened, 5—10 transverse, 11 nearly twice as long as 10; prothorax 
convex, about twice as broad as long, widest a little before the base, 
not much broader at the base than at the apex, the sides strongly 
rounded anteriorly, and sharply margined, the base bisinuate, the 
apex (viewed from above) truncate, the hind angles rectangular, the 
anterior angles obtuse, the basal fovez oblique and rather deep, 
the entire surface densely, finely punctate ; scutellum triangular, 
rather large ; elytra nearly four times as long as, and slightly 


374 Mr. G. C. Champion on the Tenebrionide 


wider than, the prothorax, subparallel to a little beyond the middle, 
regularly punctate-striate, the punctures rather fine and very 
closely placed, the interstices flat, densely, very finely punctate ; 
beneath closely, finely punctate, the metasternum smoother ; pro- 
sternum very narrow, parallel, produced behind ; epipleura ex- 
tending:to the last ventral suture; legs slender. Length 3;—3}, 
breadth 14 mm. 


Hab. N. W. Australia—Adelaide River. 


Seven examples. This species is allied to P. tetras- 
pilotum, Hope, but is smaller, flatter, and more 
parallel, the thorax rounded at the sides, and distinctly 
narrowed behind. This last-mentioned character, com- 
bined with its subparallel shape, gives it a different facies 
from most of the other representatives of the genus. 
P. bicinctwm varies in the extent of the light markings 
on the elytra, according to the predominance of the 
light or of the dark colour; the anterior fascia does not 
reach the base, but it sometimes extends to the lateral 
margin. Found under bark of a dead Acacia (Walker). 


Platydema deplanatum, n. sp. 


Oblong-oval, narrow, very depressed, black, shining, the labrum and 
antenne ferruginous, the legs rufo-testaceous. Head closely, finely 
punctate, the eyes moderately large, deeply emarginate ; antenne 
asin P. bicinctum ; prothorax convex, twice as broad as long, widest a 
little behind the middle, not much broader at the base than at the 
apex, the sides rounded and rather sharply margined, the base 
bisinuate, the apex (viewed from above) subtruncate, the hind 
angles rather obtuse, the basal fovez distinct, the surface closely 
and somewhat coarsely punctured, the punctures a little more 
scattered on the middle of the dise ; elytra about three and a half 
times as long as, and slightly wider than, the prothorax, subparallel 
in their basal half, rather coarsely punctate-striate, the punctures 
closely packed and deep, the interstices almost flat, each with an 
irregular row of fine punctures; legs slender. Length 3, 
breadth 14 mm. 


Hab. N.W. Australia—Adelaide River. 


One specimen, found under bark. Allied to P. bicinc- 
twm, but less elongate, the punctuation of the upper 
surface much coarser, the elytral interstices each with a 
single row of punctures only, the thorax more rounded 
at the sides behind, and with more obtuse hind angles, 
the upper surface shining black. 


collected in Australia and Tasmania. SA) 


ENNEBEUS. 


Ennebeeus, C. O. Waterhouse, Trans. Ent. Soc. Lond., 
1878, p. 228. 
Ennebeus, so far as at present known, is confined to 
Tasmania and tropical America,* the species from such 
widely different regions being exceedingly closely allied. 


The species of this genus are found under bark, 
according to Mr. Walker. 


Ennebeeus ovalis. 
Ennebeeus ovalis, C. O. Wat., loc. cit., p. 229. 
Hab. Tasmania—Franklin, Huon River. 
Originally described from Tasmanian examples. 


Finneboeeus australis, n. sp. (Plate VIIL., figs. 2, 2a.) 


Oblong-elliptic, pitchy-brown, shining, the legs and antenne 
rufo-testaceous ; the entire upper surface very densely, minutely 
punctured, and clothed with yellowish-cinereous pruinose pubes- 
cence. Antenne rather long and slender, joints 5—8 subequal, 
longer than broad, 9—11 forming an elongate club, 9 triangular, 
slightly longer than broad, 10 and 11 transverse, 11 very abruptly 
truncate at the apex; prothorax strongly transverse, rapidly 
narrowing from the base, the latter deeply sinuate on either side 
of the middle; elytra more elongate than in #. ovalis, with 
numerous scattered coarser punctures, which are partly arranged 
in irregular series. Length 44 mm. 


Hab. 'Tasmania—Hobart. 


Larger and more elongate than EH. ovalis; the an- 
tenn longer, and with the ninth joint more elongate ; 
the elytra with numerous scattered coarser punctures on 
the disc, the punctures showing a tendency to form 
strie. One specimen. 


EINNEB@OPSIS, n. gen. 


Last joint of the maxillary palpi moderately stout, oblong- 
ovate, obliquely truncate at the tip ; eyes moderately large, almost 
entire, rather coarsely faceted ; epistoma short, separated from the 
front by an impressed line ; labrum moderately prominent ; an- 
tenn about reaching the base of the prothorax, joint 1 stout, 2 


* Cf, Champ. Biol. Centr.-Am., Col. iv., 1, pp. 539, 540 (1893). 


376 Mr. G. C. Champion on the Tenebrionidex 


short, slender, 3 nearly twice as long as 2, slender, 4 almost as 
broad as long, 5—11 perfoliate, very strongly transverse, widening 
outwardly, 5 twice as broad as 4, 11 abruptly rounded at the tip ; 
prothorax strongly transverse, with broad median lobe at the base, 
the base deeply sinuate on either side; scutellum strongly trans- 
verse ; elytra closely embracing the base of the prothorax, and at 
the sides forming almost a continuous outhne with it ; prosternum 
strongly horizontally produced, widened behind the coxe, and 
deeply excavate beneath for the reception of the mesosternum, the 
latter convexly raised in the middle in front, and obliquely grooved 
on either side; epipleure extending to the apex of the elytra, 
moderately wide in their basal third, narrower beyond ; middle coxal 
cavities widely open externally, the trochantin large and prominent; 
hind cox transverse, narrowly separated, the intercoxal process 
narrow, triangular; tibis slightly widened and compressed, obliquely 
truncate at the apex, subequal, the spurs short ; tarsi slender, thickly 
clothed with short silky hairs beneath, the anterior pair feebly 
dilated in the male, the first joint of the hind pair a little longer 
than the following two joints united; body convex, elliptic, 
winged. 

This genus is proposed for a single species from 
Tasmania. Hnnebeopsis has entirely the facies and sculp- 
ture of Hnnebeus, Wat., but differs from it in the per- 
foliate, strongly transverse outer joints of the antenne. 
The prosternum is similarly shaped in both. Hnnebwopsis 
connects Hnnebeus with the more typical genera of 
Diaperine. 


Ennebeeopsis pruinosus, sp.n. (Plate VIL, figs. 3, 3a.) 

Pitchy-black, shining, the entire upper surface very densely, 
exceedingly minutely punctate, clothed with fine pruinose pubes- 
cence; antenne and legs rufo-testaceous. Prothorax strongly 
transverse, the base deeply sinuate on either side of the median 
lobe, and with distinct fovez, the sides converging from the base ; 
elytra more than three times as long as the prothorax, with in- 
distinct rows of coarser punctures on the disc ; beneath varying in 
colour from piceous to testaceous, very densely, minutely punctate, 
finely pubescent ; fifth ventral segment triangularly emarginate at 
the apex, and the anterior tarsi feebly dilated, in the male. Length 
3—3} mm. (¢ .) 

Hab. Tasmania—Hobart. 


Feur examples; found under the loose bark of a 
Hucalyptus stump. 


collected in Australia and Tasmania. © ate 


Dipsaconia. 
Dipsaconia, Pascoe, Journ. Ent., i., p. 123 (1860). 


Dipsaconia australis. 


Eindophleus australis, Hope, Trans. Ent. Soc. Lond., 
1845, p. 108. 

Dipsaconia bakewellit, Pascoe, Journ. Ent., i., p. 124, 

tity Lio. 

Hab. Tasmania—Hobart and Launceston. 

In none of the specimens before me are the blackish 
markings of the elytra symmetrical. I have examined 
the type of Hndophleus australis, Hope, from Adelaide, 
and also that of D. bakewelli, Pascoe. D. pyritosa, 
Pascoe, is a closely allied form, but has much shorter 
hairs on the prothorax and elytra. Under bark of 
Eucalyptus (Walker). 


ULODEs. 


Ulodes, Hrichson, in Wiegmann’s Archiv, 1842, 1], 
p. 180. 


Ulodes verrucosus. 


Ulodes verrucosus, Hr., loc. cit., p. 181, t. 5, figs. 1, a, b. 
Endophleus varticornis, Hope, Proc. Ent. Soc. Lond., 
1842, p. 78; Trans. Ent. Soc. Lond., 1845, p. 108. 

Hab. Tasmania—Hobart and Launceston. 
Many specimens, chiefly found under the dry loose 
bark of “sassafras” trees (Atherosperma moschatum). 


Erichson’s insect was from Tasmania, that of Hope 
from Adelaide. 


GANYME. 


Ganyme, Pascoe, Ann. and Mag. Nat. Hist. (4), ui., 

p-. 32 (1869). 
Ganyme sapphira. 

Boletophagus sapphira, Newm., Entom., i; p» LOA: 

Ganyme sapphira, Pascoe, Ann. and Mag. Nat. Hist. 
(4), i, p. 33. 

Hab. Tasmania—Hobart. 

"TRANS. ENT. soc. LOND. 1894,—PaRT II. (JUNE.) 2B 


378 Mr. G. C. Champion on the Tenebrionide 


One specimen, differing from the type, which is con- 
tained in the National collection, in having the elytral 
pubescence almost entirely black, with the exception of 
two small patches on each elytron. G. howitti, Pasc., 
is probably only a small variety of the same species with 
the pubescence of the prothorax and elytra almost 
entirely reddish-ochreous. 


CaaNTHUS, n. gen. 


Mentum strongly transverse ; last joint of the maxillary palpi 
stout, oval, that of the labial pair slender ; mandibles bifid at the 
tip ; gula feebly grooved behind the suture ; epistoma truncate in 
front ; eyes coarsely granulated, small, entire, projecting laterally 
beyond the antennary orbits, the latter not prominent ; antennze 
not reaching the base of the prothorax, joints 1 and 2 stout, longer 
than broad, 3--9 slender, 3 considerably longer than 4, 4—9 very 
short, 7—9 strongly transverse, 10 and 11 forming a very stout club, 
10 transverse, 11 about as broad as long, obliquely truncate at the 
tip ; prothorax longitudinally convex, strongly compressed towards 
the sides, and with the apex broadly produced in the middle over 
the head* (partly hiding it when viewed from above), the base 
with broad median lobe ; scutellum not visible ; elytra about twice 
as long as the prothorax; prosternum very narrow, declivous; hind 
coxee widely separated, the intercoxal process broad, truncate in 
front ; epipleurze broad at the base, gradually narrowing to the 
apex; ventral sutures deep; legs short, without distinct tibial 
spurs, the tibiz convex on their outer edge ; tarsi simple, clothed 
with silky hairs beneath ; the basal joint of the hind tarsi scarcely 
equalling joints 2 and 3 united, the three together not longer than 
4; body elongate, convex, thickly covered with brownish scales, 
the upper surface with coarse scattered granular elevations, 


This genus is proposed for a minute insect from 
Tasmania. ‘The general shape is suggestive of that of 
the American genus Calymmus, of the Bolitophagides, in 
which group it must be placed. The stout 2-jointed club 
of the antenna, the gibbous, anteriorly produced, com- 


pressed thorax, and the very small, unemarginate eyes 
are its chief characters. 


Caanthus gibbicollis, sp. n. (Plate VIIL, figs. 7, 7a, b.) 


Elongate-oval, narrow, moderately convex, pitchy-black, the head 
ferruginons in front; densely covered with brownish scales, the pro- 


* Our artist has omitted to show this in the figure. 


collected in Australia and Tasmania. 379 


thorax and elytra with numerous coarse, granular elevations, those 
on the elytra serially arranged, these elevations bearing long, erect, 
fine bristly hairs ; antenne and legs rufo-testaceous. Prothorax in 
the middle about as long as broad, the sides rounded and coarsely 
crenate, the anterior angles produced and prominent, the base 
rather deeply sinuate on either side of the broad median lobe, the 
hind angles obtuse, the broad anterior prolongation truncate in 
front (when viewed from above) and extending forwards nearly 
as far as the front of the head; elytra not wider than the prothorax, 
subparallel to beyond the middle, with the shoulders rounded, the 
granular elevations closely placed and forming about six irregular 
rows on each elytron; beneath, when denuded of scales, ferruginous, 
shining, the flanks of the prosternum and the middle of the meta- 
sternum with coarse, scattered granular elevations. Length 2 
mm. 


Hab. Tasmania—Launceston and Hobart. 


Two specimens, found under bark. The upper surface 
is so densely covered with adherent, earthy-looking scales 
that the granular elevations are alone visible, the rest of 
the sculpture being completely hidden. The hairs on the 
thorax and elytra are perfectly erect. 


‘TRIBOLIUM. 
Triboliwum, Macleay, Annulosa J av., 1825, p. 47. 


This cosmopolitan genus is not included in Masters’s 
Catalogue. 


Tribolium ferrugineum, 
Trogosita ferruginea, Fabr., Spec. Ins.,i., p. 324. 
Hab. Tasmania—Hobart. 


ALPHITOBIUS. 


Alphitobius, Stephens, Il. Brit. Ent., v., p. 11 (1882). 
Alphitobwus piceus. 
Helops piceus, Oliv. Hnt., iii., No. 58, p. 17, t. 2, 
figs. 13, a, b. 
Alphitobius piceus, Muls., Col. Fr., Latig., p. 237. 
Hab. N. W. Australia—Adelaide River. 
One specimen of this cosmopolitan insect has been sent 


by Mr. Walker to the British Museum. The species is 
not included in Masters’s Catalogue. 


380 Mr. G. C. Champion on the Tenebrionide 


Toxicum. 


Toxicum, Latreille, Gen. Crust. et Ins., ii., p. 167. 


Toxicum punctipenne. 


Toxicum punctipenne, Pascoe, Journ. Ent., ii., p. 454. 

Toxicum addendum, Blackb., Proc. Linn. Soc. N. 8. W., 
i. (2), p. 1481. 

Hab. N. W. Australia—Port Darwin, Adelaide River. 


Many specimens of both sexes. In the male of this 
insect the anterior femora are subangularly dilated on the 
inner side towards the base; of this no mention is made 
by Mr. Blackburn. I have examined the types of 7. 
punctipenne, Pasc.; the length given 24 millim.” is 
evidently a misprint for “54 milim.” Mr. Blackburn’s 
specimens of ‘1’. addendwm were from the northern 
territory of South Australia. Found under logs and 
loose bark (Walker). 


PARATOXICUM, n. gen. 


Mentum nearly as broad as long, widest in front and gradually 
narrowing behind ; last joint of the maxillary palpi broad ovate, 
short, very obliquely truncate at the tip, that of the labial pair 
narrow, ovate ; inner lobe of the maxille armed with a curved 
claw at the tip; labrum not prominent; mandibles bifid at the 
tip ; head unarmed, short, almost trapezoidal, sunk into the pro- 
thorax up to the eyes, the antennary orbits a little swollen, 
extending completely across the eyes behind and obliquely con- 
verging in front, the epistoma broad, truncate in front, and limited 
behind by a rather deep groove ; eyes small, oblique, moderately 
coarsely granulated, the upper and lower portions equal ; antennz 
about reaching the base of the prothorax, similarly sculptured 
throughout, stout, gradually widening outwardly, joints 1 and 2 
short, 3 about twice as long as 2 and considerably longer than 4, 
4—11 decreasing in length and increasing in width, 8—11 strongly 
transverse, 11 bluntly rounded at the tip ; prothorax transverse, 
bisinuate at the base and apex, the apex broadly, feebly lobed in 
the middle ; scutellum short, transversely triangular ; elytra nearly 
three times as long as, and at the sides forming a continuous 
outline with, the prothorax, parallel for two-thirds of their 
length, sharply margined laterally ; prosternum narrow, produced 
behind, the mesosternum not excavate for its reception; inter- 


collected in Australia and Tasmania. 381 


mediate coxal cavities open externally, the trochantin prominent ; 
intercoxal process of the abdomen subtriangular ; elytral epipleure 
entire, broad, equal in width from opposite the metathoracic epimera 
to near the apex, widened towards the base ; legs rather short ; 
anterior tibiz flattened and subtriangularly dilated, with rather 
long spurs, the four hinder tibiz with very short spurs; tarsi 
simple, sparsely clothed with long hairs beneath ; posterior tarsi 
with the basal joint short, joints 1—3 united very little longer than 
the apical one; body elongate, parallel, winged. 


This genus is proposed for a Tasmanian insect allied 
to Toxicwm (and Anthracias), from which it chiefly 
differs in having the head unarmed, the antennez 
gradually widened to the apex (their three or four outer 
joimts not forming a club nor densely punctured and 
pubescent), and the epipleurse broad and entire. Epi- 
towicum, I". Bates, is also an allied genus. ‘The head is 
not swollen on either side near the eyes as in the females 
of Toxicum, nor cornute. The antennal joints are shining 
and similarly sculptured throughout. The sex of the two 
specimens received has not been ascertained. 


Paratoxicum iridescens, n. sp. (Plate VIIL., figs. 8, 8a—c.) 


Elongate, narrow, parallel ; the head, oral organs, antenne, and 
legs ferruginous, the prothorax and elytra dull black, the prothorax 
in one specimen indeterminately ferruginous towards the sides and 
apex, the entire upper surface iridescent. Head densely, very 
minutely punctate, the epistoma a little smoother; prothorax about 
one-third broader than long, rather convex, strongly bisinuate at 
the base and feebly so at the apex, the sides parallel from about the 
middle to the base and rounded in front, the hind angles acutely 
rectangular, the anterior angles rather sharp, the base very finely 
margined and obsoletely foveate on either side, the entire surface 
densely, minutely punctate ; elytra a little flattened on the disc, 
with regular rows of fine punctures, the interstices flat, smooth ; 
beneath shining, obscure castaneous, closely and finely punctured 
along the middle, the sides of the metasternum (but not the 
episterna) with coarser punctures, the ventral segments 1—3 more 
sparsely punctured towards the sides, the flanks of the prothorax 
finely strigose. Length 52, breadth 2 mm. 


Hab. Tasmania—New Norfolk. 


‘Two specimens, found under bark of Hucalyptus. 


382 Mr. G. C. Champion on the Tenebrionidx 


HELzts. 


Helzus, Latreille, Régne Anim., ed. 1, ii., p. 301 
(1817). 


Helzxus perforatus. 


Helxus perforatus, Latr., Régne Anim., ed. 2, u1., p. 
32, t. 3, fig. 6 ; de Bréme, Essai Monogr. Cossyph., 
i. p. 55, t. 6, fig. 2, and t. 1, figs. 1, 2; 5—3); 
Macl., Proc. Linn. Soc. N.S. W., ii. (2), p. 641. 
Hab. W. Australia—Fremantle. 


Originally described from Kangaroo Island. Under 
stones in sandy places (Walker). 


PTEROHELZAUS. 


Pterohelzus, de Bréme, Essai Monogr. Cossyph., 1., 
p- 27, t. 1, figs. 1—4, B (1842). 


Pteroheleus nigricormis, 1. sp. 
Pterohelxus nigricornis, F'. Bates, in litt. 


Broad oval, moderately convex, above and beneath, the legs 
and antennz black, the upper surface opaque. Head smooth, the 
epistoma separated on either side from the front by a fine oblique 
groove, the sides of the front broadly arcuately expanded, slightly 
raised, and projecting beyond the eyes ; the latter not prominent, 
very finely faceted, and separated by a space about equalling the 
width of one of the eyes as seen from above ; prothorax almost 
smooth, strongly transverse, fully three times as wide at the base 
as it is long in the centre, very deeply emarginate in front, deeply 
bisinuate behind, the sides arcuately and very rapidly converging 
from the base, the disc moderately convex and sometimes with 
indications of a fine impressed central line, the margins broadly 
explanate, the outer edge reflexed, the anterior angles rounded, 
the hind angles sharp and prolonged backwards; scutellum smooth; 
elytra a little wider than, and nearly four times the length of, the 
prothorax, rapidly arcuately narrowing from the middle, obliquely 
truncate on either side at the base, and with rather obtuse humeri, 
the disc moderately convex, somewhat abruptly declivous beyond 
the middle; each elytron with seven or eight equidistant coste, 
which become altogether obsolete on the apical declivity, the 
interspaces each with a double row of very fine, lightly impressed, 
punctures not extending to the apex; the margins broadly ex- 


collected in Australia and Tasmania. 383 


planate, the outer limb reflexed ; beneath slightly shining, almost 
smooth, the ventral segments 1—3 feebly longitudinally wrinkled ; 
prosternum produced behind and received by the rather deeply 
excavate mesosternum. Length 173—183, breadth 123—13}3 mm. 

Hab. N.W. Australia—Port Darwin and Adelaide River. 

Three specimens, two of which have been sent by Mr. 
Walker to the British Museum; also contained in Mr. F. 
Bates’s collection, where it is labelled with the name I 
have adopted. less elongate and more oval than P. 
walkeri, de Bréme, P. piceus, Kirby, and P. cornutus, 
Macl., the elytra smooth at the apex, the legs and 
antenne entirely black. Two of Mr. Walker’s specimens 
were dead and mutilated when found. 


Pterohelzus reichet. 


Pteroheleus vreichei, de Bréme, Hssai Monogr. 
Cossyph., i., p. 35, t. 2, fig. 2; Macl., Proc. Linn. 
Soc. N. 8. W. (2), i1., p. 531. 
Hab. Tasmania—Hobart. 
Several specimens, agreeing well with de Bréme’s 
figure. Found under loose bark of Eucalyptus. 
Pterohelzus parallelus. 


Pteroheleus parallelus, de Bréme, Essai Monogr. 
Cossyph., i., p. 33, t. 2, fig. 7; Macl., Proc. Linn. 
Soc. N.S. W. (2), i, p. 542. 

Hab. W. Australia—Fremantle. 


One specimen of this species has been sent by Mr. 
Walker to the British Museum. The type was obtained 
at Swan River. 

Pterohelxus bullatus. 


Pterohelxus bullatus, Pascoe, Journ. Ent., ., p. 462; 
Macl., Proc. Linn. Soc. N. 8. W. (2), u1., p. 536. 


Hab. W. Australia—Fremantle. 
One specimen. 
Pterohelxus speltoides. 
Pteroheleus peltoides, Macl., Proc. Linn. Soc. N.8. W. 
(2), i1., p. 546. 
Hab. S. Australia—Port Adelaide. 
One specimen. 


384 Mr. G.C. Champion on the Tenebrionide 


Pterohelxus peltatus. 
Cilibe peltata, Hr. in Wiegmann’s Archiv, 1842, p. 175. 
Pieroheleus wpeltatus, de Bréme, Essai Monogr. 
Gossyph.;.1.,.p. 34, t. 2, fig. 1; Macl§ Proc. 
Luinn. Soc. N.S. W.. (2), m5 p. 540. 
Hab. 'Tasmania—lLaunceston and Hobart. 


Originally described from Tasmania. Found under 
loose bark of Hucalyptus. 


SYMPETES. 
Sympetes, Pascoe, Journ. Ent., ii., p. 464 (1866). 
Sympetes tricostellus. 


Encephalus iricostellus, White, Voy. Capt. Grey, App., 
p. 464 (1841) ; de Bréme, Essai Monogr. Cossyph., 
lpg 0 Ody, ts 0, fig.6. 

Sympetes tricostellus, Macl., Proc. Linn. Soc. N. 8. W., 
u. (2), p. 652. 

Hab. W. Australia—Albany. 


A specimen of this species in the Hope Collection. at 
Oxford bears the MS. name of latipennis, Hope, in _ his 
own handwriting. Sandy places, under stones (Walker). 


Sympetes patelliformis. 


Saragus patelliformis, Pascoe, Ann. and Mag. Nat. Hist. 
(4), v., p. 100. 


Hab. W. Australia—Fremantle. 


One example, agreeing pretty closely with Pascoe’s 
type, but differmg from it in having the prothorax more 
densely punctured and with sharper anterior angles. A 
specimen of this species is contained in the Hope 
Collection at Oxford, with the name swbrugosus, Hope, 
attached, but it does not agree with the description or 
figure of S. subrugosus, de Bréme; and there is also 
another in Mr. F. Bates’s Collection, from Clampion Bay. 


Sympetes duboulan. 


Saragus duboulai, Pascoe, Journ. Ent., 1., p. 466; 
Macl., Proc. Linn. Soc. N. S. W., 11. (2), p. 670. 


Hab. W. Australia—H. Wallaby Island in the Hout- 
mann’s Abrolhos Group, and Fremantle. 


collected in Australia and Tasmania. 385 


The type of this species was from Champion Bay. S. 
duboulait seems best placed in Sympetes, the prosternum 
being declivous behind and not received by the meso- 
sternum. At roots of bent grass, on sandhills (Walker). 


SARAGUS. 


Saragus, Hrichson, in Wiegmann’s Archiv, 1842, 1, 
pale 


Suragus levicollis. 


Silpha levicollis, Fabr., Syst. Ent., p. 73; Oliv., 
Hint.) in, 015 yp. 02) 't. 2. fea 

Saragus levicollis, Er., in Wiegmann’s Archiv, 1842, 
ft, p. 172, +.’ 4, figs. 7, a0; de Breme, ssa 
Monogr. Cossyph., i., p. 44, t. 3, fig. 1; Hope, 
Trans. Ent. Soc. Lond., 1848, p. 56, t. 7, fig. 5; 
Macl., Proc. Linn. Soc. N.S. W., ii. (2), p. 657. 

Cilibe costatus, Sol., Studi Hnt., p. 355, t. 18, figs. 
10-13 (1848). 


Hab. Tasmania—Hobart and Launceston. 


Sandy places, under stones and at roots of herbage 
(Walker). 


Saragus bicarinatus, n. sp. (Plate VILI., fig. 10.) 


Moderately elongate, broad, convex, subparallel, very obtuse 
behind, brownish-black, opaque. Head deeply sunk into the pro- 
thorax (the eyes only just visible from above), thickly and finely 
punctate, the epistoma limited at the sides and posteriorly by a 
shallow groove ; prothorax strongly transverse, deeply emarginate 
in front, the sides rapidly converging from the base, a little 
rounded anteriorly, the base slightly emarginate in the middle, 
rounded to the outer limit of the disc, and then very obliquely 
truncate to the acute, deflexed, posteriorly-produced hind angles, 
the anterior angles rather obtuse, the disc convex, obsoletely 
canaliculate in the middle, and very finely, closely punctate, with 
the interspaces (viewed under a strong lens) densely minutely 
punctured, the margins broadly, horizontally explanate, shagreened 
and minutely granulate, the exterior edge not reflexed ; elytra 
about two and a half times longer than the prothorax, and of the 
same width at the base, parallel to about the middle, very obtuse 
behind, strongly trisinuate at the base, with subacute, deflexed, 


386 Mr. G. C. Champion on the Tenebrionidx 


outwardly directed humeri, each elytron with three longitudinal 
equidistant coste extending to considerably beyond the middle, the 
inner one stout, smooth, and shining, and strongly raised, the other 
two faint and slightly crenulate, the external one curving inwards 
at the base, the space between the first costa and the suture (which 
is not raised and slightly shining) quite flat, and between it and 
the lateral margin obliquely declivous, the interspaces shagreened 
and very minutely granulate, and with rows of subobsolete punc- 
tures, a row of coarser impressions midway between the outer 
costa and the margin, the margins moderately horizontally ex- 
planate, the exterior edge not thickened ; the legs and under 
surface fusco-ferruginous ; the prosternum granulate, longitudin- 
ally wrinkled at the sides, the prosternal process rugosely punctured, 
the latter subhorizontally produced, and received by the deeply 
excavate V-shaped mesosternum; the rest of the under surface 
thickly punctured and wrinkled, the pleura smoother ; anterior 
tibiz with a short fine tooth at the outer apical angle. Length 164, 
breadth 10 mm. 


Hab. N.W. Australia—Roebuck Bay. 


One specimen. Very near S. confirmatus, Pasc., from 
W. Australia, but much larger, broader, and more robust, 
the legs much stouter; the innermost costa on the 
elytra stouter, more raised, and shining, the other costz 
faint (not being any more prominent than they are in 
S. confirmatus); the prothorax not wider than the elytra 
at the base. Another closely allied, perhaps undescribed, 
species, from Nicol Bay, is contained in Mr. F. Bates’s 
collection. S. bicarinatus is apparently not described in 
MacLeay’s monograph of the genus. 


Saragus intricatus, n. sp. 


Oblong-oval, very convex, black, subopaque. Head sparsely 
and very finely punctate, the interocular space more coarsely and 
more closely punctured ; prothorax at the base more than twice as 
broad as long, broadly and very deeply emarginate in front, the 
sides rapidly and arcuately converging from the base, the base 
feebly emarginate in the middle, broadly rounded to the outer 
limit of the disc, and then very obliquely truncate to the sharp 
posteriorly-produced hind angles, the disc transversely convex, 
shallowly longitudinally grooved in the middle behind, and thickly, 
very finely punctate, the margins strongly horizontally explanate, 
and finely shagreened, the outer edge not reflexed, the anterior 
angles rounded ; scutellum smooth; elytra slightly wider than, 


collected in Australia and Tasmania. 387 


and three times as long as, the prothorax, parallel to about one- 
third from the base, arcuately and rapidly narrowing posteriorly, 
conjointly rounded at the apex, and with rather obtuse humeri, 
each elytron with three sinuous subequidistant feeble carina (the 
first straight and more sharply raised anteriorly), and with a still 
more feeble carina midway between each of these, all connected 
here and there by transverse or oblique rami, the interspaces with 
interrupted series of coarse, exceedingly shallow punctures, the 
space between the first costa and the suture (which is not raised) 
flat, the sides abruptly declivous, the margins very slightly dilated 
at the base; beneath very sparsely minutely punctate, and also 
longitudinally wrinkled, and clothed with a few scattered hairs ; 
the prosternal process thickly punctured, strongly horizontally 
produced, and received by the deeply excavate V-shaped meso- 
sternum ; the femora very sparsely, the tibie closely and roughly, 
punctured, the anterior tibe feebly toothed at the outer apical 
angle. Length 143, breadth 9} mm. 

Hab. N.W. Australia—Adelaide River. 

Three examples, two of which are contained in the 
British Museum. Apparently distinct from al! the 
numerous described species of the genus. In the sculp- 
ture of the elytra it approaches S. reticulatus, Haag, 
from Endeavour River. 


Saragus infelix. 
Saragus infeliz, Pasc., Journ. Ent., 11., p. 466. 
Hab. Tasmania—Launceston. 


One specimen of this species has been sent by Mr. 
Walker to the British Museum. The type was from 
Tasmania, 


Saragus sp. (?). 
Hab. N.W. Australia—Montalivet I. 


One mutilated specimen (without head), perhaps be- 
longing to an undescribed species, sent by Mr. Walker 
to the British Museum. 


Saragus brunnipes. 
Celibe brunnipes, Boisd., Voyage de |’Astrolabe, Ent., 
i., p. 264. 
Cilibe brunnipes, de Bréme, Essai Monogr. Cossyph., 
I, peor, t. 8, fig. 4, 


388 MraG:C, Champion on the Tenebrionidx 


Saragus brunnipes, Macl., Proc. Linn. Soc. N.S. W. 
(2), u.,"p. 669. 
Hab. W. Australia—Cape Leeuwin. 


Three examples, probably belonging to this species. 
They closely resemble S. macleayi, Blackb., but are flatter, 
and have the expanded margins of the thorax and elytra 
ferruginous, and the humeri more angular. 


NycrTozoIuus. 


Nyctozoilus, Guérin, Voy. Coquille, Ent. 1, p. 92 
(1830). 
Sphenogenius, Solier, Studi Hnt., p. 35, 


Nyctozoilus sexcostatus,n. sp. (Plate VIII., fig. 9.) 


Oblong ovate, convex, rather broad, dull black, very sparsely 
clothed with exceedingly short, fine, appressed, yellowish-brown 
hairs. Head broad, slightly depressed in front, thickly and finely 
punctate, the epistoma feebly arcuate-emarginate at the apex, the 
antennary orbits prominent and extending more than half-way 
across the eyes; antenne piceous, ferruginous at the tip, not 
reaching the base of the prothorax, thickening a little outwardly, 
joint 3 elongate, about two and a half times as long as 4, 4—7 
decreasing slightly in length, 8 as broad as long, 9 and 10 strongly 
transverse, 11 twice as long as 10, rounded at the tip ; prothorax 
transverse, widest at the middle, a little wider at the base than at 
the apex, moderately convex, with the sides flattened and hori- 
zontally explanate, the apex broadly and deeply emarginate, the 
base (viewed from behind) very broadly and feebly arcuate- 
emarginate, distinctly margined, the sides strongly rounded at the 
middle, obliquely converging anteriorly, and constricted and deeply 
sinuate before the base, the anterior angles sharp and very pro- 
minent, the hind angles obliquely produced behind and overlapping 
the elytra, the lateral margins, and the apical margin (except in the 
middle), feebly reflexed and slightly crenulate, the entire surface 
very minutely shagreened and with shallow, scattered, fine irregular 
punctures (each bearing a short hair), which are more crowded 
towards the middle of the disc, the latter with an indistinct 
median groove and a broad shallow depression on either side of it 
a little behind the middle ; elytra very much wider than, and about 
twice as long as, the prothorax, a little rounded at the sides, 
strongly so at the base, each with three fine, sharp, slightly 
sinuous, smooth carine (the first and third almost confluent behind 


collected in Australia and Tasmania. 389 


and enclosing the shorter median one, the first curving a little 
outwards, and the second and third curving a little inwards, at the 
base), the suture smooth and similarly raised, the interspaces very 
minutely shagreened, feebly transversely wrinkled, and with 
scattered fine setiferous punctures, which become coarser towards 
the sides; beneath blackish-brown, dull, more thickly pubescent 
(except at the sides), very minutely shagreened, and somewhat 
closely impressed with fine, muricate punctures, the propleurz with 
widely scattered, simple, shallow punctures, each bearing a hair ; 
prosternum very broad, horizontal, strongly produced, rounded at 
the tip ; mesosternum broad, subangularly raised on either side in 
front; metasternum very short; intercoxal process exceedingly 
broad, subparallel, rounded in front ; epipleures very broad at the 
base, gradually narrowing behind, and extending to the apex ; legs 
pitchy-brown, rather slender, closely punctured. Length 14, 
breadth 7} mm. 


Hab. N. W. Australia—Adelaide River. 


I am unable to identify this very distinct species with 
any of those described by F. Bates, Macleay, or Haag; 
it agrees with the type, NV. obesus, Guér., in its structural 
characters. One specimen, found under a stone on a dry 


bank (Walker). 


HYpavLax. 


Hypaulax, F. Bates, Trans. Ent. Soc. Lond., 1868, 
p. 259, Nat. Hist. (4), xii., p. 16. 


Hypaulax ampliata. 
Hypaulax ampliata, F. Bates, Am. and Mag. Nat. Hist. 
(A), xiii., p. 19. 
Hab. N. W. Australia—Roebuck Bay. 


Numerous examples, found under the dry bark of 
stumps, etc. Originally recorded from W. Australia, 


Hypaulax vridescens. 
Hypaulax iridescens, Blackb., Proc. Linn. Soc. N. 8. W. 
(2), 11, p. 1433. 
? Platynotus insularis, Hope, Proc. Ent. Soc. Lond., 
1842, p. 77; Trans. Ent. Soc. Lond., 1845, p. 107. 


Hab. N.W. Australia—Port Darwin, Adelaide River. 


390 Mr. G. C. Champion on the Tenebrionidzx 


Many specimens from these localities, varying in 
length from 94—17 mm., agree very nearly with the 
Rev. T. Blackburn’s description of H. iridescens, the 
original examples of which were obtained in the northern 
territory of 8. Australia. The species is distinct from all 
those described by Mr. F. Bates. It is not improbable 
that this is the Platynotus insularis, Hope,* from Melville 
Island and Port Essington. Iam unable to find the type 
of Hope’s species in the Oxford museum. ‘The hairy 
mentum is a mark of the male sex. Under loose bark 
and also under logs lying in dry places (Walker). 

MENEPHILUS. 

Menephilus, Mulsant, Col. Fr., Latig., p. 291 (1854). 

The Australian species of this genus are found under 
dry bark and in dry rotten wood, according to Mr. 


Walker. 
Menephilus longipennis. 


Tenebrio longipennis, Hope, Proc. Ent. Soc. Lond., 
1842, p. 79; Trans. Ent. Soc. Lond., 1845, p. 110. 
Hab. Tasmania—Franklin, New Norfolk, George’s 
Bay. 
Originally recorded from Adelaide. I have examined 
Hope’s type in the Oxford Museum. 


Menephilus corvinus. 
Tenebrio corvinus, Er. in Wiegmann’s Archiv, 1842, 
eS nalioe 
Tenebrio cyanipennis, Hope, Proc. Ent. Soc. Lond., 
1842, p. 79; Trans. Ent. Soc. Lond., 1845, p. 111. 
Hab. 'Tasmania—Launceston. 
It is uncertain which name has priority, both having 
been published in 1842. richson’s type was from 
Tasmania, that of Hope from Adelaide. 


Menephilus colydiordes. 
Tenebrio colydioides, Er. in Wiegmann’s Archiv, 1842, 
Lp. 7b: 
Hab. Tasmania—Hobart, Launceston, and George’s 
Bay. 


Apparently a common insect in ‘T'asmania. 


* Incorrectly referred to Cestrinus by Pascoe and Gemminger 
& Harold. 


collected in Australia and Tasmania. 391 


Menephilus ruficornis, n. sp. 


3. Moderately elongate, convex, shining, piceous or obscure 
castaneous, the head more or less ferruginous in front, the elytra 
violaceous, zneous with a violaceous lustre, or eneous; the antennez 
rufo-testaceous ; the under surface pitchy-brown or castaneous ; 
the legs piceous with the knees and tarsi castaneous, or entirely 
castaneous. Head short, somewhat deeply sunk into the prothorax, 
coarsely, closely punctured between and behind the eyes, the 
punctures oblong in shape, the anterior portion much more finely 
punctate, the epistoma limited at the sides and behind by a rather 
deep groove ; the eyes moderately large and prominent, coarsely 
faceted, the orbits impinging on them in front; antennz moderately 
stout, thickening outwardly, scarcely reaching the base of the 
prothorax, joint 6 wider than 5, 7—11 perfoliate, much wider than 6, 
8—10 strongly transverse, 11 twice as long as 10, bluntly rounded at 
the tip ; prothorax convex, broader than long, very little wider at 
the base than at the apex, the sides sinuate behind and rounded 
anteriorly, the base strongly bisinuate, the hind angles rectangular, 
the anterior angles obtuse and declivous, the surface coarsely, closely 
punctured, the apex smoother ; scutellum subtriangular, about as- 
long as broad, faintly punctured ; elytra about three times as long 
as, and considerably wider than, the prothorax, subparallel in their 
basal half, with rows of coarse subapproximate punctures placed 
upon rather shallow striz, the punctures becoming finer towards 
the suture and coarser towards the sides, the interstices feebly 
convex, sparsely, very distinctly punctate ; beneath shining, the 
flanks of the prothorax with coarse, scattered punctures, the ventral 
segments rather coarsely and moderately closely, the metasternum 
very sparsely and finely, punctate; prosternum slightly declivous 
and strongly produced behind the anterior coxze, grooved on either 
side, the apex raised; epipleure extending as far as the last 
ventral suture, rather broad; tibize slightly curved, pubescent on the 
inner side towards the tip, the two hinder pairs feebly sinuous 
within, the anterior pair with their outer apical angles obtuse. 
Length 7—73, breadth 23 mm. 

Hab. Tasmania—Hobart and George’s Bay. 


Three examples, apparently all males. Allied to M. 
corvinus, Hr., and M. cxrulescens, Haag, but much 
smaller than either of these species, with the head, thorax, 
and under-surface much more coarsely punctured, the 
frontal suture deeper, etc. The elytra vary in colour 
from violaceous to eneous. Found under bark (Walker). 


392 Mr, G, C, Champion on the Tenebrionidee 


MENERISTES, 


Meneristes, Pascoe, Aun. and Mag. Nat. Hist. (4), i., 
p. 150 (1869). 
Asiris, Motschulsky, Bull, Mosc., xlv., 2, p. 24 (1872). 


Pascoe states that this genus only differs from Mene- 
philus in the following characters—‘“‘tibiz calcarate, 
femora incrassata.” He has overlooked the fact that the 
tibiz in his typical species, M. laticollis, are very dis- 
tinctly channelled on their outer edge; M. servulus, 
Pasc., has the tibize rounded on their outer edge, and it 
cannot be included in the same genus. 


Meneristes australis. 


abt io australis, Boisd., Voyage de l’ Astrolabe, Ent. 
, p. 204; Bless., Hore Ent. Ross. sy lj: Pv St 
ee istes laticollis, Pasc., Ann. and Mag. Nat. Hist. 
(8), avs p. loos tte fig. 2 (nec Boisd.). 
Asiris angulicollis, Motsch., Bull. Mosc., xlv., 2, p. 30. 


Hab. Tasmania—Hobart and Launceston. 
Under Hucalyptus bark, common (Walker). 


I am unable to follow Pascoe in regarding this species 
as the Baryscelis laticollis of Boisduval, though it may 
be the Tenebrio australis of that author. The Rev. T. 
Blackburn (Trans. R. Soc. 8. Austr., x., p. 282) unites 
Boisduval’s species under the one aa. Meneristes 
australis ; but to judge from the imperfect descriptions 
(Voy. de V Astrolabe, 11., pp. 2038, 254) they cannot 
possibly be synonymous. An insect sent me by Mr. 
Blackburn as the Tenebrio australis, Boisd., agrees 
with the type of M. intermedius, Pasc.; it differs from 
M. laticollis in its smaller size, and in having less acute 
hind angles to the prothorax. 


PROMETHIS. 


Promethis, Pascoe, Ann. and Mag. Nat. Hist. (4), 11., 
p. 148 (1869). 
Mederis, Motschulsky, Bull. Mosc., xlv., 2, p. 24 (1872). 


collected in Australia and Tasmania. 393 


Promethis angulata. 


Upis (Iphthinus) angulatus, Er. in Wiegmann’s Archiv, 
1842, 1, p.174. 
Promethis angulata, Pasc., loc. cit. 


Hab. Tasmania—Hobart and New Norfolk. 


Previously recorded from Tasmania and Victoria. 
Under Eucalyptus bark, generally in company with 
Meneristes (Walker). 


L&EPisPILus. 


Pachyccelia, Boisduval, Voyage de I’ Astrolabe, Ent. ii., 
p. 248 (1835) (nomen prevocc.). 
Lepispilus, Westwood, Arcana Ent., i., p. 44 (1841). 


Lepispilus sulcicollis. 


Pachycelia sulcicollis, Boisd., Voyage de |’ Astrolabe, 
Kint. u., p. 248. 

Helops sulcicollis, Boisd., loc. cit., p. 268, t. 7, fig. 5. 

Lepispilus sulcicollis, Westw., Arcana Ent., i., p. 44, 
t. 12, fig. 4; Blanch., in Dumont d’Urville’s 
Voyage au Pole Sud, iv., p. 172, t. 11, fig. 14. 

Hab. Tasmania—George’s Bay and Hobart. 


Originally described from Tasmania. ‘To judge from 
the type, L. stygianus, Pasc., appears to me to be nothing 
more than a worn (female) example of the same species. 
Found on foliage and by sweeping low plants (Walker). 


Omo.irus. 
Omolipus, Pascoe, Journ. Ent., i., p. 127 (1860). 


Omolipus oblongus. 


Omolipus oblongus, F. Bates, Trans. Ent. Soc. Lond., 
1873, p. 379. 


Hab. W. Australia—Cossack. 


Found in plenty by Mr. Walker in sandy places on the 
seashore, at roots of bent grass. These specimens differ 
from the mainland type in their smaller size, more 
shining surface, and more obsoletely striate elytra; but 
I do not think they represent more than a local form of 
O. oblongus. 'The type is from Champion Bay. 


TRANS. ENT. SOC. LOND. 1894,—PaRT I. (JUNE.) 2¢ 


394 Mr. G. C. Champion on the Tenebrionde 


Omolipus cyanetpennis, n. sp. (Plate VIII., fig. 12, 3.) 


Moderately elongate, obovate, black, the elytra cyaneous or 
cyanco-violaceous, the entire upper surface subopaque. Head very 
minutely and sparsely punctured, the epistoma separated from the 
front by a fine impressed line, the eyes small ; antennz black, about 
reaching the base of the prothorax, thickening outwardly, the 
penultimate joints transverse, the apical one twice as long as the 
tenth ; prothorax broader than long, moderately convex, very finely 
margined at the sides, more distinctly so at the base, the sides 
rounded anteriorly, obliquely converging behind, and feebly sinuate 
just before the base, the hind angles distinct, the surface still more 
minutely and more sparsely punctured than that of the head (the 
punctures only visible under a strong lens) ; elytra about twice as 
long as the prothorax, and a little wider than it at the base, sharply 
margined at the sides and with the humeri rather prominent, 
widest about the middle and then rapidly narrowing to the apex, 
punctate-striate—the striz shallow, the punctures oblong in shape, 
not very coarse, and rather closely placed, and becoming finer 
towards the apex,—the interstices smooth and slightly convex ; be- 
neath dull pitchy-black, the two basal segments of the abdomen 
with a faint cyaneous lustre, the apical two or three segments 
slightly shining, the venter sparsely and very minutely punctured ; 
the prosternum canaliculate on either side, and declivous behind, 
the apex conically raised ; legs black, shining ; the anterior tarsi 
slightly dilated in the male. Length 54—84, breadth 2}—3} mm, 
(d 2). 

Hab. N. W. Australia—Baudin Island. 


This beautiful species was found in some numbers by 
Mr. Walker, under fragments of ironstone on the summit 
of the island, at an elevation of about 250 feet, in 
company with Metyche cxrulea. ‘The island upon which 
it occurred was named by the officers of H.M.S. “ Pen- 
guin,”’ during their surveying expedition to the coast of 
N. W. Australia, O. cyaneus, Pasc., is an allied form. 


Omolipus parvus. 


Omolipus parvus, F. Bates, Trans. Ent. Soc. Lond., 
1873, p. 379. 
Hab. W. Australia—Fremantle. 


A single male example, agreeing perfectly with the 
type; the latter is from Swan River, 


collected in Australia and Tasmania. B95 


DercraLMa, 


Decialma, Pascoe, Ann. and Mag. Nat. Hist. (4), iii., 
p. 291 (1869). 


Decialma erichsoni, n, sp. (Plate VIIL., fig. 11.) 


Decialma erichsoni, F. Bates, in litt. 


d. Moderately elongate, black, the upper surface with a cwruleous 
lustre, shining, glabrous. Head coarsely, rugosely punctured, the 
vertex with a small transverse impunctate space in the middle ; 
antenns pitchy-brown, slender, extending to a little beyond the 
base of the prothorax, joint 3 nearly as long as 4 and 5 united, 9 
and 10 a little longer than broad, obconic, 11 one-half longer than 
10, bluntly rounded at the tip; prothorax short, very strongly 
transverse, convex on the disc, broadly and deeply depressed 
towards the sides, with a large broad space between the middle and 
the base more deeply excavate, the lateral margins explanate and 
strongly upturned, the sides feebly rounded, slightly convergent at 
the base and more distinctly so in front, the apex very broadly 
truncate in the middle, the anterior angles rounded, strongly, very 
broadly produced, the base deeply bisinuate, the hind angles sub- 
rectangular, the surface thickly, moderately finely punctate, a 
median line on the dise and the lateral portion anteriorly almost 
smooth ; scutellum smooth ; elytra considerably broader than, and 
about four timesas long as, the prothorax, subparallel in their basal 
half, moderately convex, thickly, irregularly, and rather coarsely 
punctate (the punctures becoming much finer at the apex), and 
with indications of irregular faint raised lines; beneath very 
shining, the metasternum at the sides and the adjacent part of the 
epipleura with coarse scattered punctures, the flanks of the pro- 
thorax feebly wrinkled, the sides of the ventval segments longi- 
tudinally wrinkled and with a few scattered punctures, the median 
part closely and more finely punctured ; prosternum horizontal, 
very strongly, convexly produced behind, and received by the 
exceedingly deeply excavate mesosternum, the latter U-shaped, 
horizontal, with the strongly raised sides vertical in front ; legs 
pitchy-black, the tarsi paler ; anterior tarsi slender ; hind tibiz on 
the inner side subangularly widened at about one-fourth from the 
base and slightly sinuous. Length 10}, breadth 4% mm. 

Hab. 'asmania—Denison Gorge, near Launceston, 


Specimens of this insect are also contained in Mr. F. 
Bates’s collection, labelled with the name I have adopted, 


396 Mr. G. C. Champion on the Tenebrionidx 


One specimen, found under bark of Atherosperma mos- 
chatum. The genus Decialma, Pasc., is probably not 
distinct from Olisthena, Er. 


TITENA, 


Titena, Hrichson, in Wiegmann’s Archiv, 1842, 1, 
p. 178; F. Bates, Ann. and Mag. Nat. Hist. (4), 
xil., p. 102. 


Titsena columbina. 


Titena columbina, Er., in Wiegmann’s Archiv, 1842, 
1, p. 179, t. 4, figs. 9, a, 6; Blanch., in Dumont 
d’Urville’s Voyage au Pole Sud, iv., Ins., p. 178. 

Strongylium rugosum, Blanch., loc. cit., t. 12, fig. 2. 


Hab. 'Tasmania—Hobart, Launceston, and Franklin. 


Not uncommon, under bark of Acacia decurrens and A. 
dealbata, locally known as wattle-trees. 


Titena alcyonea. 


Titena alcyonea, Er., in Wiegm. Archiv, 1842, 1, 
p- 180. 
Hab. 'Tasmania—Hobart. 


One specimen ; also sent by Mr. Walker to the British 
Museum. Examples of this insect are separated from 7’. 
alcyonea, Er.,in Mr. F. Bates’s Collection and labelled 
with the MS. name of cyanea, Bates; Mr. Walker’s 
insect, however, accurately fits Hrichson’s description. 


Titena tasmanica,n. sp. (Plate VIII, fig. 13.) 


Titena tasmanica, F. Bates, in litt. 


Elongate, narrow, very convex, «neous or cupreo-zeneous, the 
head and prothorax usually of a greener tint, glabrous, the antennee 
and legs rufous or pitchy-red. Head coarsely, confluently punc- 
tured, the punctures on the epistoma finer and separate one from 
another ; prothorax very convex, as long as broad, widest at 
the middle, about equal in width at the base and apex, the apex 
truncate, the base rounded at the middle and feebly sinuate on 
either side, the sides a little rounded in front and slightly con- 
verging behind, the anterior angles deflexed and obtuse, the surface 
coarsely and densely punctured, the punctures here and there 
longitudinally confluent ; elytra rather more than three times ag 


collected in Australia and Tasmania. 397 


long as, and considerably wider than, the prothorax, a little 
narrowed in front, closely and very coarsely punctured, the 
punctures variolose, here and there transversely confluent, and 
arranged in irregular, closely packed, longitudinal series ; beneath 
pitchy-brown, very coarsely punctured, the ventral segments more 
finely punctured in the middle. Length 6—6}4, breadth 2} mm. 


Hab. Tasmania—Launceston and Hobart. 


Specimens of this insect are also contained in Mr. F. 
Bates’s Collection, where they are labelled with the name 
here adopted. Much smaller, shorter, and less parallel 
than 7. columbina or T. alcyonea, Hr., the upper and 
under surfaces glabrous. Found in rotten wood, old 
posts, etc. (Walker). 


Metytrra. 
Melytra, Pascoe, Ann. and Mag. Nat. Hist. (4), iii., 
p- 34 (1869). 
Melytra ovata. 
Melytra ovata, Pascoe, loc. cit., t. 10, figs. 1, la, b. 
Hab. 'Tasmania— Hobart. 


Hymma. 
Hymea, Pascoe, Ann. and Mag. Nat. Hist. (4), i1., 
p. 39 (1869). 
Hymexa succinifera. 
Hymea succinifera, Pascoe, loc. cit., p. 36, t. 10, figs. 
3, da, b. 
Hab. Tasmania—Launceston and Hobart. 


Under bark of <Atherosperma moschatum and Lu- 
calyptus coccifera, at elevations above 2,500 feet 


(Walker). 
KcrYcHeE. 
Ectyche, Pascoe, Ann. and Mag. Nat. Hist. (4), iv., 
p. 143 (1869). 


Eetyche cerulea,n. sp. (Plate VIII, fig. 15, d.) 


Ceeruleous or violaceous ; the head and thorax dull, the elytra 
shining and of a brighter metallic colour; the upper surface clothed 


398 Mr. G. C. Champion on the Tenebrionide. 


with very long, erect blackish hairs, those on the elytra serially 
arranged and arising from the interstitial punctures. Head im- 
pressed with oblong, rather coarse, scattered punctures, the 
epistoma separated from the front by an impressed line, the eyes 
strongly transverse; antennz pitchy-brown, hairy, extending a 
little beyond the base of the prothorax, thickening a little out- 
wardly, the joints perfoliate and submoniliform, 3—7 moderately 
elongate, 8—11 shorter, 11 stouter than 10; prothorax feebly 
transversely convex, strongly transverse, as broad as the elytra, 
widest at the middle, the sides greatly rounded, explanate, and 
abruptly notched immediately before the base, the entire surface 
with scattered, oblong, coarse setiferous punctures, between which 
are some very much finer and shallower non-setiferous impres- 
sions, the dise with traces of a smooth space down the middle ; 
elytra more than twice as long as the prothorax, ovate, parallel 
anteriorly, coarsely crenate-striate, the punctures becoming finer 
towards the apex, and not very closely placed, the interstices 
moderately convex, each with a row of scattered, fine, transverse, 
setiferous impressions, the base truncate, the humeri obtuse ; 
beneath pitchy-black, very shining, with widely scattered, rather 
coarse setiferous punctures ; legs piceous or brownish, hairy, with 
intermixed much longer erect hairs. 

¢ The anterior tibiz with the single spur at the inner apical angle 
longer and sharper, the anterior and intermediate pairs more 
curved ; the first ventral segment triangularly depressed in the 
middle behind. Length 7—8 mm. (¢ 9 ) 


Hab. N. W. Australia—Parry Harbour, Troughton I., 
Baudin I., Condillac I. 


Hasily known from the allied forms by its large size 
and the bluish colour of the upper surface. LH. scabri- 
pennis, I’. Bates, from Nicol Bay, is also a large species, 
but differs from it in many respects, apart from colour. 
Not rare, under large fragments of ironstone on the 
summits of Baudin and Condillac Islands (Walker). 


Hictyche erebea. 


ee erebea, Pasc., Ann. and Mag. Nat. Hist. (4), 
i, p. 144, t. 11, fies, 1; la, v: 


pe W. Australia—Fremantle. 


Originally described from the same locality. LH. sculp- 
turata, F. Bates, from Nicol Bay, approaches this species 
very closely. 


collected in Australia and Tasmania. 399 


MIcREcTYCHE. 


Micrectyche, ¥. Bates, Trans. Ent. Soc. Lond., 1873, 
p. 362. 


These insects are found at the roots of low plants on 
sandy beaches, according to Mr. Walker. 


Micrectyche intermedia. 


Micrectyche intermedia, F. Bates, loc. cit., p. 363. 


Hab. W. Australia—H. Wallaby Island in the Hout- 
mann’s Abrolhos Group. 


One specimen, agreeing perfectly with the type from 
Champion Bay. 


Micrectyche ferrugmea. 


Micreciyche ferruginea, F'. Bates, loc. cit., p. 364. 
Hab. W. Australia—Fremantle. 


Three specimens. The type is stated to be from Swan 
River. 


PHENNIS, 0. gen. 


Head short and broad, sunk into the prothorax up to the eyes, 
with the sides of the front obliquely converging and not at all 
prominent (not projecting over the points of insertion of the 
antennz) ; the epistoma very short, truncate at the apex, limited 
behind by a shallow groove ; labrum prominent, separated from 
the epistoma by a coriaceous space ; the eyes small, transverse, 
moderately prominent, rather finely faceted, feebly emarginate ; 
last joint of the maxillary palpi ovate ; antennz hairy, moderately 
slender, extending to a little beyond the base of the elytra, joints 
1—8 more or less ovate, 1 stout, 2 about as long as 4, 3 nearly as 
long as 4 and 5 united, 4—8 equal in thickness but gradually de- 
creasing in length, 9 nearly twice as long as, and very much wider 
than, 8, triangular, longer than broad, 10 triangular, as broad as 
long, 11 oval, narrower than 10, blunt at the tip ; prothorax trans- 
versely cordate, subtruncate in front and behind, coarsely, irregularly 
dentate at the sides, the latter slightly expanded; scutellum strongly 
transverse, transversely convex; elytra fully one-half broader than, 
and about three times the length of, the prothorax, truncate at the 
base, parallel in their basal half, each with ten rows of .coarse deep 
punctures ; anterior coxal cavities closed behind ; legs rather short, 


4.00 Mr. G. C. Champion on the Tenebrionidx 


hairy, moderately slender, the penultimate joint of all the tarsi 
small, simple, the tarsi clothed with long hairs beneath, the first 
joint of the hind pair not longer than the following two joints 
united, the tibial spurs short but distinct; the upper surface clothed 
with long, erect bristly hairs, with intermixed short, appressed, fine 
hairs, the latter forming well-defined patches on the elytra. 


This genus is proposed for a single species from 
Tasmania. It seems to be best placed near LHctyche. 
The clothing of the upper surface resembles that of the 
“ Amphidorides,” except that the finer hairs on the elytra 
form well-defined patches. The form of the antenne is 
peculiar—the ninth to the eleventh joints being stouter 
than those preceding, the ninth and tenth triangular, the 
ninth longer than the eighth or tenth. 


Phennis fasciculata, n. sp. (Plate VIII., fig. 14.) 


Oe Moderately elongate, rather convex, pitchy- brown, the front 
of the head and the anterior margin of the prothorax paler; the 
elytra with an oblong humeral patch and the suture indeterminately 
testaceous, the interstices also paler towards the base ; the antenns 
fusco-testaceous, with the three apical joints piceous ; the legs 
fusco-testaceous, with the femora darker towards the base; the 
upper surface sparsely clothed with long, erect, bristly, brownish 
hairs, which are serially arranged on the elytra, and with intermixed 
short, appressed, yellowish-cinereous hairs, the latter forming well- 
defined patches on the elytra ; the legs and antennz clothed with 
long, fine, bristly hairs. Head thickly, shallowly punctate ; pro- 
thorax transversely cordate, much narrower at the base than at the 
apex, transversely convex, coarsely and irregularly dentate at the 
sides (with about six teeth on each side), the surface thickly covered 
with coarse, rounded, very shallow punctures ; elytra parallel to 
beyond the middle, flattened on the disc, with rows of coarse, deep, 
closely placed punctures, the punctures subtransverse on the disc, 
rounded towards the sides, the interstices narrow, sparsely, very 
minutely punctate, the third, fifth, andseventh subcostate. Length 
43, breadth 13 mm. 


Hab. Tasmania—Hobart. 


Of this species Mr. Walker has sent three specimens 
to the British Museum, and one to myself; the description 
is entirely taken from the latter. They were all found 
under the bark of Hucalyptus. 


collected in Australia and Tasmania. 4.01 


CoRIPERA. 
Coripera, Pascoe, Journ. Ent., ii., p. 483 (1868) ; F. 
Bates, Ann. and Mag. Nat. Hist. (4), xui., p. 112. 
The name Pseudhelops, Guér., is incorrectly adopted 
for this genus in the Munich Catalogue, cf. I’. Bates, 
op. cit. 
Coripera deplanata. 
Adelium deplanatum, Boisd., Voyage de |’Astrolabe, 
Eni. u1., p. 277, Ins., t. 7, fig. 6 (1835). 
Hab. Tasmania—Launceston and Hobart. 


Not uncommon. Under logs and stones in rather 
damp places, especially between 2000 and 4000 feet 
elevation, on Mount Wellington (Walker). 


ADELIUM. 
Adeliwm, Kirby, Trans. Linn. Soc., xu., p. 420 (1818). 


Adeliwm licinordes. 

Adeliwm licinoides, Kirby, Trans. Linn. Soc.,xu.,p. 421; 
Boisd., Voyage de |’Astrolabe, Ent. 11., p. 276. 
Adelium cisteloides, Er. in Wiegmann’s Archiv, 1842, 

1, p. 176; Blessig, Horze Ent. Ross., i., p. 101. 


Hab. Tasmania—Hobart, Launceston, and Franklin. 


Not uncommon. Erichson’s A, cisteloides was from 
Tasmania, 


Adeliwm vicarium. 
Adeliwm vicarium, Pasc., Journ. Ent., ., p. 480. 
Hab. W. Australia—Darlington. 
Two specimens, apparently belonging to this species ; 
they are much larger than the type. 
Adelium scytalicum. 


Adeliwm scytalicum, Pasc., Ann. and Mag. Nat. Hist. 
(4), i., p. 137, 

Hab. W. Australia—Fremantle. 

Three specimens, perhaps referable to this species, 


from the type of which they differ in their less shining 
thorax and elytra. Described from Swan River. 


4.02 Mr. G. C. Champion on the Tenebrionidx 


Adelium abbreviatum. 
Adeliwm abbreviatum, Boisd., Voyage de |’Astrolabe, 
Ent. i, p. 281. 
Adelium impressum, Blanch. in Dumont d’Urville’s 
Voyage au Pole Sud, iv., Ins., p. 177, t. 11, fig. 18. 
Hab. Tasmania—Hobart, Launceston, and Franklin. 
Not uncommon. Ascends to 4,000 feet on Mount 
Wellington (Walker). 


Adelium latwm. 
Adelium latum, Pasc., Journ. Ent., ii., p. 482. 
Hab. Tasmania—Launceston. 
Four specimens, agreeing with the type. This species 


is a very close ally of A. abbreviatum, but it has the 
thorax more convex and with more obtuse hind angles. 


Adeliwm brevicorne. 


Adelium brevicorne, Blessig, Hore Ent. Ross., i., 
p. LOL, tS, tis. 12. 

Adelium neophyta, Pasc., Ann. and Mag. Nat. Hist. 
(4), ui., p. 136. 

Hab. SS. Australia—Port Adelaide. 


Numerous examples. 


Adeliwm porcatum. 
Carabus porcatus, Fabr., Syst. Ent., p. 239 ; Ent. Syst., 
L, p. 147; Oliv., Hnt., i., 35, p. 37, t. 7, fig. 84. 
Calosoma porculatum, Fabr., Syst. Hleuth., i., p. 211. 
Adeliwm caraboides, Kirby, Trans. Linn. Soe., xii., 
p. 466, t. 23, fig. 7; Boisd., Voyage de l’Astrolabe, 
Ent. 11., p. 274. 
Hab. 'Tasmania—Hobart. 
Adelium strigipenne. 
Adelium (Setrotrana) strigipenne, F. Bates, Trans. 
Ent. Soc. Lond., 1873, p. 365. 

Hab. N.W. Australia—Adelaide River. 

Three specimens, apparently referable to this species, 
and differing a little zmter se in the intricate elytral 
sculpture. The type of A. strigipenne, which I have 
examined, is labelled ‘* Australia.’’ 


collected in Australia and Tasmania. 4.03 


Adeliwm sp. (?) 
Hab. W. Australia—Albany. 


One specimen, perhaps an extreme form of A. lindense, 


Blackb., from Port Lincoln. 


Adeliwm commodum. 


Adelium commodum, Pasc., Ann. and Mag. Nat. Hist. 
(4), iii., p. 139. 


Hab. Tasmania—Hobart and New Norfolk. 


Not uncommon. Described from Tasmania. The 
entire upper surface is brassy in all the specimens 
received, Pascoe describes the insect as black, with the 
elytra eneous. 


Adelium tasmanicum, n. sp. 


Rather elongate, narrow, moderately convex, black or pitchy- 
brown, with a brassy lustre, feebly shining; the oral organs, 
antenne, the base of the tibiz, and the tarsi and coxe ferruginous. 
Head broadly flattened anteriorly, thickly and finely punctured, the 
epistoma arcuate-emarginate in front and limited behind by a well- 
defined groove ; antenne rather elongate in the male, about reaching 
the base of the prothorax in the female, the penultimate joints 
transverse, the apical one stout; prothorax as long as broad, 
truncate in front (as viewed from above), broadly and feebly 
emarginate at the base, slightly narrower at the base than at the 
apex, the sides a little rounded anteriorly, gradually and somewhat 
obliquely converging behind, the anterior angles rounded and 
declivous, the hind angles obtusely rectangular, the base obliquely 
foveate on either side, the surface closely, finely, distinctly 
punctured, and with a few widely-scattered setiferous impressions ; 
elytra about two and one-third times as long as the prothorax, and 
wider than it, feebly rounded at the sides, which are sharply 
margined anteriorly, a little flattened on the disc, with rows of fine, 
distinct, approximate punctures placed upon almost obsolete strix, 
the interstices flat, thickly and minutely. punctured, the second, 
fourth, sixth, and eighth with a few widely scattered setiferous 
impressions, the humeri very obtuse ; beneath thickly and finely 
punctured ; anterior tarsi with the four basal joints moderately 
widened in the male. Length 74—10, breadth 3—34 mm. 


Hab. Tasmania—Hobart and Launceston. 


4.04, Mr. G. C. Champion on the Tenebrionidx 


Three examples. Closely allied to A. commodum, 
Pasc., and occurring at the same localities, but differ- 
ing from it in having the thorax closely and distinctly 
punctured, the elytra with rows of fine but distinct punc- 
tures placed upon almost obsolete strize (in A. commodum 
the striz are sharply cut and indistinctly punctured) ; the 
setiferous punctures on the elytra are less distinct than in 
A. commodum, and placed on the second, fourth, sixth, 
and eighth, instead of on the third, fifth, seventh, and 
ninth interstices. 


Adelium nodulosum, n. sp. 


Q. Rather elongate, narrow, moderately convex, pitchy-brown, 
the elytra obscure castaneous in their outer half, the entire upper 
surface with a brassy lustre and feebly shining; the oral organs, 
antenn, the base of the tibie, and the tarsi ferrugimous. Head 
somewhat flattened anteriorly, thickly and moderately finely 
punctured, the epistoma feebly arcuate-emarginate in front and 
limited behind by a well-defined groove ; antennz extending to a 
little beyond the base of the prothorax, the penultimate joints as 
long as broad ; prothorax as in A. tasmanicum, but more densely 
and more distinctly punctured ; elytra as in A. tasmanicum, but less 
rounded at the shoulders, with rows of fine, distinct, approximate 
punctures placed upon almost obsolete striz, the interstices flat, 
thickly, very distinctly punctate, and each with a series of feeble 
nodular elevations, which become more distinct and subtuberculiform 
towards the apex. Length 83, breadth 35 mm. 


Hab. ‘Tasmania—Launceston. 


One example. This insect closely resembles A. 
tasmanicum, but differs from it in the denser punctuation 
of the thorax, and in having a series of rather closely 
placed nodular elevations on each of the elytral interstices, 
these elevations becoming tuberculiform at the apex. 
From Brycopia tuberculifera (which has somewhat similar, 
but fewer, elevations on the elytra) it may be known by 
the more transverse and less prominent eyes, the much 
finer punctures of the elytral striz, the coarser punctures 
of the interstices, etc. The elytra are subtruncate at the 
base. 


Dryoria. 


Dinoria, Pascoe, Ann. and Mag. Nat. Hist. (4), ii, 
p. 141 (1869). 


collected in Australia and Tasmania. 4.05 


Dinoria picta. 
Dinoria picta, Pase., loc. cit. 
Hab. 'Tasmania—Hobart. 


Apparently a common insect at Hobart. Under bark 
and among chips in damp situations (Walker). 


Dinoria celioides. 


Dinoria celioides, Pasc., Ann. and Mag. Nat. Hist. 
(4), v., p. 1038. 


Hab. Tasmania—Hobart. 


Originally recorded from Queensland. Same habits as 
D. picta (Walker). 


Brycopia. 


Brycopia, Pascoe, Ann. and Mag. Nat. Hist. (4), i., 
p. 141 (1869). 
Brycopia tuberculifera, nu. sp. 

Brycopia tuberculifera, F. Bates, in litt. 

Moderately elongate, rather narrow, convex, pitchy-black with 
an eneous lustre, shining; the oral organs and antenne ferru- 
ginous, the basal joints of the latter usually darker ; the legs 
pitchy-brown, the tarsi ferruginous. Head sparsely, finely, irre- 
gularly punctate, and also with a few coarse setiferous impressions 
in front, the epistoma not clearly defined, feebly arcuate-emarginate 
in front, and limited behind by a rather broad transverse 
depression ; the eyes moderately large, coarsely granulated, convex, 
prominent, rounded as seen from above ; antennz rather elongate, 
thickening outwardly, the apical joint very stout; prothorax 
subquadrate, convex, as long as broad, subtruncate in front (as 
viewed from above), feebly bisinuate at the base, the sides 
moderately rounded anteriorly and sinuous and converging behind, 
the hind angles acute, the anterior angles obtuse and declivous, the 
base obliquely depressed and foveate towards the sides, the surface 
very finely and rather closely punctured, and with a few coarse, deep, 
widely scattered setiferous impressions ; elytra about two and one- 
third times as long as the prothorax, and wider than it, somewhat 
oval, with rows of moderately coarse, subapproximate punctures 
placed upon almost obsolete strie, the interstices flat, sparsely, 
finely punctate, and with scattered, feeble tubercular prominences, 


406 Mr. G. C. Champion on the Tenebrionidex 


which become more distinct towards the sides and apex, the third, 
fifth, and seventh each with three or four setiferous impressions, 
the humeri very obtuse. Length 8?—9j, breadth 3;—3} mm. 


Hab. Tasmania—Hobart, Launceston, and Franklin. 


Four specimens have been sent to me by Mr. Walker, 
and others are contained in Mr. F. Bates’s collection, the 
latter bearing the name I have adopted. LB. tuberculifera 
has very much the facies of Adeliwm commodum, Pasc., 
an insect occurring in the same localities ; but it may be 
easily known from that insect by the prominent and 
more rounded eyes, the imperfectly defined epistoma, the 
tuberculate elytra, ete. 


CHALCOPTERUS. 


Chalcopterus, Blessig, Horee Ent. Ross., 1., 1, p. 103 
(1861); Blackburn, Proc. Linn. Soc. N.S.W. (2), 
vil, p. 415 (1892), and viii., pp. 53—56 (1893). 


Chalcopterus longipennis. 


Amarygmus longipennis, Hope, Proc. Ent. Soc. Lond., 
1842, p. 79; Trans. Ent. Soc. Lond., 1845, p. 109 
(? 2ec Blackburn). 


Hab. W. Australia—Fremantle. 


One specimen, also another in the British Museum-set, 
agreeing with Hope’s type. The Rev. IT’. Blackburn in 
his description of this species (Proc. Linn. Soc. N.S.W. 
(2) vii, p. 456) states that the tarsi are clothed with 
black hairs beneath ; in the insect before me the hairs are 
fulvous, but as I did not examine the tarsi of Hope’s 
type, the identification with his C. longipennis is perhaps 
not quite certain. 


Chalcopterus howittr. 


Amarygmus howitti, Pasc., Ann. and Mae. Nat. Hist. 
(4), ii., p. 348. 

Chalcopterus cwpripennis, Blackb., Proc. Linn. Soe. 
N.S.W. (2), vi., p. 422 (nec Hope). 

Hab. Tasmania—Hobart. 


Not hitherto recorded, I believe, from Tasmania. It 
is a common species in Southern Australia. 


collected in Australia and Tasmania. 407 


Chalcopterus wridicolor. 


Chalcopterus iridicolor, Bless., Horse Ent. Ross., 1., 
p. 107, t. 4, figs. 8, 6 (1861) ; Blackb., Proc. Linn. 
Soc. N.S.W. (2) viii., p. 67. 

Hab. Tasmania—Hobart. 


I refer with some doubt a very variable Chalcopterus 
obtained in plenty in Tasmania by Mr. Walker to this 
species, the type of which was from Melbourne. The 
Rev. T. Blackburn in his recent papers on the genus 
Chalcopterus does not mention any species specially as 
from ‘l'asmania. 


Chalcopterus semiticus. 


Amarygmus semiticus, Pasc., Ann. and Mag. Nat. Hist. 
(4), ii., p. 349 (1869). 
Hab. N. Australia—Cape Bougainville. 


One specimen of this species has been sent by Mr. 
Walker to the British Museum. It agrees well with the 
type from Port Denison. C. semiticus is apparently 
unknown to Mr. Blackburn (cf. Proc. Linn. Soc. N.S.W. 
(2), viii., p. 69). 


AMARYGMUS. 


Amarygmus, Dalman, Anal. Ent., p. 60 (1823) (par- 
tim); Blackburn, Proc. Linn. Soc. N.S.W. (2), 
vii., p. 415 (1892), and viii, p. 87. 


Amarygmus tyrrhenus. 


Amarygmus tyrrhenus, Pasc., Ann. and Mag. Nat. 
Hist. (4), v., p. 105; Blackb., Proc. Linn. Soe. 
N.S. W.. (2), vii., p. 91. 


Hab. W. Australia—Albany. 
One specimen. 


Amarygmus perpleaus. 
Amarygmus perplewus, Blackb., Proc. Linn. Soc. 
N.S.W. (2), viii., p. 102 (1893). 


Hab. N.and N. W. Australia—Port Darwin, Adelaide 
River, Roebuck Bay. 


4.08 Mr. G. C. Champion on the Tenebrionide. 


Sent in plenty by Mr. Walker. The numerous speci- 
mens from Adelaide River have the elytra more or less 
cyaneous, instead of black, as in the type and in the 
examples from Port Darwin. Mr. Blackburn’s specimens 
were from the North Territory of 8. Australia; he has 
examined some of Mr. Walkerv’s examples. 


EXPLANATION OF Prats VIII. 


Fic. 1. Diphyrrhynchus ellipticus, @. 
2. Ennebeus australis. 


2a. ~ % antenne. 

3. Ennebeopsis pruinosus. 

3a. 3 85 antenne. 

4. Diphyrrhynchus apicalis, @. 

5. Edylius canescens, 2. 

5a. fe i labium. 

5D. 98 » maxilla and maxillary palpus. 


6. Lyphia tasmanica. 
7. Caanthus gibbicollis. 


7a. 5 e maxilla and maxillary palpus. 
7b. a FF antenne. 

8. Paratoxicum iridescens. 

8a. 5 e maxilla and maxillary palpus. 
8b. " e labium. 

8c. * 4) antenne. 


9. Nyctozoilus sexcostatus. 
10. Saragus bicarinatus. 
11. Decialma erichsoni. 
12. Omolipus cyaneipennis, ¢. 
13. Titena tasmanica. 
14. Pheennis fasciculata. 

15. Ectyche cerulea, 3. 


0005 


XII. Description of the Larva and Pupa of Papilio 
homerus, Fab. By Cuarues B. Taytor, F.E.S. 


[Read February 28th, 1894.] 


Axsout two or three years ago, a lady residing in the Bath 
district of the Parish of Saint Thomas, and a diligent 
collector of insects, discovered seven examples of a large 
Caterpillar of a beautiful green colour, on a tree bearing 
large subovate glossy green leaves (botanical name un- 
known) growing near a certain locality known as the 
“Fountain Road.” Of these she was good ‘enough to 
send me four specimens, which unfortunately, however, 
never reached my hands, as they were lost in transit, 
while I shortly afterwards heard that the other three 
examples had died before assuming the pupal state, as a 
result of confinement and want of proper attention. 
Towards the middle of last year Mrs. Swainson found 
another of these Caterpillars, which she sent down to the 
Museum of the Jamaica Institute for identification. It 
arrived at a time when there was no experienced 
Entomologist in charge (the post of Curator being then 
temporarily vacant), but on seeing it I ventured to express 
the belief that it was the larva of P. homerus. This 
individual also died during the process of casting the 
larval skin. Through the kindness of Lady Blake I was 
enabled to make a special expedition to Bath during 
November last, in search of this larva, and the day after 
my arrival (the 27th) had the good fortune to find an 
almost full-fed specimen, which developed a male 
homerus on the 29th of December following. It is 
worthy of note that all these Caterpillars were obtained 
from the same tree, notwithstanding that there were many 
others of the same species in the immediate neighbour- 
hood. 

Larva. Length about 2} inches, general appearance incrassated, 
the body being thickest from the 3rd to the 5th segments, from 


TRANS. ENT. SOC. LOND. 1894,—PaRT 1. (JUNE.) 2D 


A10 = On the larva and pupa of Papilio homerus. 


thence tapering gradually to the anal extremity, which is abruptly 
truncated. Head small, pale ochreous or brown, deflected, and, 
when the larva is at rest, almost completely obscured by a pro- 
jection of the anterior margin of the 1st segment. 

Colour protective, harmonizing closely with the tint of the leaf 
of the food plant. Upper surface a rich deep chrome-green 
extending down to about the spiracular region, where it terminates 
with a more or less irregular or sinuous outline, most pronounced 
towards the posterior segments, where the green is sharply divided 
from the colour of the under surface by a narrow edging of 
white. 1st segment with a short transverse dorsal bar of pale dun 
or ochreous brown; a broad, curved band of the same colour, 
bearing a few lines and spots of black, runs in a posterior direction 
across the dorsal surface of the 3rd segment, and down to the sides 
of the 4th segment, where, curving slightly upwards, it ends in 
club-shaped dilations, enclosing a simple ocellus coloured as 
follows: On a comma-shaped ground patch of black is drawn a 
rather broad ring of dun, bordered below with bluish-grey followed 
by a black iris, with a bluish-white pupil. Across the posterior 
margin of the 5th segment is drawn a most conspicuous luniform 
stripe of white, having a raised appearance, as if laid on with 
several coats of body colour, and which is in turn marked with 
little transverse bars of pale lavender, five in all; the horns of the 
crescent have this time a slightly anterior direction, and terminate 
at the sides between the 4th and 5th segments just at the junction 
of the green with the brown of the under surface. 

Anterior margin of the fifth segment, immediately in front of 
the white stripe, finely irrorated with white ; between the 5th and 
6th segments is a broad transverse band of deep black, more or less 
completely hidden by the segmental fold, except when the larva is 
in motion, when it is very conspicuous. Ventral surface, including 
abdominal legs, dull dark brown. A broad band of this colour, 
suffused with a lavender-grey tint, bearing a few scattered spots of 
blue-grey, and having a very irregular or jagged outline, is con- 
tinued obliquely upwards in a posterior direction from the 8th 
segment, and, meeting on the dorsal surface of the 10th segment, 
descends again, obliquely, narrowing rapidly however before joining 
the brown of the under surface, thus enclosing an irregular 
diamond-shaped patch of green. Segmental folds strongly marked 
on the ventral surface. 

Pura, of usual shape, not greatly angulated. Colour, wholly 
dark wood-brown, with two puirs of white tubercles on the dorsal 
surface of the middle abdominal segments. 


€ 4h 5 


XIII. Notes on Dorydium (?) westwoodi, Buchanan 
White, with observations on the use of the name 
Dorydium. By Wituiam F. Kirsy, F.L.S., F.ES., 
Assistant in Zoological Department, British 
Museum (Nat. Hist.), South Kensington. 


[Read February 28th, 1894.] 


A. rew days ago I received a letter from Mr. Herbert 
Clark, of Christchurch, New Zealand, enclosing two 
specimens of a very curious Homopterous insect, which I 
have succeeded in identifying with Dorydiwm westwoodi, 
Buchanan White. 

Mr. Clark writes, ‘‘ I found it on the rushes which grow 
in damp situations. In colour and shape it so much 
resembles a piece of dried rush that I have never been 
able to find a specimen except by switching the net 
amongst them. I think they are scarce and very local, as 
the specimens I have captured were taken in a space a 
few yards square, and I have never been able to collect 
them elsewhere. The place where they were found was in 
a plantation of Pinus insignis about a quarter of a mile 
from the sea, and a few chains from the river. The time 
of appearance is from the beginning of November to the 
end of the year.” 

The insect belongs to the family Jasside, and the 
references are as follows: 


Dorydium westwoodi. 


Buchanan White, Ent. M. Mag., xv., p. 215 (1879) ; 
Signoret, Ann. Soc. Hnt. France (5), x., p. 43, 
pl. 1, fig. 38, details (1880). 

One of the specimens received from Mr. Clark is a 
perfect insect (a male, I think), and the other a pupa, as 
may be seen by the rudimentary tegmina. The mature 
specimen measures 9 lines in length, and agrees with 

TRANS. ENT. SOC. LOND. 1894.—ParT Il. (JUNE.) 


412 Mr. W. F. Kirby on Dorydium (?) westwood, 


Buchanan White’s description better than with Signoret’s, 
except in size (Signoret gives 11 mm., and Buchanan 
White 14). The insect is probably variable in size and 
markings, or in the absence of markings, or there may 
be more than one allied species; without a good series, 
and perhaps observation of the living insects, it might be 
difficult to clear up this point. 

Buchanan White received three specimens from Wake- 
field, which had been taken by Fereday near Christ- 
church. He remarks, ‘‘'This curious species strongly 
resembles the seed of one of the larger grasses.” 

Signoret received his specimens from John Scott ; they 
appear to be darker than the types; but the allied 
Australian species described by G. R. Waterhouse 
(Trans. Ent. Soc.,i1., p. 195) under the genus Cephalelus, 
differ much in size and colour. They differ, too, from the 
species referred to Dorydiwm by the longer and more 
tapering frontal prominence, 

There is a curious confusion relating to the genera 
Cephalelus and Dorydiwm which I will try to clear up. 
In 1832, Percheron (Mag. Zool. Cl. ix., pl. 48) described 
and figured a species from an unknown locality, which he 
called Cephalelus imfumatus. 

In 1839, Burmeister (Handb. Ent., u1., p. 106) de- 
scribed an insect from the Cape under the name of 
Dorydium paradovwm, but at p. 1006 stated, that his 
insect was identical with Percheron’s, and shortly after- 
wards figured it as such in his “Genera Insectoram.” 
Instead of dropping the generic name, as he was bound 
to do, he used it again for an allied insect from Sicily, 
which he described and figured as Dorydiwm lanceolatum. 
This figure is referred to at p. 1006, and appears to have 
been published while Part II. of the ‘‘ Handbuch” was 
passing through the press, and would, therefore, have 
priority over it. 

Nor does the confusion end here. In 18387, Herrich- 
Schiffer described (Panzer, Heft 144, pl. 6) an insect 
from Nuremberg as Jassus paradoxus; oddly enough, 
another species with a long frontal prominence, though, 
judging from the figure, I should say that it is certainly 
not congeneric with either Dorydiwm I. (Cephalelus), or 
Dorydium I. Signoret (Ann. Soc. Ent. France (5), 
ix., pp. 259-265) confuses everything. Firstly, he gives 
the genus Cephalelus with four species : infumatus, 


with observations on the name Dorydium. A13 


Perch., and percheroni, Guér.,* from South Africa, and 
C. marginatus and brunneus, G. R. Waterhouse, from 
Australia. I may say that the two latter are closely allied 
to, if not actually congeneric with, Dorydium (?) west- 
woodi, and do not belong to the South African genus 
Cephalelus. Next to Cephalelus, Signoret places the 
genus Dorydiwm, with the two species, D. lanceolatum 
and paradovum, Burm.; but for the latter he gives the 
locality Paris; and his insect is evidently paradowus, 
Herrich-Schiiffer ; Burmeister’s species being, as we have 
seen, Cephalelus infumatus, Perch., or a closely-allied 
species. 

It is clear that D. lanceolatum, Burm., must be 
regarded as the type of Dorydiwm, and that most of 
the species referred to Dorydiwm will have to be 
removed either to Cephalelus or new genera formed for 
them, if necessary. But this task I will leave to the next 
specialist who takes up the Jassidz. 


** T cannot find a reference to this species, and suspect it ma 
) Pp 
be an error. 


oan. i 


XIV. Some new species of Membracide. By the Rev. 
Canon Fow ter, M.A., F.L.S. 


[Read 14th March, 1894.] 


I wave for some time past been engaged in working out 
the Membracidz for the Biologia Centrali-Americana, and 
in the course of my work I have received a large number 
of specimens from the Vienna Museum through the 
kindness of Dr. Ganglbauer and Herr Handlirsch, and 
from the Belgian Museum through the kindness of M. 
Severin; I am also much indebted for help to Dr. 
Aurivillius of the Stockholm Museum, but as_ the 
specimens he has sent me are exclusively Central Ameri- 
can there are none that I can here notice; among the 
others, however, there are several interesting species 
which lie outside the scope of the Biologia, and a few of 
these I have described below. 

At present there is great confusion with regard to 
certain of the genera of this very extensive and interesting 
group of insects, partly owing to the fact that Walker’s 
work, though embracing a large number of genera and 
species, was very slight, and in some cases most con- 
fusing, as he places the most dissimilar insects under one 
genus, and partly because Stal, if in doubt, appears to 
have disregarded Walker’s work, and redescribed the 
species. 

Tropidoscyta transiens, sp. n. 

Parva, brevis, testacea, aureo-pubescens, capite magno, antice 
rotundato, pronoto inzequali, antice in cornu brevissimum rotun- 
datum fuscum producto, carinis duabus supra humeros valde 
elevatis, humeris ipsis prominulis, dorso remotius punctato, carina 
centrali instruct, a latere viso pone medium fortiter sinuatim 
depresso, apice fusco perobtuso ; tegminibus apicem prothoracis 
longe superantibus, hyalinis, venis testaceis, maculisque quibusdam 
obscure brunneis ; corpore subtus fusco, postice testaceo ; pedibus 
testaceis. 

TRANS. ENT, SOC. LOND. 1894,—PART II. (JUNE.) 


416 Rey. Canon Fowler on some 


A small short species, testaceous, with the metopidium, the very 
short bluntly rounded horn of pronotum, the apical portion of the 
two strong caring above shoulders, and the very blunt apex fuscous: 
the dorsum if viewed from the side is much depressed behind 
middle, and is extended into a short, blunt, very slightly deflexed 
apex ; if viewed from the front the metopidium is large and broad, 
and appears to be furnished with five points formed by the anterior 
. horn, the apex of the two suprahumeral carinz and the slightly 
prominent shoulders ; tegmina hyaline, with obscure dark markings, 
underside fuscous, hinderpart testaceous ; legs testaceous. Long, 
cum tegm., 5 mm. ; lat. int. hum., 3 mm. 


Hab. Rio Janeiro. ¢. (Belgium Museum Collection.) 


This species is closely allied to Hrecthia bicolor, Walk., 
and also resembles Tropidoscyta torva, Stal (Membracis 
torva, Germ.) ; it is one of the species that complete the 
transition from Membracis to Bolbonota; no two genera 
can well be less alike than these, yet connecting forms 
occur which make it almost impossible to separate the 
intervening genera satisfactorily. 


Bolbonota rufo-notata, sp. n. 


Parva, nigra, pronoto carinato, a latere viso fere sequali, apice 
depresso, humeris, dorsoque plus minusve, rufo-ferrugineis ; 
tegminibus pellucidis, basi nigré vel nigro-brunned, venis fuscis ; 
pedibus rufis vel ferrugineis. 

A small species, variable in colour, black with the shoulders and 
more or less of the disc of the pronotum of a ferruginous red 
colour, sometimes the black and sometimes the red colour pre- 
vailing ; the pronotum is carinate and uneven if viewed from ahove, 
but if viewed from the side the central carina is almost level to 
apex, where it is depressed ; punctuation coarse but obscure; 
tegmina transparent, hyaline, with the base black or brownish-black, 
sometimes ferruginous internally, veins dark and conspicuous; legs 
red or ferruginous. Long. 2 mm. ; lat. int. hum., 13 mm. 

Hab. New Granada (Nolken), (Vienna Museum 
Collection) ; five examples. 


The species is allied to B. pusto, Germ., but the latter 
species is considerably larger, longer in proportion, quite 
black, and more coarsely punctured, with the dorsum 
more level and the apex differently formed; it is also 
closely akin to B. inxqualis, Fab., but, apart from the 
colour, it is much more finely punctured than that species 
and has the dorsum more level if viewed from the side. 


new species of Membracide. ALT 


Bolbonota auro-sericea, sp. n. 


Opaca, dilute-testacea, pube aureo-sericed dense vestita ; capite 
pronotoque sat dense subtiliter punctatis, illo elongato sat angusto, 
hoc carina centrali integra, carinisque tribus utrinque supra humeros 
instructo, dorso a latere viso «quali, recto, mox ante apicem de- 
presso, apice utrinque carin’ brevi obliqué ; tegminibus flavis, ad 
medium dilutioribus, ad basim apicemque brunneo-variegatis ; 
corpore subtus pedibusque testaceis, pectore femoribusque ad 
partem infuscatis, 


A rather large light testaceous species, thickly clothed with fine 
golden pubescence ; pronotum very finely punctured with the 
central keel and three abbreviated keels on each side above 
shoulders, and a small oblique keel on each side of apex, distinct, 
the rest of the surface even, dorsum almost straight, if viewed from 
the side, depressed just before apex ; tegmina yellow, dark before 
apex and near base; underside and legs testaceous, chest and 
upper part of femora infuscate. Long. cum tegm.,5 mm. ; lat. 
int. hum., 3 mm. 


Hab. North America. ¢. Coll. Signoret (Vienna 
Museum Collection). 


As far as I know this is the only species of Bolbonota yet 
recorded from North America; the genus is omitted in 
Dr. Goding’s synopsis (Trans. Am. Ent. Soc., xix., 253) ; 
it is a very distinct species. 


Triquetra obtusa, sp. n. 


Testacea, unicolor, pronoto tectiformi, sat fortiter ad latera 
subrugosius punctato, dorso recto antice altissimo in cornu haud 
producto, sed obtuse angulato, metopidio longo, fere recto, cornubus 
humeralibus porrectis, acutis, apice longo, acuto usque ad apicem 
tegminum extenso ; tegminibus totis hyalinis, venis dilute testaceis; 
pedibus testaceis. 


A large species of a unicolorous testaceous colour, with the pro- 
notum tectiform, rather strongly and at the sides subrugosely 
punctured, with the dorsum continued in a straight line from apex 
(which is long, sharp, and acute) to the metopidium, which is then 
abruptly declivous in almost a straight line ; at the place where the 
dorsum and metopidium meet there is no trace of a horn as in the 
allied species; tegmina entirely hyaline, with the veins light 
testaceous ; legs testaceous, Long. 14—15 mm.; lat. int. corn. 
hum., 10} mm, 


418 Rev. Canon Fowler ow some 


Hab. New Granada (Nolken), (Vienna Museum Collec- 
tion), three specimens; Ecuador (H. de Ville), (Belgium 
Museum Collection), four specimens. 

This species may easily be distinguished from 1’. grossa 
(with which latter species 7. virgata and T’. virescens are 
synonymous) by the shape of the front part of the 
prothorax. I found the specimens above referred to 
among the unnamed material of the Stockholm and 
Vienna Museum Collections, and set on one side as a 
new species; it may possibly be referred to Triquetra 
grossa as a hornless variety, but I have examined a very 
large number of the last-named species, and have not 
found one specimen like 7’. obtusa among them. 


Potnia jansoni, sp. n. 

Testacea, capite sat magno, metopidio supra caput plicato, 
pronoto antice in cornu brevissum. obtusum, rufescens late nigro- 
marginatum extenso, fortiter punctato, cariné centrali, costisque 
utrinque quattuor vel quinque sat regularibus instructo, humeris 
leviter prominulis, lateribus sensim in processum longum extensis ; 
tegminibus prothoracem vix superantibus testaceo-hyalinis, apice 
ipso fumoso ; corpore subtus pedibusque testaceis. 

Somewhat elongate for a Potnia, testaceous, with the pronotum 
strongly punctured, produced in front into a very short blunt horn, 
which is reddish, with the margins broadly black, the black colour 
being extended for a short distance on the dorsum, central keel 
and four or five regular coste on each side well marked, apical 
process long, very slightly deflexed; tegmina produced very slightly 
beyond prothorax, testaceous hyaline with the apical margin 
smoky ; legs and underside testaceous. Long. cum tegm., 8 mm. ; 
lat. int. hum., 3} mm. 


Hab. Demerara (Janson), ¢. (Belgium Museum 
Collection.) 


Potnia perobtusa, sp. n. 
Precedenti affinis, sed minor, colore dilutius testaceo, pronoto 


antice minus porrecto et perobtuse angulato, processuque postico 
breviori et rectiori, apice magis obtuso, distinguendus, 


Very like the preceding, but smaller and lighter coloured and 
less strongly punctured, with the pronotum very obtusely angled 
and not extended into a horn, and with the posterior process 
shorter and straighter ; the tegmina are hyaline with the veins 
testaceous. Long, cum tegm, 7 mm. ; lat. int, hum., 3 mm. 


new species of Membracide. 419 


Hab. Rio Janeiro and Botafogo, Brazil; a fair series. 
(Belgium Museum Collection.) 


Both these species have the facies of Hnchotype, rather 
than of Potnia, the pronotum being distinctly and 
regularly ribbed ; but the presence of four apical areas on 
the wings prevent their being classed with the former 
genus as at present constituted. 


Enchotype concimna, sp. n. 

Parvula, castanea, pronoto fortiter rugose punctato, linea cen- 
trali aliisque utrinque irregularibus utrinque elevatis, humeris a 
fronte viso obtuse prominulis, cornu antico fere recto, brevi, lato, 
acuto, unicarinato, apice postico longo, acuto ; tegminibus hyalinis 
venis testaceis, apicem versus fuscis ; pedibus testaceis. 

A small dark castaneous species,, with the pronotum coarsely 
and rugosely punctured, with the central line and others on each 
side elevated, and with the shoulders, if viewed from the front, 
obtusely prominent ; anterior horn situated just above metopidium, 
short, broad, and almost upright, and sharp at apex; tegmina 
vitreous, with the veins testaceous towards base, darker towards 
apex; legs testaceous. Long. cum tegm.6mm,, lat. int. hum. 4 mm. 

Hab. Cuba (Mayr., Vienna Museum Collection). 

This species is closely allied to Hnchotype fairmairet, 
Stal. (Hoplophora fairmairei, Guér.), but differs in the 
shape of the prothoracic horn, which is much shorter, 
less porrect, and more upright, and also in the smaller 
size, less strong pubescense and darker venation of the 
tegmina. 

Enchotype is a sub-genus of Stal, which is closely 
allied to Potnia, from which it is chiefly separated by the 
fact that the wings have three instead of four apical 
areas ; it is very distinct from Hoplophora, of which it is 
by some authors reckoned as a sub-genus. ‘The species 
above described is plainly an Hnchotype, but it has 
besides the three apical alar veins a very minute fourth 
one on the exterior margin, which appears to be a thick- 
ening of a vein and to be quite distinct from the exterior 
area of Potnia. 


Hoplophora unicolor, sp. n. 
Longa, lata, parallela, capite et pronoto toto rufo-ferrugineo 
unicolori, ille parvo, hoe fortiter punctato, humeris prominulis, 


420) Rev. Canon Fowler on some 


dorso a latere viso fere recto, haud elevato, lined centrali et tribus 
vel quattuor aliis utrinque sat regularibus, elevatis, apice acuto ; 
tegminibus elongatis, testaceo-hyalinis, immaculatis, venis totis 
testaceis ; corpore subtus infuscato, pedibus rufo-testaceis. 

A long, broad, parallel species, with the head and pronotum 
entirely rufo-ferruginous, unicolorous ; head small, thorax very 
coarsely punctured, with a central raised line and three or four on 
each side which are almost regular and very little broken, apex 
sharply pointed; tegmina long, testaceo-hyaline, without any dark 
spots, and with all the veins testaceous ; underside infuscate, legs 
rufo-testaceous. Length sine. tegm. 10 mm. ; cum tegm. 15 mm. ; 
lat. int. hum. 7 mm. 


Hab. Colombia. 


Two female specimens (Vienna Museum Collection). 
This species is related to H. gigantea, and is very closely 
allied to a new species which I have described from 
Mexico, under the name of H. signoretv. 


Hille ecuadorensis, sp. n. 


Elongata, brunnea, pronoto dense et fortiter subrugose punctato, 
carina centrali, lineisque quibusdam elevatis utrinque instructo, 
cornu dorsali lato obtuso tricarinato infuscato, in metopidium 
dorsumque sensim descendente ; tegminibus brunneo-hyalinis, parte 
dimidia basali fortiter punctat&, opac&; corpore subtus nigro ; 
femoribus infuscatis. 

A small elongate castaneous-brown species, with the pronotum 
thickly and strongly and subrugosely punctured, with a central 
keel and four or five elevated lines on each side, and with an obtuse 
dorsal horn, which is tricarinate and infuscate and slopes gradually 
down to the metopidium and the dorsum; tegmina brownish 
hyaline, with the basal half strongly punctured ; underside black. 
Long. 7 mm. ; lat. int. hum. 3} mm. 


Hab. Hceuador (Coll. Signoret. . Vienna Museum 
Collection). 


Telamona prealta, sp. 0. 


Fusco-brunnea vel fusca, carin’ centrali prothoracis nigra, 
capite pronoto multo angustiori, illo fortiter punctato, humeris 
prominentibus, subauriculatis, ad latera rugose carinato, protu- 
berantid dorsali magn4, prealtd, apice antice late rotundato, postice 
minus alto margine recto, angulo postico obtuso, ad basim utrinque 
fortiter impressd ; tegminibus externe hyalinis, ceteris, preecipue 
ad apicem, fumosis ; corpore subtus pedibusque fusco-brunneis. 


new species of Membracide. 421 


Dark fuscous browa, with the central carina of the pronotum 
and sometimes apical portion of the dorsal protuberance black ; 
pronotum strongly punctured with a very large and elevated dorsal 
protuberance ; this is strongly declivous behind and in front, and 
in front is broadly sinuate before metopidium ; its apex is broad, 
rounded and elevated in front, less high and with a straight margin 
behind, and with the posterior angle obtusely rounded, but marked; 
tegmina hyaline externally, with the base and apex dark and the 
central portion smoky ; underside and legs fuscous. Long. cum 
tegm., J—10 mm.; lat., 5—6 mm. 

Hab. Saguenay, Brazil (V. Huart); several speci- 
mens (Belgium Museum Collection), 


Telamona ruficarinata, sp. n. 


Sat angusta, capite nigro rufo-variegato, pronoto rugoso, fortiter 
punctato, brunneo-testaceo fusco-variegato, cariné centrali rufa a 
capite usque ad apicem extensa, et carinis quibusdam insequalibus 
et rugosis utrinque instructo, protuberantié dorsali paullo ante 
medium, magna, laté, rotundaté, cornubus anterioribus ad humeros 
latis, porrectis, subauritis, paullo reflexis, ad apicem rotundatis ; 
tegminibus nigro testaceoque variegatis, venis rufescentibus, apicem 
versus dilutioribus; abdomine, basi segmentorum flavescenti 
excepta, nigro ; pedibus testaceis, ad partem infuscatis. 

Comparatively narrow, with the pronotum testaceous-brown 
variegated with black, and with a red carina running from just 
above head to apex; tegmina variegated with fuscous and testaceous 
and with the veins rufescent ; abdomen black with the apex of the 
segments yellowish ; chest black ; legs testaceous, partly infuscate; 
prothorax rugose, strongly punctured, irregularly carinate on each 
side of the central line, with a large broad protuberance a little 
before middle, descending in front abruptly to the dorsum, and so 
leaving a flat and level space before metopidium ; anterior horns 
porrect, broad, rounded at apex; apex of prothorax extending 
nearly to apex of tegmina. Long. cum tegm., 10 mm. ; lat. int. 
hum, 5 mm. 


Hab. Bogota (Coll. Signoret. Vienna Museum Collec- 
tion) ; one male specimen. 


Lucilla intermedia, sp. n. 

Elongata, angusta, ab humeris pronoti leviter obtuse prominulis 
ad apicem sensim angustata, griseo-brunnea, capite sat magno, 
pronoto perobscure fusco-variegato, carina centrali, lineisque 
utrinque tribus vel quattuor, sat regularibus, instructo, dense et 
fortiter punctato, dorso a latere viso late et persensim ad medium 


422 Rev. Canon Fowler on some 


elevato postice depresso; tegminibus hyalinis ad basim externe 
punctatis, opacis, pedibus testaceis. 

An elongate narrow species, of a griseous brown colour, with a 
few obscure fuscous markings on pronotum, which is almost uni- 
colorous ; head rather large, ocelli distinctly nearer to one another 
than to eyes, central keel strong and distinct throughout from just 
above head to apex, which reaches beyond or at least as far as apex 
of tegmina and is very gradually and bluntly pointed ; the lateral 
carine are fairly regular, three or four on each side ; the dorsum if 
viewed from the side is very broadly and slightly elevated, depressed 
towards apex, and almost imperceptibly and broadly truncate to 
metopidium ; tegmina hyaline, punctured towards base ; legs testa- 
ceous. Long., 7 mm. ; lat. int. hum., 3 mm. 


Hab. Ecuador (Coll. Signoret. Vienna Museum 
Collection). 

I have referred this species to Stal’s genus Lucilla, as 
it is very closely allied to Oxygonia viridula, Fairm., 
which Stal mentions as his type of the genus. I am 
somewhat doubtful, however, as to the generic value of 
Hnnya and Lucilla, and they cannot be separated from 
the Hille group on the character assigned by Stal (v., 
Ofv. af. Kongl., Vet. Ak. Férh., 1867, No. 7, p. 555), 
*corio areola discoidali destituto,” as a discoidal area is 
certainly sometimes present. JL. intermedia differs from 
L. viridula in its more eiongate form and in the shape of 
the dorsum, which is much flatter and highest behind 
middle, whereas in the last-named species it is highest 
before middle; the sculpture is also different. The two 
types of Oxygonia viridula in Signoret’s collection, from 
the Vienna Museum, are before me. I believe, how- 
ever, that they belong to two different species. 


Paradarnoides, nu. g. 


Oblongus, capite longitudine latiori, fronte angust4, ocellis 
inter se quam ab oculis distincte magis remotis, oculis promi- 
nentibus ; pronoto ad humeros leviter obtuse prominulos latissimo, 
deinde sinuatim angustato et in processum longum, ad basim latum, 
postice subparallelum, a latere viso leviter deflexum, acutum, 
utrinque ad latera ab humeris usque ad apicem profunde im- 
pressum, producto, dorso a latere viso ad basim processus leviter 
sinuato, carina centrali per totum percurrenti ; tegminibus apicem 
prothoracis vix superantibus, areis apicalibus quinque, discoidalibus 
tribus ; alis areis apicalibus quattuor. 


new species of Membracide. 423 


Oblong, subparallel, with the ocelli evidently further removed 
from one another than from eyes, which are prominent ; pronotum 
with the shoulders, which are set far back, obtusely prominent, 
behind these gradually narrow, with a gentle sinuation and con- 
tinued into a long process, which is strongly and broadly impressed 
at margins from shoulders to apex, and if viewed from the side is 
gradually deflexed and evidently carinate, the carina being con- 
tinued to the frontal margin of the metopidium ; tegmina slightly 
exceeding the apex of prothoracic process, with five apical and three 
discoidal areas, the areas being for the most part subequal and 
oblong ; wings with four apical areas ; legs rather slender. 


This genus appears to be allied to Darnoides as re- 
presented by its type Darnoides limbata, but may be 
easily known by the position of the ocelli, the more 
prominent eyes, and the fact that the tegmina have three 
discoidal areas; the form, moreover, is more subparallel 
and less narrowed behind, and the shape and sculpture 
of the prothorax behind shoulders is very different. 


Paradarnoides severini, sp. n. 

Major, capite fusco, rugoso, pubescenti, metopidio rugosé punc- 
tato, fusco-testaceo, maculis duabus fuscis antice convergentibus, 
interdum fere deficientibus, pronoto pone humeros fortius punctato 
vitt’ lata fused, deinde dilute testaceo, apice longo fusco ; tegmini- 
bus fusco-hyalinis, ad basim punctatis, venis testaceis ; corpore 
subtus fusco pubescenti; pedibus testaceis plus minusve infuscatis. 


Head rugose, pubescent, fuscous, pronotum testaceous in front, 
then broadly fuscous behind shoulders, and then light testaceous, 
the apex being fuscous or reddish fuscous; on the metopidium 
there js a broad fuscous patch on each side of the central carina 
(which is testaceous) ; these however are sometimes confused or 
almost wanting ; punctuation of prothorax rugose, strongest be- 
hind shoulders ; tegmina fusco-hyaline, with the base more or less 
broadly fuscous and punctured, veins testaceous, underside fuscous, 
strongly pubescent in front; legs testaceous with the femora and 
tibiz more or less infuscate. Long. cum tegm., 9 mm. ; lat. int. 
hum., 3} mm. 

Hab. Guadaloupe (Delauney), (Belgian Museum Collec- 
tion); three female specimens. 

There is a larva of this species in the Belgian Museum 
Collection ; it is onisciform, broad and subparallel, but 
rather wider behind, with the front parts uneven, and the 


424 Rev. Canon Fowler on new species of Membracide, 


abdominal segments furnished at the sides with broad 
oblong processes, fringed with setz ; the colour is dirty 
testaceous, but has probably considerably faded. 


Paradarnoides ignipes, sp. n. 


Preecedenti affinis, sed minor, colore nigro,rmetopidio longiori, 
processu pronoti ad medium albido-marginato, prothorace dis- 
tinctius et minus rugose punctato, pedibusque miniatis facile 
distinguendus. 

Allied to the preceding, but smaller, with the shoulders set very 
far back so that the metopidium is only slightly declivous; of a 
black colour, with the margins of the pronotum narrowly edged with 
white behind shoulders, and the apex ferruginous; the eyes are 
very prominent, and the front part of the head is clothed with 
golden pubescence ; the pronotum is more closely and less rugosely 
punetured than in the preceding species; the tegmina have the 
base of the claws and the external margin of carina towards base 
black, and the veins large and fuscous ; the underside is black with 
the front parts strongly pubescent, and the legs are bright scarlet, 
with the exception of the coxe and tarsal claws, which are fuscous. 
Long. cum tegm., 7} mm. ; lat. int. hum., 3 mm. 


Hab. Guadeloupe (Delauney), (Belgian Museum Collec- 
tion) ; one male specimen. 

It is just possible that this may be the male of the 
preceding, but I do not think that this can be the case, 
as the insect differs considerably in the structure of the 
front part of the pronotum, which in P. ignipes is much 
longer from the shoulders to the eyes than in P. severini; 
in general appearance they are quite distinct. 


( 425 ) 


XV. Temperature Hxperiments in 1893 on several species 
of Vanessa and other Lepidoptera. By Fxreperic 
MerririEbp, F.E.S. 


[Read March 14th, 1894.] 
PuaTE IX, 


In my last paper (Trans. Ent. Soc. Lond. 1893, p. 57) I 
described some experiments on pupex of P. napi, off- 
spring of the spring emergence, and mentioned that I 
had some pup offspring of the summer emergence. 
The parents consisted of two males and two females, 
taken at Hailsham, and kindly sent to me by Mr. Vine 
on the 30th July, and two females taken at Petworth on 
the 5th August, and kindly given to me by Mr. Fletcher. 
From these I had several hundred eggs, which were laid 
much more freely on cardamines than on cabbage, 
though the larva seemed to feed as willingly on the 
latter as on the former. 

Early in September nearly all pupated. Ten were 
placed at 90° for ten days, a period amply sufficient to 
have caused their emergence had they belonged to the 
earlier brood, but it produced no effect either on the date 
of their emergence, when afterwards placed out of doors, 
or on their markings or colouring. The rest were kept 
out of doors. Some were forced about the middle of 
February, and some more early in March, emerging in 
from eight to seventeen days. The difference in appear- 
ance between these and the rest, which emerged out of 
doors between the 20th April and 9th May (except one 
which emerged 6th June), is the same in kind (though 
somewhat less in degree) as the difference between 
those of the summer emergence, which were forced all 
_ through, and those of that emergence which were cooled 
for the greater portion of their pupal period, as 
described Trans. Ent. Soc. Lond. 1893, p. 57. 

Consequently I may apply to the winter pupating 
brood the general remark made there, as to the brood 
which pupates in the summer, viz., that a part but not 
all of the characteristic seasonal colouring of this species 
depends on the temperature to which the individual is 


TRANS. ENT. SOC. LOND, 1894,—PaRT III. (SEPT.) 25 


426 Mr. F. Merrifield on Temperature Heperiments 


subjected, adding that the spring emergence appears 
to be less sensitive than the summer emergence is to 
temperature. Neither of the broods experimented on 
by me has proved so sensitive as those operated on 
by Prof. Weismann, and described by him (“Studies in 
Heredity ”’). 

Pararge egeria. This insect has two well-known climatic 
forms, the light spots in the South Kuropean form 
having the bright ochreous colouring of P. megera, 
instead of the straw colour of the English var. 
(egerioides). But the experiments tried afford little or 
no reason for supposing that these differences in appear- 
ance are the direct result of temperature. 

I obtained, between the 25th May and the middle of 
June, more than one hundred pup, which were sub- 
jected to various temperatures from between 80° and 
90° down to 33° (for many weeks), with various trans- 
fers from the lower to the higher temperatures. 

The chief difference, in general appearance, is between 
Classes I. and II. (forced) on the one hand, and Classes 
IV.-XIV. (those at 56° and under). ‘The former have 
the light spots smaller and less clearly defined, and the 
dark ground colour considerably lighter, and im many 
cases freckled with small dark brown spots. Class III. 
(open air, at about 66°) are not quite so dark as Class 
TV., but much nearer to them than to the forced. 

Though the light spots in those forced are smaller 
than in the others, they are somewhat more numerous. 
Class VIII. (iced and then forced) have the ground 
colour dark, but the light spots as numerous as in those 
which were forced, and as large as are the spots in those 
at the lower temperatures. One of these indeed, which 
I exhibit, has an inner row of light spots or traces 
of them on the secondary wings in nearly all the 
interspaces, and on the underside a perfect submarginal 
chain of six light spots, pupilled with dark brown, ona 
light ground colour. 

A few examples I reared from eggs laid in August 
showed the same effects generally as those from the 
eggs laid in April and May, so that there does not 
appear to be any marked constitutional difference in this 
respect between the spring and summer emergences of 
this species; my experiments would, however, lead me 
to expect those which emerge in spring to be in general 


on Vanessa and other Lepidoptera.’ 427 


more vivid in their markings and colouring than those 
which emerge later, after a spell of hot weather. 

Cidaria silaceata. 'This is known to be very variable 
in its markings, and I was tempted to experiment on it 
by the remark of M. Guenée that the spring and summer 
broods appear to vary asin the Selenias, and by other 
observations (Ent. Record, i1., 297), to the effect that 
the summer brood is smaller, and the band across the 
forewings less broken. 

Mr. Nicholson kindly gave me some eggs early in 
April, from which I had pup, which were subjected to 
about the same variety of temperatures as were the 
pupe of P. egeria, above mentioned. 

The main difference is between those at or over 80° and 
the rest; the latter being more strongly marked than 
the former, the transverse band perhaps showing a 
shghtly greater tendency to be broken, and _ their 
light markings being of a rather more ochreous tint ; as 
a consequence, those at 80° or over have a duller and 
more uniform appearance than the others. 

But the most distinctive feature is in the size. 
Those at or over 80° are, as a class, smaller than 
the others. This species, therefore, must be added to 
those in which temperature, during the pupal period, 
affects the size of the imago. ‘The difference in colour- 
ing and marking is hardly as great as I should have 
expected, and seems scarcely so great as that some- 
times met with between the spring and summer natural 
emergences, though it is in the same direction. I 
intended to experiment on the winter pupating brood, 
but a brood which I had from the second emergence 
came out as a third emergence, and were nearly all 
spoiled before I discovered that they had emerged. 

Araschnia levana. Desiring to experiment with pup 
of the summer emergence (var. prorsa) for their whole 
pupal period, I obtained, through Mr. Edwards, from 
North Germany, a large number of pupz from which I 
had, towards the end of April, thirteen good pairs, which 
I placed over growing nettle, in headless casks, and 
fed on orange, etc. All circumstances, including abun- 
dant sunshine, appeared most favourable; but I only 
obtained thirty-two eggs, laid (I believe by a single 
parent) mostly in strings of from two to eight, pro- 
jecting from the undersides of the leaves. Only eleven 


428 Mr. F. Merrifield on Temperature Experiments 


hatched, beginning 15th May, and all of them pupated. 
Three were forced at 80°, producing in from six to 
seven days the characteristic black prorsa form; four 
others were, at from one to eight hours’ old, placed in 
the refrigerator on the 18th June, and remained there, 
at about 48°, till 80th or 31st August (seventy-three or 
seventy-four days), when, observing signs of emergence, 
I placed them in the room at about 65° to 70°, and there 
three of the four emerged in a day or two, the fourth not 
until thirty-two days more, 7.e., on the 2nd October. 

The remaining four were placed at 33° till 29th Sep- 
tember (eighty-four days), then moved to the refrigerator 
at about 48°, whence, after twenty days more, they were 
moved to the room, about 59°, three of them emerging 
respectively in ten, eighteen, and twenty days more 
(total 114-124 days). ‘The first of them was a cripple, 
and the fourth died. This left six of the eight subjected 
to a low temperature. These six all emerged in perfect 
condition, and were unmistakably of the true levana type ; 
two of those cooled, without having been iced, showing 
slight traces of the intermediate porima form, but the other 
four being of the pure levana type, and nearly as different 
in colouring from the three that were forced as one of 
the common fritillaries—say A. selene—is from a “ white 
admiral” (Z. sibilla). I exhibit examples of the two forms. 


I now proceed to describe some experiments on four 
species of the genus Vanessa. In reference to these I 
have had the great advantage of submitting the speci- 
mens experimented on in the manner described in this 
paper, to the careful examination of Dr. Dixey, whose 
paper on the phylogenetic significance of the wing- 
markings in certain genera of the Nymphalidx will be 
found in Trans. Ent. Soc. Lond. 1890, p. 89, and he has 
favoured me with valuable observations upon them, 
which I am permitted to append ; they are distinguished 
by being placed within brackets. The “series” and 
“spots 7” referred to in these observations are described 
in Dr. Dixey’s paper, and may be indicated generally as 
follows, V. urtice and V. polychloros being convenient 
examples for the purpose:—Four dark patches on the 
costa, continued in series more or less perfectly across 
the wings, distinguished by the Roman numerals I., II., 
III.,1V., the latter including the dark submarginal band. 


on Vanessa and other Lepidoptera. 429 


Four lighter areas, A, B, C, D, alternating with the dark 
ones, A being the innermost, and coming before I.; these 
also continued in series more or less across the wings. 
Vanessa polychloros. There were sent me on the 6th 
May, a brood of between 130 and 140 larvee found on 
sallow in the New Forest; they were about three- 
eighths of an inch in length. I found they would eat 
cherry and birch, but seemed to prefer willow, on which 
accordingly I placed them, at first in two, and after- 
wards in four, large sleeves. On the 27th May, one had 
begun to spin, and all were brought indoors and fed on 
cut willow. There were 138; by the 31st 127 had spun 
up, and the rest followed in a few days. Their pupz 
were subjected to temperatures ranging from 100° to 
32°, being in many cases transferred, after a time of 
varying length, from the lower to the higher tempera- 
yi vice versa. I proceed to give their treatment in 
etail :— 


Class I., forced at from 90° to 100° (Plate IX., fig. 1). Most of 
these died, but afew at 90°, or a few degrees lower, did well; forced 
as pupating larve or pupe under twelve hours, they emerged in seven 
days. A second lot of ten (Class II.) were put in a shady place 
out of doors, where the temperature averaged about 62°, and all 
emerged in from twenty to twenty-two days. A third lot 
(Class III.) were placed in a cellar at a temperature averaging 
about 56°, where all emerged on the fortieth day. A fourth lot 
(Class IV.) were placed in the refrigerator at about 48°, and, after 
periods ranging from fourteen to forty-six days, transferred to 
(a) the forcing box at 80°-90°, emerging in from three to five 
days more; (b) the cellar at 58°, emerging in from twelve to 
twenty-five days more ; or (c) the room, at 68°-75°, where they 
emerged in from four to five days more. A fifth lot (Class V.) 
were iced for periods ranging from fourteen to forty-two days, and 
then (a) placed at 80°-90°, emerging in five to seven days more ; 
(b) in the cellar at about 59°, emerging in twenty-seven to thirty 
days more ; or (c) the refrigerator at about 49°, for from six to 
thirty-two days, and then, either the cellar at 58°, emerging in from 
twenty-two to twenty-three days more, or the room at about 
68°-75°, emerging in two to twenty-eight days more. With the 
exception of a few that were injured by accidents, of those that 
were killed by excessive heat, as mentioned before, and of four or 
five among those longest iced, all emerged, and except among some 
of those iced, there were no cripples. 


430 Mr. F. Merrifield on Temperature Hxpervments 


The effect on colouring was as follows :— 

Class I. (forced at about 90°, or upwards), emerging in seven 
days). The ground colour of a lighter and yellower hue of brown 
than is normal, with many yellowish clouds and broad streaks, 
especially in the interspaces of the nervures on the outer half of 
the forewings. [Black spots generally are more sharply defined 
than in normal specimens. ‘There are no blue submarginal cres- 
* cents in the forewings, but many bluish scales on the extreme 
margin. The spots in “Series D” are particularly well-defined 
near the costa, and are not pupilled. | 

Class IT. (shade, out of doors, at 519-69°, averaging about 62°, 
emerging in twenty to twenty-two days). Ground colour of a 
redder brown, and with few yellowish clouds ; the yellow sub- 
marginal outer line is especially reduced, and the dark band inside 
it widened and darkened. 

Class ITT. (cellar at 54°, rising to 58°, averaging 56°, emerging 
in forty days). Effects intensified ; the yellow submarginal line 
has almost disappeared, and there is a scattering of dark spots on 
the ground colour, in some cases forming a streak in front of the 
inner edge of the forewings. [There are indications of blue sub- 
marginal crescents in the forewings, but less blue in the fringe or 
extreme margin than in I. The spots in “Series D” sometimes 
bear minute black pupils. ] 

Class IV.a (refrigerator, about 49°, fourteen to forty-six days ; 
then forced at 80°-90°, emerging in five days). Effect rather a 
mixture of those in Classes I. and III.; the scattering of dark 
spots exists, but the yellowish clouds and yellow submarginal 
streaks are partially restored ; in those exposed to the low tempe- 
rature for forty-two days, there were several that died or were 
crippled, and the dark markings in some others are varied with a 
paler hue, giving rather a “greasy” appearance to these dark parts. 
[The spots of “Series D” often with minute black pupils ; 
‘Series C” is indicated in the hindwings by a row of black points; 
a new dark spot tends to be formed between “II.8” and “TITI.8.”] 

Class IV.b (refrigerator at about 49°, fourteen to forty-six days; 
then cellar at 56°-60°, emerging in twenty-five to twelve days, or 
room 65°-75°, in five to four days). Much like Class IIT., except that 
the ground colour is duller, and the submarginal blue tends to be 
supplanted by black ; in those longest exposed to cold, the dark 
parts tend to spread. [Tendency to formation of new dark spots 
continues, | 

Class V.a (iced at 33°, fourteen to thirty-eight days, then forced 
at 80°-90°, emerging in seven to five days). These, unless iced 


ow Vanessa and other Lepidoptera. 431 


twenty-nine days or more, are very like IV.a ; iced for that ora 
longer period, they are darker; in all cases they show a return of 
the yellowish markings. 

Class V.b (iced at 33°, fourteen to forty-two days, then at various 
temperatures, such as cellar at 59°, emerging in twenty-seven to thirty 
days ; or refrigerator at about 49°, six to thirty-two days, and then 
in cellar or room, emerging in twenty-one to two days more). These 
are classed together, because the effects seem to depend on the 
duration of the icing. No great effect is produced under twenty- 
nine days’ icing ; the extreme darkness, often without crippling, is 
produced by icing thirty-six days, followed by the refrigerator 
at 49°, for six to nine days (Plate IX., fig. 3); but some taken 
straight from the ice to the cellar are nearly as dark. Of those 
iced from thirty-eight to forty-two days nearly all died, or were 
more or less crippled ; one of the latter has nearly all the four spots 
on the forewings obliterated ; and it may be observed that the 
icing for thirty-six days or more, followed by the refrigerator, 
which produces the extreme dark effect, has a tendency to cause 
the normal spot near the inner edge, which is nearest the base of 
the forewing, to disappear. 

As regards the general appearance of those which show the 
extreme effect of the low temperature, it may be said that they 
much resemble V. xanthomelas. [Tendency to formation of new 
dark spots continues. An additional dark spot may also appear in 
cell (forewing) below “I” and “II.” The border may become 
uniformly dark. ] 

A second company of V. polychloros, just changing their last 
skin, reached me on the 2nd June, also on sallow. Sleeved on 
cherry they did very well. Some were forced, emerging in six and 
a half to seven days. Others, placed out of doors at a temperature 
ranging from 67° to 59°, averaging about 64°, emerged in sixteen 
to seventeen days. This was a rather lighter coloured company 
of butterflies. There is the same kind of difference in appearance 
between the forced and the others, as there is between Classes I. 
and II. of the first company, but it is less in quantity. 


V. polychloros, general conclusions. The colouring is 
considerably affected by temperature in the pupal stage, 
low temperatures producing a deepening of the ground 
colour and an extension of the dark markings; and high 
temperatures producing a lightening of the ground colour 
and an extension of the yellowish markings. The blue 
and bluish markings are strongest in those at moderately 


432 Mr. I. Merrifield on Temperature Hxperiments 


low temperatures, Classes III. and IV., in many of which 
they form some rather bright crescents on the forewings ; 
but at the extremely low temperatures they tend to be 
supplanted, in some cases entirely so, by black. 

[Forcing invariably tends to produce yellow, whether 
pupa previously warmed or cooled. Refrigeration pro- 
duces increased breadth of dark brown, whether followed 
or not by forcing.|_ Imay add that among the specimens 
I exhibit, one belonging to Class LV.a (refrigerator thirty- 
eight days, then cellar four days, and forced three days) 
(Plate IX., fig. 2), to which my attention has been directed 
by Dr. Dixey’s observations, is particularly interesting, 
showing “ Series D” as a nearly complete chain of faint 
yellowish spots, or rather clouds, on both forewings and 
hindwings, the anterior three or four on the forewings, 
and all those on the hindwings having each a small black 
point in the centre. It seems as if it required cold, 
succeeded by heat, to cause this chain of yellowish spots 
centred with black to be brought out. 

Nothing has been said about the colouring of the 
under sides. ‘This varies moderately in darkness or 
lightness, but I have not been able to associate this 
variation definitely with temperature. 

Vanessa atalanta. In looking carefully, and aided by 
a strong light, at the V. atalanta upon which experiments 
were made in 1892, as recorded, Trans. Ent. Soc. Lond. 
1893, pp. 58-62, I noticed a feature which had escaped me 
before, viz., that eight out of the ten which were subjected 
to the high temperature of 80°-90° had a few dull orange 
scales on the upper side of the forewings, between the 
large white costal blotch and the row of smaller white 
spots nearer the hind margin. 

I determined to develop this tendency by exposing 
some pup to a greater heat. The effect was a great 
development of this orange colour, both in intensity, it 
becoming distinctly scarlet, and in quantity, so as to form 
a scarlet cloudy patch sufficient to attract attention on 
casual observation. I exhibit several examples. This 
patch is between the large white costal patch and the 
third of the row of white spots beyond, and tends to form 
a scattered ring around this third spot; other scales, from 
golden brown to scarlet, are developed along the outer 
part of the costa and near the base, and elsewhere on the 
forewing. 


on Vanessa and other Lepidoptera. 433 


The following is a description of the treatment to 
which the pupe were subjected, and of its results :— 


A considerable number of pup were placed at a temperature of 
about 100°, at which nearly all died, after progressing so far as to 
show their imaginal colouriug. I then lowered the temperature to 
about 90°-95°, with the result that the great majority of them 
show these markings, the scales being scarlet instead of dull orange, 
and in several cases being so increased in quantity as to form a 
scarlet clouding sufficient to attract attention when the insect is 
held at arm’s length and more. These scarlet scales follow the 
nervure which separates the second from the third of the row of 
white spots above referred to; they are not on the nervure, but on 
each side of it, and in some cases extend to the nervure next below, 
and have a tendency to form a scattered ring round the third of the 
white spots. These scarlet scales are also to be found on the costa, 
extending in some cases from the beginning of the large white 
costal patch almost to the apex of the wing (an ochreous colouring 
in this region is to be observed in captured specimens). Asso- 
ciated with these markings is an increase of the brightness and 
warmth of the golden-brown colouring of the costa and nervures 
in the basal part of the wing ; in some crippled specimens this 
golden-brown is very vivid, and makes some approach in hue 
towards the scarlet band across the wing ; in some of these also a 
patch of the scarlet scales is to be found between the middle of the 
scarlet band and the large white costal spot. 

In my paper on the experiments of 1892, I mentioned that on 
the underside of two out of ten specimens at from 80° to 90°, a 
new small scarlet spot appeared between the scarlet band across the 
forewings and the inner edge. In only six individuals out of those 
subjected to a high temperature in 1893 do I find scarlet in this 
region. I find it occurs in two places, both below the median 
nervure, viz., (a) just before it forks, (4) a little below the lower 
branch of the same nervure. One specimen (No. 4) shows both 
(a) and (b); three (Nos. 21, 39, and 49) show (a) only, and 
two of them but slightly; and two (Nos. 48 and 188) show 
(b) only. 


The scarlet scales on the upper side are found in the great 
majority of those (about thirty) that were subjected to a tempe- 
rature of 90°, or upwards, during their whole pupal period, and in 
three out of thirty-three that were subjected to 80°-100° during 
the earlier part of their pupal period ; they are not found in any 
of the forty or so that were subjected to lower temperatures. The 


434 Mr. F. Merrifield on Temperature Hxperiments 


scarlet patches on the under side are only found in six out 
of forty-eight which were subjected to 99° or upwards, and five 
out of the six were so exposed for practically their whole pupal 
period. 


In the opposite direction of low temperature, I tried 
further experiments with V. atalanta, the most marked 
results of which I exhibited at the meeting of the Ento- 
mological Society on the 8th November, 1895. ‘They con- 
firm, and in some cases carry further, the results obtained 
in the previous year. The low temperature causes much 
substitution of white, lavender, or metallic blue-green 
scales (one of these colours seeming readily in this insect 
to pass into another of them) for the black in normal 
specimens ; the large white spot on the costa is greatly 
enlarged and spread, and the tendency of the third of the 
row of submarginal spots to ocellation which is above 
referred to, as caused by a high temperature, is shown in 
a different manner by the low temperature, which tends 
to form a whitish ring round, and very near to it 
(this third spot is on the underside ocellated in normal 
specimens), 

The extreme low temperature forms are, on the 
whole, so decidedly smaller than the average, that I am 
inclined to think the low temperature is a cause of 
reduced size in this species, more especially as those at 
the high temperatures, even where this is so extreme as 
to kill some of them, are all of full size. 


[ Yorced.—Resolution of inner margin of red band fairly 
marked. 

Refrigerated.—Marginal blue much extended, especially 
about anal angle of hindwing and in centres of spots of 
“Series IV.” ; spots of “Series D”’ often ringed with 
pale blue, apart from ocellation; indications present of 
bluish centres to ‘ Series III.” in hindwings; greater 
general blackness. | 


Mr. J. J. Walker has kindly given me a specimen from 
Gibraltar which resembles, in its main features, some of 
my earliest individuals. It was taken, recently emerged, 
on the 17th February, and I find that the mean tem- 
perature of January and February at Gibraltar may be as 
low as 48°7° and 50°9°, Dr. Chapman has kindly sent 


on Vanessa and other Lepidoptera. 435 


me some atalanta of a very late brood reared last 
October and November at Hereford, which also present 
some of the characteristic appearances of my cooled 
specimens. 


Vanessa (Grapta) c-album. Owing to the kindness of 
Mrs. Hutchinson, who sent me some eggs laid by hiber- 
nated butterflies in the spring, and of Mr. Nesbitt, of 
Llandogo, who sent me larvz of the second brood at the 
end of July, I have been able to ascertain that while both 
broods are affected by temperature in the pupal stage, the 
first brood is much the more sensitive of the two. It is 
remarkable that there should be so great a difference in 
constitution between these two broods, as, under natural 
conditions, the pupz of both broods are exposed to tem- 
peratures differing by a very few degrees, the one passing 
the pupal stage in England about June, and the other 
about August. It is entirely consistent, however, with 
Mr. W. H. Edwards’ experiments and observations on 
the two closely-allied American species, Grapta inter- 
rogationis and Grapta comma, as described in the 
“Canadian Naturalist’? for 1877 and 1878, and much 
light is thrown on the subject by Prof. Weismann’s 
observations on Mr. Edwards’ experiments, in the Pro- 
fessor’s “‘ Studies in Heredity,” by Prof. Meldola, vol. i., 
p. 149. The individuals experimented on, eight of the 
first brood and eleven of the second brood, were not 
sufficiently numerous to justify me in describing the 
results in detail; but I hope to try experiments on a 
much larger scale during the present year. 

Vanessa to. Mr. Morris, of Lewes, kindly gave me a 
company of about one hundred and twenty larvee, all in 
their last skins, or nearly so, on the 15th June, which in 
the extraordinarily early season of 1893 was late for 
them. I was much occupied in other ways, and perhaps 
it was owing to insufficient attention that I did not 
obtain more than about sixty pupz, and those were a 
little under the full size. They were subjected to various 
temperatures from about 100° downwards. ‘Those at 
100° all failed to emerge. Sixteen, which were at $0° 
for six days, and then at 80°, all emerged in one day 
more, making seven days. As the temperature was 
lowered, there appeared a gradual tendency to disinte- 
gration of the ocellus on the forewing, until in one 


436 Mr. F. Merrifield ov Temperature Experiments 


(Plate IX., fig. 4), iced at 33° for twenty-two days, then in 
the refrigerator for twenty days, and then in the cellar for 
eighteen days, it ceases to be an ocellus, being resolved 
into a chain of small white spots, which are bright, with 
only a very slight bluish shade about them, and affording 
a remarkable confirmation of Dr. Dixey’s views of the 
origin of that ocellus, as exemplified in the plate attached 
to his paper in the Transactions for 1890. In these 
iced and cooled specimens the blue becomes more vivid, 
and a narrow dusky marginal band, slightly darker in hue 
than the chestnut brown ground colour, appears, with a 
submarginal incomplete row of small dusky spots, very 
distinct. On the hindwing there is little change, but a 
tendency to disintegration of the blue in the ocellus. 

Dr. Chapman kindly sent me, on the 30th August, 
part of a second brood he had found rather more than 
half grown. From about forty larve I obtained thirty- 
eight fine and healthy pupz, but nearly all were killed 
by the severe cold to which I subjected them, though 
only a little in excess of that to which No. 6L was 
exposed. 

[I. Forced: a tendency shown to the development of 
dark spots at the apices of the interspaces (‘Series IT.”’); 
tendency towards fusion of bluish constituents of ocellus 
in hindwing. II. Cooled: tendency of “IV.” (marginal 
chain) to separate from “D” (light apical spots) ; 
“JV.” rendered more distinct in forewing. III. Iced: 
separation of ‘“D” and “IV.” asin II. ‘ Claw-mark ” 
tends to lose regular curve, and to become angulated. 
Bluish constituents of ocellus in hindwing tend to be- 
come separated into two parallel series—“III.” and 
“TV.,” @.e., a marginal and submarginal. | 

V. antiopa. About seventy pupx, mostly rather 
fresh, were obtained for me from near Berlin by Mr. 
Edwards, on the 19th July, and were subjected to 
various temperatures from about 100°, emerging in from 
three to five days, and 80°, when they took a day or two 
more, downwards. ‘The most severe temperature sur- 
vived, without injury, was twenty-seven days in the 
refrigerator, at about 47°. All that were placed in ice 
(33°) for twenty days or upwards died, except one that 
was a cripple. The results are negative, as none show 
any marked differences in marking or colouring that can 
be assigned to temperature. The absence of positive 


on Vanessa and other Lepidoptera. 437 


results is very probably owing to the circumstance that 
the pupe were all of them several days old when they 
reached me. 


The experiments now recorded confirm in general the 
conclusions drawn from such as have preceded them, and 
some of which may be briefly enumerated as follows :— 
(1) The effects of temperature are different when applied 
at different periods of the pupal stage. (2) A great range 
of temperature may cause but little difference in appear- 
ance, while a very few degrees near the top or bottom of 
the range the imsect will bear may cause a great 
difference. (3) There may be a great constitutional 
difference in sensitiveness to temperature between two 
seasonal emergences of the same species. (4) This may 
be so even when both pass the pupal period at about the 
same temperature (this is in accordance with Mr. W. H. 
Edwards’ observations above referred to). (5) While 
some kinds of effect seem to be what may be called the 
direct result of temperature, in others, and perhaps the 
most important, temperature appears to operate by 
causing the individual to “throw back” to some 
ancestral form; this last circumstance has been con- 
sidered to explain the reason why a low temperature in 
some species causes darkening of the colours, and in 
other species produces the opposite effect. (6) In these 
cases of “‘reversion,” the kind of effect produced appears 
to depend on the stimulus applied, low temperatures 
producing one class of effects and high temperatures 
a different class of effects. 

The whole subject is one of much complication, and 
calls for further experiments in many directions. The 
direction which mine have taken, following in the lines 
initiated by Weismann and W. H. Hdwards, especially 
if pursued with species belonging to regions where the 
seasonal or other occasional differences of temperature 
are extreme—North America, Siberia, Japan, or the 
vicinity of mountains—will help to trace, and separate 
from the rest, such of the causes of variation as depend, 
directly or indirectly, on temperature. Systematic 
experiments on a number of well-selected species 
belonging to countries where the seasonal difference is 
hygrometric rather than thermometric, would probably 
produce valuable results. The nature of the food-plant, 


438 Mr. F. Merrifield on Temperature Experiments. 


which undoubtedly influences size and vigour, and is 
generally considered also to influence markings and 
colour, offers another line in which experiments of a 
more systematic and comprehensive character than any 
yet tried would undoubtedly well repay the labour 
attendant upon them. ‘There are other natural sur- 
roundings, most potent for many purposes, the effect 
of which might usefully be tested by experiment, such 
as light and electricity and magnetism. As to light, I 
tried, in 1891, some experiments on B. cynthia and 
S. illustraria, recorded in the Trans. Ent. Soc. for 1892, 
p. 42; but, so far, with negative results. And in 1891. 
I tried some experiments with strong magnetic currents 
on some Lepidoptera in all their stages; but these 
yielded no positive result. 

Norre.—As this paper and Dr. Dixey’s, which succeeds it, con- 
tain observations on some of the same facts by two different and 
independent observers, there is necessarily some repetition, but in 
order to reduce this as much as possible, I have greatly condensed 
my own observations. 


EXpLANATION OF Prate IX. 


Fic. 1, Vanessa polychloros: pupa at 90°-95°, emerging in seven 
days. 

Fic. 2. V. polychloros: pupa about 49°, for thirty-eight days, then 
about 58° for four days, then about 85°, emerging in 
three days more. 

Fic. 3. V. polychloros: pupa at 33° for thirty-six days, then about 
49° for nine days, then about 58°, emerging in thirteen 
days more. 

Tia. 4. V. io: pupa at 33° for twenty-two days, then about 49° 
for twenty days, then about 60°, emerging in eighteen 
days more. 


( 439 ) 


XVI. Mr. Merrifield’s Experiments in Temperature- 
Variation as bearing on Theories of Heredity. By 
Freperick A. Drxey, M.A., M.D., F.E.S., Fellow of 
Wadham College, Oxford. 


[Read March 14th, 1894.] 


Tue results of Mr. Merrifield’s experiments on the varia- 
tions produced in butterflies by the exposure of the pupa 
to different conditions of temperature, are in themselves 
of great interest. But the interest becomes enhanced 
when it is recognized that many of the new features 
which make their appearance under these conditions are 
identical with those occurring normally in other species 
more or less closely allied to the subjects of experiment ; 
that in not a few instances the disturbance of natural 
temperature-conditions appears to have caused reversion 
to an earlier stage in the phylogenetic history of the 
species; and further, that the ancestral.features thus 
revived seem to vary with the nature of the disturbance. 

Examples of these phenomena, from a previous series 
of experiments, were given in ‘Trans. Ent. Soc. Lond. 
1898, p. 55, and were commented on by me (Ibid. p. 69). 
The latest results obtained by Mr. Merrifield, besides 
confirming many of the former, furnish further instances 
of the same nature, as follows :— 


I. VANESSA ATALANTA. 
A. Warmed. 


(1) The occurrence of red scales in the dark ground- 
colour between the middle of the scarlet band and the 
large white costal spot c. This is an approach to the 
condition in V. huntera and V. myrinna, and more 
remotely to that in Grapta and Argynnis. A correspond- 
ing feature is seer in V. to, which in this respect is 
more ancestral than V. atalanta. 

(2) The tendency towards the formation of a scattered 
ring of red scales round the spots 8 and ¥y of Series D. 

TRANS. ENT. SOC. LOND. 1894,—parT III. (SHPT.) 


440 Dr, F. A. Dixey on Mr. Merrifield’s Experiments 


This again recalls a common condition in V. myrinna and 
V. huntera. 

(3) The appearance of a new red spot on the under- 
side of the forewing, just below the stem of the median 
nervure before bifurcation. ‘This represents a pale patch 
of various shades in V. callirrhoe, V. myrinna, V. hwntera, 
V. cardwi, etc. 

(4) The appearance of another red spot on the under- 
side of the forewing, just below the first median nervule. 
This represents a patch visible on both surfaces of V. 
callirrhoe, and fully developed in all the species nearly 
allied to V. carduwt. 

(5) The tendency towards resolution of the inner 
margin of the red band, as in V. callirrhoe. 

(6) The suffusion of the dark ground colour with 
golden brown, also as in JV. callirrhee. 


Of these, Nos. (1) (2) and (3) are points now 
observed for the first time; while (4) (5) and (6) are 
confirmations of previous results. (See a former paper 
by the author in Trans. Ent. Soc. Lond. 1893, p. 70.) 


B. Cooled. 


(1) Much substitution of lavender or metallic blue- 
green scales for black. This points to the ancestral 
condition seen in the females and parts of the males in 
many species of Argynnis, e.g., A. paphia, var. valesina, 
A. sagana ¢, A. niphe 3 and ¢, and A. diana ¢. 

(2) The presence of minute patches of bluish scales 
near the margin of the dark ground-colour in the hind- 
wing, indicating the blue centres of the almost com- 
pletely merged Series I11.—an ancient feature of Vanessa 
and Grapta. 

(3) The marked increase of marginal blue, especially 
about the anal angle of the hindwing. This appears to 
represent the condition seen in Argynnis niphe, and 
ultimately to point back to the primitive Argynnid 
colouring of A. valesina and A. diana &. 

These are all confirmations of former results. 


II. VANgEssa 10. 


In this species, warming tends to revive, in the fore- 
wing, a series of dark spots (ID), occurring normally in 


in Temperature- Variation. 44] 


Araschnia levana. Cooling tends in the first place to 
separate certain constituents of the ocellus, and when 
carried to a high extent has the remarkable effect of 
causing an unmistakable resolution of the ocellus in the 
forewing, the appearance finally produced being that of 
the ordinary Vanessa character in a comparatively un- 
modified form. It is interesting to see how completely 
these cooled specimens bear out the views which I ven- 
tured to express, some years ago, on the origin and 
constitution of this ocellus (Trans. Ent. Soc. Lond., 
1890, pp. 99, 100, pl. i, fig. 12). The ocellus of the 
hindwing is also affected in the same direction. 


III. VaNnEssA POLYCHLOROS. 


Cooling tends to produce several features which 
appear to be ancestral. The chief of these are (1) the 
pupilling with black of the spots of Series D in the fore- 
wing; (2) the occasional indication of Series III. in the 
hindwing; and (3) the tendency towards the formation 
of a new dark spot between II.8 and III.8. These 
points approximate towards the condition in Grapta. 


TV. GRapra C-ALBUM. 


In both broods cooling tends to induce or increase a 
darkness of ground-colour; this being undoubtedly an 
ancestral character.* 


In all cases of this kind the obvious question occurs— 
are we to consider these phenomena as true instances of 
reversion, or is it merely that like causes have produced 
de novo a like effect in descendant and ancestor? ‘I'he 
latter explanation may account for some of the facts, but, 
I think, not for all. It may perhaps give the reason for 
a general diffusion of bluish scales, or for a change of the 
ground-colour from black to brown, but it is scarcely 
adequate to explain the special formation of a definite 
pattern, as of Series III. with its blue centres in JV. 
atalanta, or the reduction of the ocellus in V. 1o to the 
primitive Vanessa condition. Without raising the vexed 
question of sexual selection, we may yet affirm that 


* The observations on the three last species are new ; those on 
V. atalanta, as has been seen, are partly new and partly old. On 
the whole subject of the ancestral markings in Argynnis and 
Vanessa, see the author’s paper in Trans. Ent. Soc. Lond., 1890. 


TRANS. ENT. Soc. LOND. 1894,.—parv 1. (SEpr.) 2 F 


412 Dr. F. A. Dixey on Mr. Merrifield’s Experiments 


among the features induced or revived by altered tem- 
perature-conditions, there is at least a residuum which 
must have owed its first origin to causes other than the 
direct action of temperature on the organism. Nor, 
again, are these to be considered as cases of “arrested 
development”; for the stages reproduced are stages in 
the phylogeny of the species, not in the ontogeny of the 
individual. 

If, then, these revived features are really ancestral, - 
how is their revival to be accounted for? The whole 
subject of reversion abounds with difficulty. An expla- 
nation commonly offered is that the characters last 
developed in the history of a species, or of an individual, | 
are less stable than those that have a longer history 
behind them, and that have become firmly established, 
under the operation of a long-continued process of 
heredity. Any disturbance—such as an exceptional con- 
dition of temperature—of the normal course of growth, 
may therefore be expected to act in the first place on the 
newer and less stable features, interfering with their 
usual line of development, and shaking back the species 
as it were to an earlier and more firmly-founded stage 
of its development—just as in an earthquake the freshly- 
built wing of a house, where the mortar was not yet dry, 
might fall and leave the older portions standing. Such 
an explanation, however, is in itself at best but partial, 
for it gives no real reason why the newer features should 
be less stable than the old; and indeed it comes to little 
more than restating the difficulty in another form. 

The two attempts to find a more definite explanation 
of reversion which may be said at present to hold the 
field, are those which pass respectively under the names 
of Darwin and Weismann. If the Darwinian assumption 
of centripetal “ gemmules” be granted, the commonest 
case of reversion, that namely which results from hybridi- 
zation, especially between recently-established species, is 
capable of explanation under the hypothesis of pan- 
genesis. But it may be questioned whether pangenesis 
as stated by Darwin is capable of accounting for such 
cases as the present, inasmuch as in them the condition of 
full maturity is almost reached before the introduction of 
the modifying disturbance. Although the ovum from which 
the individual has originated may under the Darwinian 
hypothesis have contained numerous gemmules of an 


in Temperature-Variation. 443 


ancestral type, which though usually dormant might 
under certain circumstances become active in the onto- 
genetic process, it would yet seem a legitimate conclusion 
from the hypothesis, that the introduction of any cause 
analogous to hybridization in its action on the developing 
organism must belong to a far earlier stage in the onto- 
geny than the beginning of the pupal condition ; it must 
belong, in fact, to the stage of fertilization of the ovum. 
There are, however, a few facts on record, such as the 
assumption of ancestral characters by an old hen (Darwin, 
« Animals and Plants under Domestication,” 1868, vol. ii., 
p. 54), and the appearance of an earlier vertebrate con- 
dition in limbs of Amphibia reproduced after amputation 
(Ibid., 11., p. 15), which seem in some respects analogous 
to the present instances, as being apparently cases in which 
a disturbance of normal conditions at a comparatively 
late ontogenetic stage has in some way led to reversion 
in the course of the individual growth. These cases are 
regarded by Darwin as not incompatible with pangenesis, 
though not fully explained by it. 

Tf, on the other hand, we postulate with Weismann the 
existence of “ids”? and ‘‘ determinants,’”? endowed with 
the nature and properties that he supposes, the instances 
that we are considering become more explicable. For 
according to this theory every feature in the structure of 
the individual organism is the result of a “ struggle of the 
ids” in ontogeny, the final character of each histological 
unit being fixed at the moment of the liberation of its 
proper determinants by the disintegration of the “ids.” 
The competition between the carriers of heredity, many of 
which must under the theory be ancestral in character, so 
far from being confined to the ovum, is being waged 
throughout the entire ontogeny, and is renewed at every 
successive stage of development. This being the case, it 
is to be expected that any external influence, such as 
temperature, on coming into force at any given stage, 
should be able to exert an effect upon the struggle pro- 
ceeding at that particular time between determinants 
which are just beginning to play their parts in the onto- 
geny, and should in consequence be able to modify pro 
tanto the resulting adult organism. It would be, more- 
over, natural to expect the different determinants to be 
affected by different temperatures, nor would it be sur- 
prising to find that temperature-conditions, which are 


444 Dr. F. A. Dixey on Mr. Merrifield’s Experiments 


ex hypothesi diverse from those normal to the species, 
should favour one or other set of ancestral determinants 
at the expense of those more proper to the species. ‘This 
would explain why the effect of heat differs from that of 
cold, though both lead to reversion. 

There is, however, one fact which shows that the above 
explanation is not entirely adequate—the faci, namely, of 
the hereditary transmissibility of certain temperature 
modifications, as determined in the case of Polyommatus 
phleas by Weismann himself (“ ‘The Germ-Plasm,” 1893, 
p. 399). This phenomenon admits of a ready explanation 
under the theory of pangenesis ; the point that pangenesis 
fails to explain is the reversionary character of the original 
change, unless, indeed, we suppose a ‘‘ struggle of gem- 
mules,” analogous to the “ struggle of determinants,” and 
continued, like the latter, throughout the ontogeny; in 
which struggle certain conditions favour the ancestral 
rather than the modern gemmules. But just as the 
theory of pangenesis seems to require some such addition 
as that suggested, so also, under the rival hypothesis, it 
seems necessary to supplement the explanation above 
given with another supposition already propounded by 
Weismann, namely, that the temperature-conditions are 
capable, in some cases, of actually altering the consti- 
tution of unexhausted determinants wherever they occur, 
even in the germ plasm of the ovum itself. 

Jam myself inclined to think that, granting Weismann’s 
general theory of heredity, the more special cases of 
reversion are to be chiefly explained, as above, by tie 
critical influence of the temperature-conditions on the 
struggle of the determinants, rather than by an intrinsic 
effect on the determinants themselves. The latter may 
account for such cases as a general lightening or darken- 
ing of the ground-colour, as in Weismann’s P. phleas, 
which strictly speaking are not really but only acci- 
dentally reversionary ; it will not, however, account in 
my opinion for the special ancestral marks shown by 
Mr. Merrifield’s V. atalanta and J. io. 

The point is capable of verification. If it be true that 
there is a selective influence which is exerted upon the 
actual struggle of the determinants, that influence would 
find a different expression in the adult according to the 
particular stage in the ontogeny at which the influence 
was applied, as it would affect those determinants only 


in Temperature-Variation. AAS 


between which at that time the struggle was being 
waged,* If, on the other hand, there is no such influ- 
ence, but the effect is entirely a direct one and modifies 
the individual determinant, then all the as yet unex- 
hausted determinants that are capable of reacting to this 
particular disturbance would be affected in some degree ; 
though no doubt, as Weismann supposes, to a greater 
extent if they had reached the point of disintegration than 
otherwise. 

Again, it seems to me to be of great importance to 
ascertain if possible which of these modifications are 
transmissible to descendants. If all the modifications, 
including those which I have supposed to be produced 
in the first way, can be shown to be hereditarily trans- 
missible, this would amount to a demonstration that the 
second explanation is adequate ; and the first may then 
be abandoned as unnecessary. Should only some be 
inheritable, the presumption would be in favour of the 
co-existence of both modes of action; moreover, the 
greater the number of non-transmissible variations that 
can be produced, the more will the case be strengthened 
against pangenesis, and in favour of the “ centrifugal” 
theory. . 

I am anxious to see, if possible, the results of breeding 
experiments on specimens like these for yet another 
reason. It seems to me that by comparative experi- 
ments, with and without artificial selection, on such 
variations as may be transmissible, a measure might be 
obtained of the relative importance of selection and the 
mere action of external influences in the transformation 
of a species. I think, too, that no better group for such 
experiments as these of Mr. Merrifield’s could be selected 
than the Vanessas. For, in the first place, it is only 
among poikilothermic animals that the direct effects of 
temperature can be fully studied ; then among these the 
Lepidoptera are pre-eminent for the extremely delicate 
register of variation afforded by their wings ; and, lastly, 


* The fact that in V. polychloros forcing invariably tends to pro- 
duce a certain effect, whether preceded by warming or cooling ; 
while refrigeration brings about another definite effect, whether 
followed or not by forcing, seems so far favourable to this hypo- 
thesis. See above, p. 432. 


446 Dr. F. A. Dixey on Temperature-Variation. 


among the Lepidoptera the Vanessas belong to an 
assemblage the phylogeny of which may claim to be at 
any rate partially known.* It is hardly necessary to point 
out how much service may be rendered to researches of 
this kind by the careful working out of the true internal 
affinities of Lepidopterous groups. In proportion as their 
phylogeny is placed on a secure basis, we shall be able to 
pronounce with confidence on the real character, whether 
reversionary or not, of these remarkable variations; and 
shall accordingly be able to estimate at its proper value 
the evidence they bring towards the solution of the great 


problem of Heredity. 


* See the author’s paper already referred to, in Trans. Ent. 
Soc. Lond., 1890, p. 89. 


XVII. Description of new Cicindelidee from Mashunaland. 
By Louis Pirinevey, F.E.S. 


[Read March 28th, 1894.] 


In my ‘Catalogue of the South African Cicindelidee” 
(Trans. South Afric. Phil. Soc. vii, 1893), I expressed, 
as my opinion, that this catalogue could not be regarded 
as final, as Gazaland and Ovampoland would, in all 
probability, yield new forms of the wingless genera, 
Myrmecoptera, Dromica, and Cosmena; but I was not 
prepared to receive, a few months later, no less than five 
Myrmecoptera and one Cosmema, hitherto undescribed, 
and all found in one locality of the newly-opened-up 
northern territories of South Africa, which will be known 
hereafter as Zambezia. 

All these species were captured round Salisbury at 
the beginning of the rainy season, by my esteemed cor- 
respondent, G. A. K. Marshall, Hsq., who has gone lately 
to reside in Mashunaland. Matabeleland and Mashunaland 
would seem to be the home of Myrmecoptera, for besides 
the five new species here described, Mr. Marshall has 
also captured the hitherto extremely rare M. polyhir- 
moides, Bates, and M. bilunata, Dohrn. Mr. F. HW. 
Selous, while recruiting at Buluwayo from the effects of 
a wound received in the Matabele war, captured there 
the equally rare M. mauchi, Bates, and M. limpopoiana, 
Pér. These two species were very abundant, [ am in- 
formed by Mr. Selous, who, however, captured only one 
example of each species, as he had no means of storing 
them. Mr. Marshall also writes that M. ivicta and 
some of the other species were fairly common. I have 
now recorded twelve species of Myrmecoptera from 
Matabeleland and Mashunaland, and two Cosmema. 

The other Cicindelide captured near Salisbury by Mr. 
Marshall were Megacepvhala regalis,* Cicindela clathrata, 


* Occurs also near Buluwayo. 
TRANS. ENT. SOC, LOND. 1894.—part 111. (SEPT.) 


448 M. Louis Péringuey on descriptions of 


C. disjuncta, Jansenia angusticollis, and Cosmema lepida, 
He had not met with any Mantichora as yet. 


Gen. CictnpeLa, Linn. 
C. mashuna, sp. n. 


Supra obscure cuprea, subnitida, labro flavo utrinque bipunc- 
tato ; prothorace quadrato, subtiliter granulato, albido piloso ; 
elytris granulosis, margine sat lata, utrinque breviter tri-hamata 
maculisque tribus pallide flavis notata. 


Obscure bronze on the upper part, with the underside of the 
prothorax glowing-red, and the abdomen dark blue, legs glowing- 
red with the tarsi green; labrum elongate, convex, tridentate 
at the apex, white, and with two setigerous punctures on each side, 
one near the median tooth, the other near the outer angle; head 
aciculate, with a few white decumbent hairs; prothorax short, 
quadrate, rugose, covered with white decumbent hairs; elytra 
elongate, subparallel, finely granulose, and having on each side a 
moderately broad, pale-yellow marginal band, sending out three short, 
blunt rami, and also three round spots of the same colour on each 
side of the suture, the posterior one of which is nearly connected 
with the apex of the median marginal ramus, but the median one 
is not at all connected with the humeral ramus, and the basal one 
is placed far above it ; the marginal band and also the dorsal round 
spots are edged with dark bronze. Length, 10; width, 3mm. 


This species, which might be taken at first sight for a 
variety of (. marginella, Dej., is easily distinguished 
by having a sub-basal spot on each side of the suture, 
which could not be the interrupted end of the lateral 
humeral ramus, whereas the other two spots following 
the sub-basal one might be the continuation of the two 
lateral spurs. 


Hab. Mashunaland (Salisbury). 


Gen. Myrmecortpra, Germ. 
M. angusticollis, sp. n. 


Supra eneo-nigra, subnitida, subtus cyanea; capite strigoso, 
labro albido, nigro-marginato ; prothorace cylindrico, angusto, 
valde elongato, transverse plicato ; elytris elongatis, pone medium 
ampliatis, profunde punctatis, utroque plaga humerali elongata, 


new Cicindelidee from Mashwnaland. 449 


fascia transversa post medium posita vittaque postica supra- 
marginali albis ornato. 


Dark metallic blue, moderately shining on the upper part, 
underside cyaneous ; labrum white, narrowly edged with black in 
the male, broadly in the female ; joints of antenn foliate from 
the fifth to the apical one ; head strigose; prothorax very narrow, 
cylindrical, nearly twice as long as broad, very slightly narrowed 
in front and behind, and finely plicate transversely ; elytra very 
narrow at the base, gradually ampliated from the base to two-thirds 
of the length, convex with the sutural part ending on each side in 
a spine, longer and sharper in the male, broadly and deeply pitted, 
with the punctures deeper in the interior part, and having on each 
side a white humeral narrow band, a discoidal transverse post- 
median band narrowed in the centre, and reaching from the outer 
margin to two-thirds of the width of the disk, and a supra- 
marginal narrow longitudinal band of the same colour, reaching 
from the rounding of the posterior part to the apex ; legs cyaneous, 
with the knees slightly reddish. Length, 124-13 ; width, 3-33 mm. 


In shape, size, colour, and sculpture this species 
resembles much IM. spectabilis, Pér.; the joints of the 
antennee in M. angusticollis are foliate from the fifth to 
the apical one, whereas in M. spectabilis only the 4th, 5th, 
and 6th joints are foliate ; the prothorax is still narrower 
than in the last-named species, and the punctures on the 
elytra are not so broad; the humeral white band is 
similar, but the posterior one is shorter, and does not 
unite with the transverse discoidal one, 


Hab. Mashunaland (Salisbury.) 


M. mashuna, sp. n. 

Aingo-nigra, supra subnitida; capite strigoso, labro nigro, 
macula albida in medio; prothorace cylindrico, transverse plicato ; 
elytris elongato-ovatis, utrinque quinque-costatis, costis longis, 
sub-undulatis, interstitiis alveolatis. 


Black, moderately shining, with a bluish tinge on the head and 
prothorax, elytra dark bronze ; labrum black, witha median yellow- 
ish white patch in the male, this patch being hardly distinct in the 
female ; head conspicuously plicate; prothorax cylindrical, longer 
than broad, distinctly constricted in front and behind, and trans- 
versely plicate ; elytra elongated, gradually ampliated from the base 
to about two-thirds of the length, convex, the male with two 
moderately long sutural spines, the female without any, and having 


450 M. Louis Péringucy on descriptions of 


on each side five long raised lines, the outer two reaching from the 
base to about three-fourths of the length, the three discoidal ones 
somewhat shorter, apical part shagreened, intervals broadly foveate, 
the fovex with ‘a golden sheen, no supra-marginal white spot or 
band ; underside and legs very dark blue. Length, 17-19 ; width, 
43-6 mm. 


The nearest ally to this species is M. bertoloni; the 
shape of the prothorax is the same, but in M. mashuna 
the transverse folds are more raised and with a slightly 
broader interval, the costew of the elytra reach much 
nearer to the apex, and the alveolx on the disk are more 
regular, 


Hab, Mashunaland (Salisbury). 


M. marshalli, sp. n. 

/Enco-nigra, elytris weneis, subnitidis; capite strigoso, labro 
nigro, in medio albo-maculato ¢ ; prothorace subcylindrico, antice 
posticeque valde constricto, transverse plicato, vitta media albo- 
tomentosa ; elytris elongato-ovatis, utrinque quinque-costatis, costis 
post medium productis, interstitiis foveato-reticulatis, vitta basali 
alba in elytro singulo, plaga discoidali post medium posita vittaque 
postica supra-marginali albis. 

Black with a bronze tinge, elytra brassy, moderately shining, 
underside and legs black, with a bluish tinge; labrum black, 
with a broad median yellowish-white patch in the ¢; head 
very obliquely plicate between the eyes, and transversely so behind ; 
prothorax subcylindrical, only a little longer than broad, with 
the discoidal part somewhat raised on each side, narrowed in 
front and behind, transversely plicate, and with a median band 
of white hairs reaching from apex to base; elytra elongated, 
gradually ampliated from the base to about two-thirds of the 
length, ending in two sharp, long, sutural spines, convex, with 
five cost reaching from the base to two-thirds of the length, 
but the first and fifth longer than the three discoidal ones, inter- 
vals foveato-reticulate, apical part shagreened, on each side a white 
line running on the second costa from the base to about one-fourth 
of the length, a more or less sublunar or rounded patch of the same 
colour placed at the apex of the three discoidal raised lines, and a 
narrow, supra-marginal posterior band, extending from the same 
height as the discoidal patch to the apex. Length, 19; width, 
5 mm, 


I know the male only. The female will probably prove 


new Cicindelidee from Mashunaland, 451 


to be more ovate and more convex, with the labrum 
nearly totally black. 

In general facies, this species approximates to M. poly- 
hirmoides, but the markings are different from those of 
any other Myrmecoptera known up to date. 

Hab, Mashunaland (Salisbury). 


M. formosa, sp. n. 


/Eneo-nigra, supra subnitida, capite strigoso, labro nigro, in 
medio albo-vittato ; prothorace subquadrato, antice posticeque 
constricto, disco utrinque elevato, transverse plicato ; elytris elon- 
gato-ovatis, uirinque quinque-costatis, costis undulatis, interstitiis 
secundo quartoque profunde-alveolatis, vitta basali alba medium 
disci utriusque attingente plagaque triangulari ad apicem posita 


albis. 


Bronze-black, moderately shining on the upper part, dark 
metallic-blue underneath ; head conspicuously strigose; labrum 
black, with a median whitish longitudinal band in both sexes ; pro- 
thorax nearly quadrate, narrowed in front and behind, with the 
disk raised on each side and plicate ; elytra gradually ampliated 
from the base to about. two-thirds of the length, convex, ending in 
two moderately long apical spines, longer and sharper in the male 
than in the female, each elytron with five coste, the first and second 
near the suture very wavy, and reaching from the base to two- 
thirds of the length, the third costa equally long, while the fourth 
and fifth extend further, reaching to three-fourths of the length ; 
the intervals between the suture and the first costa and those 
between the second and fifth are broadly plicate, that between the 
first and second costs divided into six or seven broad, deep fovez, 
and the space between the fifth costa and the outer margin also 
broadly foveate ; in the fourth interval runs a basal yellowish-white 
band, reaching about the median part of the disk, and there is a 
supra-apical, more or less triangular, broad patch of the same 
colour, placed below the fourth and fifth costs; legs very dark 
blue. Length, 18-20; width, 5-7 mm. 


Allied to M. mauchi, Bates, but differentiated by the 
broad and deep fovezw in the second interval on the 
elytra; the dorsal longitudinal white band is longer in 


proportion, and the apical one more broadly triangular, 
Hab. Mashunaland (Salisbury). 


452 M. Louis Péringuey on descriptions of 


M. invicta, sp. n. 

/Eneo-nigra, supra-subnitida ; capite strigoso, labro nigro medio 
albo-vittato ; prothorace quadrato, antice posticeque constricto, 
disco utrinque elevato, transverse plicato ; elytris elongato-ovatis, 
quinque-costatis, interstitiis reticulatis linea basali medium disci 
haud attingente, plagaque supra-apicali lata in singulo elytro albis. 


Bronze-black, moderately shining on the upper side, underside 
and legs cyaneous-black; head strigose ; labrum black, with a 
median yellowish-white longitudinal band in both sexes ; prothorax 
quadrate, narrowed in front and behind, with the disk raised on 
each side and plicate ; elytra gradually ampliated from the base to 
about two-thirds of the length, convex, ending in two acute spines, 
long in the male, short in the female, each elytron with five cost 
reaching from the base to two-thirds of the length, with the first 
and second a little shorter than the other three, and the intervals 
reticulate ; on the second costa runs a narrow white line, running 
from near the base to about one-third of the length, and behind, 
edging the dorsal costz and extending more or less diagonally 
from the outer margin to the inner costa, isa sinuate silvery patch ; 
this patch is generally broader in the ¢ than in the ¢?, and I have 
seen a g¢ example where it had disappeared altogether. Length, 
15-20 ; width, 5-6 mm. 

A near ally to M. mauchi, Bates, but half the size; the 
sculpture of the elytra is identical, but the costz are 
better defined ; the inner one is shorter than the others, 
the fourth and fifth unite towards the apex in the male, 
but seldom in the female, while in the ¢ of M. mauchi the 
first and fifth costee are the longest, and unite below the 
others, and the posterior patch in the last-named species 
is triangular, short, not transverse, nearer the apex, and 
below the fifth costa. 

Hab. Mashunaland (Salisbury). 


Gen. Cosmema, Bohem. 
C. dolosa, sp. n. 


Elongata, nea, elytrorum lateribus cyaneis; capite strigoso, 
antennis subcompressis, foliatis; prothorace cylindrico, postice 
constricto, transverse plicato; elytris elongatis, postice leviter 
ampliatis, crebre profundeque punctatis, convexis, ad apicem 
utrinque bi-mucronatis, puncto humerali maculisque rotundatis 
supra marginem positis albis. 


new Cicindelidee from Mashunaland. 453 


Dark bronze, with a greenish tinge; the lateral parts of the 
elytra cyaneous; underside and legs dark greenish-blue, tibiz 
rufescent ; antenne subcompressed and slightly foliate ; head finely 
strigose diagonally ; labrum black, with the apical part somewhat 
yellowish in the male, totally black in the female ; prothorax narrow, 
cylindrical, constricted behind, finely transversely plicate ; elytra 
elongated, covered with deep, closely set, moderately broad punc- 
tures, convex, with the sutural part ending in two short spines on 
each side, as in C. lepida, Boh., but much shorter, and having on 
each side three ovate, supramarginal white spots, the first one 
humeral and very small, the median one a little elongated, and the 
posterior one ovate. Length, 10; width, 3 mm. 


This species, owing to the tendency of the antennz to 
become compressed and subfoliate, connects Cosmema 
with Myrmecoptera; like (. lepida, it has two spines on 
each side of the suture, but these spines are not so much 
developed ; the supramarginal white markings are not 
unlike those of C. elegantula, Boh., but in C. dolosa the 
posterior one is a round or ovate patch, not a white line 
or band. 


Hab. Mashunaland (Salisbury.) 


: a bsioes ved 
4 


Bae oh nt Len 
ee b Val on WW 


XVI. On Pyralidina from the Malay Archipelago. 
By Epwarp Meyrick, B.A., F.Z.S. 


[Read May “nd, 1894.] 


THE collection which forms the subject of this paper 
was made by Mr. W. Doherty, and is in the possession 
of Mr. H. J. Elwes, who entrusted me with types of the 
species for examination. The localities from which the 
specimens came comprise the Island of Sambawa, 
Celebes, Pulo Laut (Borneo), and §.E. Borneo. Many 
of the species are new; the large number of new and 
curious species of Oligustigma is especially remarkable. 
The fauna of Sambawa has evidently felt the effects 
of isolation, several of the species that occur there 
varying in a marked manner from the normal type of 
the same species occurring elsewhere. 


PYRAUSTIDA. 


ANISOCTENA, n. g. 


Forehead oblique ; ocelli present ; tongue developed. Antenne 
4,in ¢ with a sinuation at 4, basal fourth with short very uneven 
pectinations, longest on sinuation, twice fluctuating between that 
and base, with a notch above basal joint surmounted by a strong 
tooth. Labial palpi rather long, curved, ascending, second joint 
much thickened with dense projecting scales, terminal joint 
moderate, thick, obtuse. Maxillary palpi moderate, terminally 
dilated with scales. Thorax in ¢ with patagia forming a dense 
central tuft. Abdomen in g with lateral hair-tufts beyond middle, 
and dense black genital tuft. Posterior tibiee with outer spurs 
about half inner. Forewings in g with inner margin densely tufted 
with hairs towards base ; 3, 4, 5 approximated at base, 10 approxi- 
mated to 9. Hindwings densely haired above towards inner margin 
and on lower edge of cell towards base ; 3, 4,5 approximated at 
base, 7 out of 6 near origin, anastomosing with 8 to +. 


Allied to Margaronia, but with many curious characters. 
The hairy margin of cell of hindwings is, of course, 
abnormal in the Pyraustide. 


TRANS. ENT. SOC. LOND. 1894.—PART III. (SEPT.) 


456 Mr. E. Meyrick on 


A. synomotis, sp. n. 

3. 389 mm. Forewings somewhat elongate, hind margin oblique ; 
rather dark fuscous, purplish-tinged ; a darker discal mark, and an 
indistinct dot near before it. Hindwings rather dark fuscous. 


Celebes; one specimen. 


Margaronia laticostalis, Gn. Pulo Laut, Sambawa. 

M. diaphanalis, Walk. Pulo Laut. 

M. glauculalis, Gn. §8.E. Borneo. 

M. aquosalis, Snell. Celebes, 8.E. Borneo. 

M. pomonalis, Gn. Sambawa. 

M. costiflexalis, Gn. Sambawa. 

M. principalis, Walk. 8.H. Borneo. This species is 
closely allied to M. prothymalis, Swinh., but differs in 
the form of the large postmedian blotch of the forewings, 
which is rounded-oblong, the two ends about equally 
broad, whereas in prothymalis the blotch is triangular, 
the upper end being very much broader. 

M. badialis, Walk. S.E. Borneo. 

M. bwwitralis, Gn. S.E. Borneo. 

M. actorionalis, Walk. Pulo Laut. 

M. conclusalis, Walk. Sambawa. 

M. bicolor, Sv..ins. Sambawa. 

M. stolalis, Gn. Pulo Laut, 8.E. Borneo. 


M. sphenocosma, sp. n. 

dg. 19—21 mm. Antennal cilia 2. Forewings moderate, hind 
margin rather oblique ; fuscous-whitish; two straight, oblique, 
dark fuscous fasciz near base ; beyond these a parallel white fascia, 
edged with dark fuscous ; then a large transverse-oval dark fuscous 
spot, reaching from near costa to submedian fold ; beyond this a 
narrow triangular white spot from costa beyond middle, edged 
with dark fuscous, reaching half across wing ; second line slightly 
curved, dark fuscous ; terminal area beyond this wholly fuscous ; 
cilia fuscous, tips at anal angle white. Hindwings white ; a dark 
fuscous streak from costa near base to anal angle; a dark fuscous 
median fascia, furcate above so as to enclose a triangular spot ; 
second line straight, dark fuscous, terminal area wholly fuscous ; 
cilia fuscous, tips white on lower half of hind margin. 


Pulo Laut, S.H. Borneo; two specimens. 


M. scapulalis, Ld. (lymphatalis, Swinh.). Pulo Laut. 
M. tolumnialis, Walk. Pulo Laut. 


Pyralidina from the Malay Archipelago. 457 


Omiodes hiracia, sp. n. 

é6- 30 mm. Forewings rather elongate, hindmargin rather 
strongly oblique ; fuscous, costa somewhat lighter ; markings dark 
fuscous ; first line straight, second serrate and slightly curved on 
upper half, strongly broken inwards in middle, thence straight ; a 
dot beyond first line, and a discal mark : cilia fuscous, tips towards 
anal angle white. Hindwings grey ; a darker discal dot ; second 
line dark grey, upper half moderately curved ; cilia whitish, base 
dark grey. 

Pulo Laut; one specimen. 


O. erythrias, sp. n. 

dé. 20 mm. Forewings moderate, hindmargin somewhat 
oblique ; deep brown-reddish, ochreous-tinged anteriorly, more 
fuscous posteriorly ; lines indistinctly darker, first nearly straight, 
oblique, second strongly broken inwards in middle ; a darker discal 
mark : cilia snow-white, base dark fuscous. Hindwings dark grey, 
mixed with dark red in disc, costa whitish, inner margin suffused 
with whitish ; cilia as in forewings. 

Celebes; one specimen. 


Coptobasis radicalis, Walk. Pulo Laut. 
C. lunalis, Gn. Celebes, 8.E. Borneo, Sambawa. 
C. monochromalis, Walk. Celebes. 


TYLOSTEGA, g. 1. 


Face rounded ; ocelli present ; tongue developed. Antenne 3, 
in ¢ ciliated. Labial palpi moderate, curved, ascending, second 
joint with dense somewhat rough scales beneath, terminal joint 
moderate, obtuse or somewhat pointed. Maxillary palpi moderate, 
filiform. Abdomen rather long. Posterior tibize with outer spurs 
less than half inner. Forewings in ¢ above with impression in 
cell containing a small flat scale-tuft, beneath with depression 
covered by dense pecten of scales from upper edge (in 9 also 
present, but less developed) ; 3, 4, 5 closely approximated towards 
base, 10 closely approximated to 9. Hindwings with 3, 4, 5 closely 
approximated towards base, 7 out of 6, anastomosing with 8 to 
middle. 

Nearly allied to Aripana, from which it differs mainly 
in the scale-pecten of the forewings, and the shorter and 
less acute terminal joint of palpi. Type 7’. chrysanthes. 

TRANS. ENT. SOC. LOND. 1894,—Part 111. (SEPT.) 24 


458 Mr. HE. Meyrick on 


T’. chrysanthes, sp. n. 

6. 17—19 mm. Antennal cilia of g 3. Forewings rather 
elongate, hindmargin somewhat obliquely rounded ; orange, irregu- 
larly blotched with fuscous ; lines rather irregular, dark fuscous, 
forming blackish spots on costa, first hardly curved, second at ? 
acutely angulated to below middle of disc, and again acutely 
angulated to inner margin ; a blackish discal mark ; an almost hind- 
marginal row of blackish spots: cilia orange, base obscurely 
fuscous-spotted. Hindwings as forewings, but first line absent. 


Pulo Laut; two specimens. 


T. mesodora, sp. 0. 

¢. 19mm. Differs from 7. photias as follows : antennal cilia 
14; forewings somewhat shorter and broader, costal streak more 
orange-tinged, hindmargin with a slender orange streak preceded 
by a series of blackish spots ; hindwings shorter and more rounded, 
marked as in forewings, except that there is a broad ochreous- 
whitish antemedian band. 


Pulo Laut; one specimen. 
I 


T. photias, sp. n. 

¢. 20—21 mm. Antennal cilia 1. Forewings rather elongate, 
hindmargin somewhat obliquely rounded ; fuscous ; inner margin 
suffused with ochreous towards middle ; a suffused pale ochreous 
streak along costa, with a projection beyond second line ; lines 
dark fuscous, forming blackish spots on costa, first hardly sinuate, 
second sinuate, acutely angulated from near anal angle to below 
middle of disc, thence again to inner margin, buat obsolete at 
extremity ; a blackish discal mark ; a blackish hindmarginal streak, 
including a series of whitish dots: cilia whitish-grey, base dark 
grey. Hindwings whrtish: base fuscous; a broad rather dark 
fuscous hindmarginal band, anterior edge suffused with ochreous ; 
a fuscous mark in disc near before this ; a hindmarginal series of 
connected white dots ; cilia as in forewings. 


Pulo Laut, S.E. Borneo ; two specimens. 


T. (?), poltisalis, Walk. Pulo Laut; a 2; the generic 
reference is therefore doubtful. 
T. schematias, sp. n. 
$@. 19—21 mm. Antennal cilia of g¢ minute. Forewings 
somewhat elongate, hindmargin rather obliquely rounded ; 
ochreous-yellow ; five small round black spots, viz., one on costa 
at 3, two on inner margin before middle and towards anal angle, 


Pyralidina from the Malay Archipelago. 459 


one near hindmargin below middle, and one at apex; a black 
transverse mark from costa beyond middle; cilia ochreous- 
yellowish, tips paler. Hindwings pale ochreous-yellowish ; a small 
black spot at apex, and another on middle of hindmargin . 

Pulo Laut, S.E. Borneo, Sambawa, Celebes; four 
specimens. In the Celebes specimen the spots are 
partially indistinct. 

Aripana meritalis, Walk. Sambawa. 

A. abdicalis, Walk. Pulo Laut, S.E. Borneo, Sambawa. 

A. lactiferalis, Walk. (paucipunctalis, Snell.). Pulo 

Laut, Celebes. 

A, glaucias, sp. n. 

g-. 28 mm. Forewings white; a greyish-ochreous subcostal 
streak from base to beyond middle ; black basal and subbasal dots 
on costa, one beneath costa beyond these, and one on inner margin 
towards base ; lines broad, fascia-like, pale greyish-ochreous, first 
obsolete on upper half, second hardly curved, regular; a round 
black discal ‘spot; a black dot on costa at 2, and another, whitish- 
centred, on costal end of second line; a pale greyish-ochreous 
hindmarginal streak, preceded by a black dot below middle : cilia 
pale greyish-ochreous, tips paler. Hindwings white ; a black discal 
dot ; second line and posterior markings as in forewings. 

Pulo Laut; one specimen. Specially distinguished by 
the large size, and broad fascia-like regular second line; 
nearest to A. lactiferalis, but differs from it also by the 
black dorsal dot towards base. 


Rehimena phrynealis, Walk. Pulo Laut. 

Conogethes clioalis, Walk. Sambawa. 

C. haemactalis, Snell. Pulo Laut, Sambawa. 

C. punctiferalis, Gn. Pulo Laut, Celebes, Sambawa. 
C. iopasalis, Walk. Sambawa. 

C. nilalis, Walk. (— usalis). Pulo Laut. 

C. sublituralis, Walk. Celebes, Sambawa. 


Nosophora scotaula, sp. n. 

6. 27 mm. Forewings moderate, hindmargin obliquely 
rounded ; dark fuscous, Hindwings dark fuscous. 

Pulo Laut; one specimen. ‘The sexes in the genera 
Nosophora and Analthes appear to differ strikingly in 
some cases at least, and their relationships are at present 
imperfectly understood. I consider it just possible that 
this might be the d of the very different N. congenitalis, 
of which I have seen only ?s. 


460 Mr. E. Meyrick on 


N. ochnodes, Meyr. Pulo Laut. 
N. chironalis, Walk. Pulo Laut. 
N. congenitalis, Walk. Pulo Laut. 


Analthes semitritalis, Ld. Pulo Laut. 

A. idyalis, Walk. (disjunctalis, Walk.) Pulo Laut, 
Celebes. I consider it nearly certain that the 
forms described under these two names are sexes 
of the same species. 


A, pyrrhocosma, sp. 0. 
Q. 19 mm. Forewings rather elongate, hindmargin rather 


oblique ; yellow ; markings deep brown-red ; a costal streak, costal 
edge dark fuscous ; a basal spot, connected by a subcostal streak 
with first line ; lines irregular, first nearly straight, second at 2 
abruptly bent round to connect with discal spot, thence again 
abruptly bent to inner margin; discal spot transverse-oblong, 
containing a pale mark, and touching costal streak ; a broad, 
irregular hindmarginal band, partially touching second line: cilia 
blackish. Hindwings deep brown-red; basal half, a large oval 
postmedian spot, and a spot on middle of hindmargin yellow; apex 
suffused with blackish : cilia blackish. 


S.E. Borneo ; one specimen. 


Meroctena tullalis, Walk. Pulo Laut. 
M. Staintonw, Ld. Pulo Laut. 


MEROTOMA, g. n. 


Face rounded ; ocelli present ; tongue developed. Antenne 3, 
in g¢ moderately ciliated. Labial palpi moderate, subascending, 
second joint much thickened with dense projecting scales beneath, 
terminal joint very short, concealed, Maxillary palpi moderate, 
filiform. Posterior tibiz in g extremely short, ~ of tarsi, fur- 
nished with large tuft of hairs above, outer middle spur absent, 
inner middle spur greatly dilated and enlarged. Forewings with 
3, 4, 5 approximated at base, 10 approximated to 9. Hindwings in 
g with inner marginal pencil of hairs ; 3, 4,5 approximated at 
base, 7 out of 6 near origin, anastomosing with 8 to . 


M. dairalis, Walk. Pulo Laut, Celebes. 


SAGARIPHORA, g. 0. 


Forehead rounded ; ocelli present ; tongue developed. Antenne 
4 in @ moderately ciliated. Labial palpi moderately long, curved, 
ascending, second joint with dense rough projecting scales beneath, 
terminal joint rather long, with triangular taft in front. Maxillary 


Pyralidina from the Malay Archipelugo. 461 


palpi moderate, filiform. Abdomen rather long. Posterior tibiz 
in ¢ strongly dilated with dense scale-tufts above and beneath on 
jower half, basal joint of tarsi with dense rough scales beneath. 
Forewings with 3, 4,5 approximated at base, 10 approximated to 9. 
Hindwings in ¢ with inner marginal tuft of hairs; 3, 4, 5 
approximated at base, 7 out of 6 near origin, anastomosing with 8 
to +. 


Nearly allied to Agrotera. 


S. heliochlaena, sp. n. 

g. 28 mm. Forewings moderate, hindmargin rather strongly 
oblique ; ochreous-brownish, orange-tinged ; basal half ochreous- 
yellow mixed with ferruginous-orange, limited by a rather curved 
line ; second line formed by blackish dots, irregular : cilia ochreous- 
brown, tips darker. Hindwings yellowish, paler anteriorly; a 
cloudy longitudinal streak of blackish irroration from middle of 
disc to hindmargin. 


Sambawa; one specimen. 
Agrotera effertalis, Walk., Celebes. 
Aetholix flavibasalis, Gu., S.H. Borneo. 


SAROSCELIS, g. n. 


Characters of Notarcha, but posterior tibie in ¢ with dense tuft 
of hairs. 


S. nicoalis, Walk. Sambawa, S.H. Borneo. 

Dichoerocis pandamalis, Walk. (frenatalis, Ld.) Pulo 
Laut, S.E. Borneo, Sambawa. ‘This species varies 
considerably in size, and in the form and position 
of the second line of forewings. The Sambawa 
specimens have the lines finer than those from 
Borneo. 


Notarcha (?) nigrofimbrialis, Snell. Celebes. 

N. quaternalis, Z. Sambawa. 

N. compsogramma, sp. n. 

Q. 23mm. Forewings moderate, hindmargin rather obliquely 
rounded ; whitish-yellowish; two straight orange subbasal trans- 
verse streaks, first black-dotted on costa ; lines rather thick, orange, 
marked with black spots on costa, first straight, second moderately 
broken inwards below middle ; a round black discal spot ; a suffused 
orange subterminal streak : cilia whitish, with blackish basal and 
grey median lines. Hindwings as forewings, but without subbasal 


462 Mr. H. Meyrick on 


lines or black costal spots, discal spot orange, not touching either 
line. 

Sambawa; one specimen. Differs from N. tigrina, 
Moore, chiefly in the discal spot of hindwings not being 
connected with the second line. 


N. pyrrhalis, Walk. Pulo Laut. 

N. pyranthes, sp. n. 

2. 21mm. Forewings anes hindmargin rather obliquely 
rounded ; ochreous-yellow ; markings blackish ; subbasal and _ first 
lines straight; a transverse discal spot outlined with blackish, 
connected by a line with inner margin; an oblique streak from 
costa near apex, reaching half across wing ; a straight streak from 
apex to inner margin before anal angle, and a hindmarginal streak, 
confluent at apex: cilia shining grey, base yellow. Hindwings 
ochreous-yellow, with an orange anal blotch ; discal spot blackish, 
connected by a bent line with anal angle ; an oblique blackish spot 
before apex, and a narrower mark before middle of hindmargin ; a 
black hindmarginal streak : cilia as in forewings. 


S.HE. Borneo ; one specimen. 


N. rigidalis, Snell. Pulo Laut. 
N. xuthalis, Walk. (— usalis) (onustalis, Snell.) 8.E. 
Borneo. 

N. multilinealis, Gn. Pulo Laut, Celebes, Sambawa. 

N. charesalis, Walk. (octasema, Meyr.) 8.H. Borneo. 

N. homomorpha, sp. n. 

d. 27 mm. Legs white, anterior pair banded with black. 
Forewings moderate, hindmargin rather obliquely rounded ; light 
ochreous-brown ; lines blackish, interrupted, first rather irregular, 
second serrulate, sinuate on upper 2, thence abruptly broken 
inwards to below middle of disc; a blackish dot in disc imme- 
diately beyond first line, and a small blackish discal spot; an 
interrupted blackish hindmarginal streak: cilia light brownish, 
basal half barred with blackish. Hindwings light ochreous-grey, 
darker grey towards apex ; a darker grey discal mark ; second line 
faintly darker, formed as in forewings; a blackish hindmarginal 
streak : cilia grey-whitish, basal half dark grey. 


Pulo Laut; one specimen. 
N. unitalis, Gn. Pulo Laut. 
N. iophanes, sp. n. 


6d. 35 mm. Abdomen with two expansible tufts of hairs at 
base above. Forewings rather elongate, hindmargin rather oblique; 


Pyralidina from the Malay Archipelago. 463 


fuscous, purplish-tinged, costa and terminal area dark fuscous ; a 
small purple-whitish spot in disc before very indistinct first line ; 
a small dark fuscous spot in disc beyond first line, and a transverse 
dark fuscous discal spot, separated by a larger square purple- 
whitish spot, and followed by a small purple-whitish spot ; two 
obscure purple-whitish spots between veins beneath central spot ; 


= 


second line dark fuscous, upper 2 curved outwards, edged pos- 
teriorly by a whitish line ending in a rather large subcostal and 
small subdorsal spot: cilia dark fuscous. Hindwings fuscous, with 
a broad dark fuscous terminal band; a dark fuscous discal spot, 
and twice strongly sinuate second line: cilia dark fuscous, towards 
tips whitish on upper half of hindmargin. 

S.E. Borneo; one specimen. 

N. decialis, Walk. (— usalis). Pulo Laut. 


Pleuroptya awrantiacalis, F.R. Pulo Laut, S.E. Borneo. 


MErToporRTHA, g. n. 


Forehead rounded, with long projecting spreading tuft of hairs 
from between antenne ; ocelli present ; tongue developed. An- 
tenn 3, in’ ¢ ciliated, with a sinuation and slight thickening of 
rough scales on stalk rather near base, basal joint enlarged. Labial 
palpi rather long, ascending, second joint rather shortly rough- 
scaled, terminal joint moderate, obtuse. Maxillary palpi mederate, 
terminally dilated with scales. (Legs broken.) Forewings with 3, 
4, 5 approximated at base, 10 approximated to 9. Hindwings with 
3, 4, 5 approximated at base, 7 out of 6 near origin, anastomosing 
with 8 to 5. 

M. dolopsalis, Waik. Pulo Laut. 

Phlyctaenia itemalesalis, Walk. Sambawa. 


Pyrausta inscisalis, Walk. Sambawa. 

P. celatalis, Walk. Pulo Laut. 

. ciniferalis, Walk. Pulo Laut, 8.H. Borneo. 

. ablactalis, Walk. §S.E. Borneo, Celebes, Sambawa. 
. damastesalis, Walk. Sambawa. 

. detritalis, Gn. Pulo Laut. 

. marginalis, Moore. §S.H. Borneo. 


asnelachege) a 


CryYPSIPTYA, 2. n. 


Characters of Pyrausta, but antenne $, thorax in ¢ beneath 
with large lateral extensible plate of greatly elongated scales. 


}, nereidalis, Ld. Pulo Laut, S.E. Borneo. 


46 | Mr. E. Meyrick on 


Tsocentris aequalis, Ld. Celebes. 


Ischnurges gratiosalis, Walk. Pulo Laut, S.H. Borneo. 
I. octoguttalis, Feld. S.H. Borneo. 


Dasyscopa, g. n. 


Characters of Scoparia, but hindwings in g with subdorsal 
groove, hairy above and filled beneath with dense hairs and with a 
dense tuft of hairs from middle of its inner edge, outer edge with 
long fine spreading hairs, 4 and 5 widely remote, parallel. 


D. homogenes, sp. un. 

gd. 16 mm. Forewings elongate, hindmargin rather oblique; 
pale grey, mixed with whitish and coarsely sprinkled with blackish ; 
a short black irregular streak from middle of base; lines whitish, 
first nearly straight, edged posteriorly with blackish, second ob- 
tusely angulated above middle; two small blackish spots repre- 
senting orbicular and claviform touching black edge of first line ; 
discal spot irregular, blackish; a broad irregular whitish sub- 
terminal line, interrupted in middle. Hindwings pale grey ; 
tufts of subdorsal groove blackish. 


Sambawa ; one specimen. 


IscHNOScopA, g. n. 


Characters of Scoparia, but forewings with 7 absent, hindwings 
with 5 absent. 


I. chalcozona, sp. n. 

36 @. 9—10 mm. Labial palpi elongate, pointed. Forewings 
elongate, hindmargin rather strongly oblique; dark fuscous, 
purplish-tinged ; lines rather thick, yellow, first hardly curved, 
second angulated in middle. Hindwings dark grey. 


Sambawa, over 3,000 feet; three specimens. 


ENDOGRAPHIS, g. 0. 


Forehead rounded ; ocelli present ; tongue developed. Antenne 
4, in ¢ moderately ciliated. Labial palpi moderate, ascending, 
second joint with long dense projecting scales beneath, terminal 
joint rather short, obtuse. Maxillary palpi moderate, filiform. 
Abdomen in ¢ with two or three long filaments from each side 
above middle. Posterior tibie with outer spurs half inner. Fore- 
wings with 3, 4,5 approximated at base, 10 approximated to 9. 
Hindwings in ¢ with deep subdorsal groove, inner margin folded 
over beneath and prolonged into a stiff projection at anal angle ; 


Pyralidina from the Malay Archipelago. 465 


4 and 5 short-stalked from point with 3,7 out of 6 near origin, 
anastomosing with 8 to middle. 


A genus of uncertain affinity. 


Hi. acrochlora, sp. n. 

¢. 15mm. Forewings moderate, apex rectangular, hindmargin 
rather oblique ; whitish-fuscous ; costa suffused with whitish- 
ochreous, more clearly posteriorly ; lines dark grey, indistinct, 
forming blackish marks on costa, first curved, second strongly 
curved on upper 3 ; an indistinct dark grey discal mark : cilia grey, 
with a blackish line, base ochreous-white. Hindwings grey, more 
whitish towards base and inner margin; second line dark grey, 
partially indistinct : cilia as in forewings, on lower half of hind- 
margin wholly whitish-grey. 

Pulo Laut ; one specimen. 

Stenia fauculalis, Walk. 8.H. Borneo. 


Metasia hilarodes, sp. n. 

¢. 20 mm. Antennal cilia minute. Forewings moderate, 
hindmargin obliquely rounded ; yellow-ochreous ; lines fuscous, 
first indistinct, second black-dotted on costa, followed by three 
semi-transparent whitish dots on upper third, abruptly curved 
outwards on middle third, thence strongly broken inwards and 
obsolete to beneath middle of disc, whence it is continued to inner 
margin ; a fuscous discal mark, preceded by a quadrate semi- 
transparent whitish spot, beneath which is another similar an- 
teriorly dark-edged spot preceding second line ; three dark fuscous 
dots on costa posteriorly : cilia ochreous-yellow, tips paler. Hind- 
wings ochreous-yellow ; second line as in forewings, but without 
whitish spots. 


Borneo (?); one specimen. 
M. tampialis, Walk. (— usalis). Pulo Laut, 8.E. 
Borneo. 


M. didasalis, Walk. Sambawa. 
M. melesalis, Walk. S.H. Borneo. 


MB=TASIODES, g. 0. 
Characters of Metasia, but face without prominence, labial palpi 
ascending, rough-scaled beneath, terminal joint short, obtuse. 


M. achromatias, sp. n. 
¢. 14—16mm. Antennal cilial?. Forewings elongate, hind- 
margin rather strongly oblique ; grey-whitish, irregularly mixed 


466 Mr. H. Meyrick on 


with dark grey ; some black scales towards base ; lines black, first 
nearly straight, followed by a small black subcostal spot touching 
it, second irregular, from + of costa to near anal angle, thence 
strongly curved round to below middle of disc, and again bent to 
inner margin at ? ; a transverse discal spot outlined with black ; 
three small blackish spots-on posterior half of costa besides 
extremity of second line; a dark grey spot on middle of hind- 
margin ; a hindmarginal row of blackish dots: cilia whitish, basal 
half spotted with blackish-grey. Hindwings with colour, second 
line, hindmarginal dots, and cilia as in forewings. 
Sambawa ; two specimens. 


M. calliophis, sp. n. 

dg. 13 mm. Antennal cilia 1. Forewings rather elongate, 
hindmargin oblique ; ochreous-yellow ; a black spot on base of 
costa; lines black, forming spots on costa, first nearly straight, 
second irregularly sinuate from 3 of costa to near anal angle, 
thence strongly curved round to beneath middle of disc, and again 
bent to inner margin at 2; orbicular and transverse discal spots 
outlined with black ; a small black spot on anal angle, and four 
black dots on upper half of hindmargin : cilia whitish, with a 
dark grey line. Hindwings with colour, second line, anal spot, 
and cilia as in forewings ; a black discal dot; a black streak along 
upper half of hindmargin. 

Pulo Laut; one specimen. 


M. lilliputalis, Snell. (?) Sambawa. 
M. tholeropa, sp. n. 


g. 9mm. Antennal cilia 4, Forewings elongate, hindmargin 
rather oblique; whitish-fuscous, irregularly mixed with dark 
fuscous ; costa blackish towards base ; lines black, forming spots 
on costa, first nearly straight, second rather irregular from + of 
costa to anal angle, which it touches, thence acutely angulated to 
beneath middle of disc, and again strongly curved round to inner 
margin at 2 ; a blackish subcostal spot beyond and touching first 
line; discal spot transverse-oblong, outlined with black ; costa 
ochreous-whitish on posterior half, with a small black spot in 
middle, and a dot between this and second line; a hindmarginal 
series of connected black dots : cilia whitish, with a blackish some- 
what interrupted line. Hindwings with colour, second line, and 
cilia as in forewings; a black discal dot ; a waved black hind- 
marginal line. 


Sambawa ; one specimen. 


Pyralidina from the Malay Archipelago. 467 


M. ceavalis, Walk. S.E. Borneo. 


Tabidia insanalis, Snell. Celebes. 

T’. aculealis, Walk. (trisignata, Moore) Pulo Laut. 

T’. craterodes, sp. n. 

$ @. 19—22 mm. Differs from 7. insanalis as follows :— 
colour more whitish, markings thicker and darker, especially hind- 
marginal band ; forewings with upper half of discal spot almost 
wholly dark; hindwings with first line connected by suffused 
longitudinal streaks with base, a well-marked blackish discal spot 
beyond and more or less connected beneath with first line, 
second line distinctly curved (in insanalis straight); apex of 
abdomen in ¢@ rather dark ‘fuscous beyond the silvery-white 
subapical ring. 

Pulo Laut; two specimens. 


SYNTOMODORA, g. n. 


Forehead rounded ; ocelli present ; tongue developed. Antennz 
4, in ¢ serrate, shortly ciliated. Labial palpi moderately long, 
curved, ascending, second joint with rather long dense projecting 
scales beneath, terminal joint moderately long, pointed. Maxillary 
palpi moderate, filiform. Abdomen rather long. Posterior tibiz 
with outer spurs }—2 of inner. Forewings with 3, 4,5 approxi- 
mated at base, 10 approximated to 9. Hindwings with 4 and 5 
stalked from near 3, 7 out of 6 near origin, anastomosing with 8 to 
middle, 


Apparently allied to Tabidia. 
S. thoasalis, Walk. Celebes. 


Ptilaeola collaris, Walk. Pulo Laut, Sambawa. 
P. zebinalis, Walk. S.E. Borneo. 


Archernis callixantha, Meyr. Pulo Laut. 

Diathrausta profundalis, Ld. Sambawa. 

Diasemia grammalis, Dold. 8.E. Borneo, Sambawa. 

Agathodes ostentalis, Hb. Sambawa. 

Syngamia castoralis, Walk. ( purpurescens, Moore). 
Pulo Laut. 

Cnaphalocrocis medinalis, Gn. §.H. Borneo. 

Stegothyris diagonalis, Gn. Sambawa. 

Pagyda salvalis, Walk. Pulo Laut. 

P. amphisalis, Walk. Pulo Laut. 


468 Mr. EH. Meyrick on 


Platamonia ampliatalis, Ld. Celebes. 

P. camillalis, Walk. Pulo Laut. 

P. ptochura, sp. n. 

3S. 23mm. Abdomen with two dark fuscous subapical rings, 
not marked with white. Forewings moderate (broader than in 
camillalis), hindmargin rather obliquely rounded ; pale ochreous- 
yellowish ; a broad costal suffusion, broadest before middle, and a 
broad hindmarginal band dark fuscous; lines blackish, partially 
indistinct, first hardly curved ; second rectangularly but obsoletely 
broken inwards in middle; a blackish linear discal mark: cilia 
whitish, with a grey line. Hindwings with colour, second line, 
hindmarginal band, and cilia as in forewings. 


Pulo Laut; one specimen. 


Pleonectusa macaralis, Walk. Pulo Laut, Sambawa. 


Cometura lirisalis, Walk. (picrogramma, Meyr.) S.E. 
Borneo. 


Nistra caelatalis, Walk. S.E. Borneo, Sambawa, 
Celebes. 


Hydriris elutalis, Walk. Pulo Laut. 

H. ornatalis, Dup. Pulo Laut, S.E. Borneo, Sambawa. 

H. retractalis, Walk. (opalina, Moore). Pulo Laut, 
S.E. Borneo, Sambawa. 


Caprinia felderi, Ld. Pulo Laut. 
Pinacia fulvidorsalis, Hb. Sambawa. 


Sirtocauta testulalis, Hb. Sambawa. 


Nausinoe globulipedalis, Walk. (columalis, Snell.) 
Celebes. 

N. ewroalis, Swinh. (rivulalis, Snell.) Pulo Laut. 

N. conchylia, sp. n. 

¢. 20 mm. Forewings elongate, hindmargin very oblique, 
sinuate above anal angle ; cell with an irregular impression beneath, 
2—5 approximated at base, 10 out of 9; prismatic whitish, thinly 
scaled ; markings rather dark fuscous, ochreous-tinged ; first line 
irregular, indistinct, furcate towards inner margin, second forming 
an angular projection posteriorly in middle ; discal spot formed by 
a large oval ring, lower extremity connected with second line ; an 
irregular subterminal fascia, dilated to touch hindmargin in middle. 
Hindwings broad, veins 2—5 closely approximated at base, 7 
anastomosing with 8 almost to apex ; prismatic whitish, thinly 


Pyralidina from the Malay Archipelago. 469 


scaled; markings rather dark fuscous; a discal spot, lower ex- 
tremity touching a large dorsal suffusion ; an apical blotch; a 
triangular spot on hindmargin below middle. 


Pulo Laut; one specimen. 


N. trogalis, Walk. (ommatalis, Snell.) Pulo Laut. 
N. lindalis, Walk. © Sambawa. 


SYMMORACMA, &. Nn. 


Forehead somewhat prominent ; ocelli present; tongue de- 
veloped. Antenne 1, in ¢ slender, simple, in 2 minutely ciliated. 
Labial patpi moderate, porrected, clothed with dense projecting 
scales. Maxillary palpi moderate, filiform. Abdomen in ¢ very 
long. Posterior tibiz with outer spurs about half inner. Fore- 
wings with 3, 4, 5 approximated at base, 10 out of 9. Hindwings 
with 3, 4, 5 approximated at base, 7 out of 6 near origin, thence 
coinciding with 8 to apex. 


Probably related to Nawsinoe. 

S. spodinopa, sp. n. 

6 2. 19—20 mm. Forewings elongate, hindmargin oblique ; 
purplish-grey, irrorated with black; a small white spot on sub- 
median fold before middle ; a white crescentic discal mark ; second 
line whitish, middle third curved outwards ; a hindmarginal row 


of white dots: cilia grey. Hindwings grey ; a fine darker post- 
median line, formed as in forewings. 


Sambawa ; two specimens. 


Orphnophanes euceralis (— usalis) ( productalis, Ld.) 
S.E. Borneo. 


PmrRISYNTROCHA, g. 0. 


Forehead rounded ; ocelli present ; tongue developed. Antennze 
almost 1, in ¢ very shortly ciliated. Labial palpi moderate, 
ascending, second joint rough-scaled beneath, terminal joint short, 
obtuse. Maxillary palpi short, filiform, pointed. Abdomen in ¢ 
long. Posterior tibize with outer spurs about half inner. Forewings 
with 4 and 5 stalked, 10 out of 9. Hindwings with 4 and 5 
stalked, 7 out of 6 near origin, anastomosing with 8 to middle. 


Allied to Nausinoe. 


P. alienalis, Walk. (cuneolalis, Snell.; picata, Butl.) 
Pulo Laut. 


470 Mr. E. Meyrick on 


Mixophyla venatalis, Walk. (— usalis) (erminea, 
Moore). Sambawa. 


Pterygisus fedalis, Gn. Sambawa, Pulo Laut. 
Cymoriza fulvobasalis, Snell. §.H. Borneo. 


Hydrocampa oxygona, sp. n. 

gd. 21 mm. Forewings fuscous ; lines slender, white, first 
curved, second angulated beneath costa, at 2 abruptly curved up- 
wards to middle of disc, thence again abruptly curved downwards, 
angulated above dorsum ; a subterminal row of uneven white dots. 
Hindwings fuscous, lighter in disc; extreme base and a subbasal 
line whitish; lines slender, white, first hardly curved, second 
forming an acute angular prominence in middle ; a round blackish 
discal spot ; a subterminal row of white dots. 


Sambawa; one specimen. 


Nymphula irialis, Walk. (—usalis). Pulo Laut. 

N. responsalis, Walk. §.H. Borneo. 

N. turbata, Butl. S.E. Borneo. 

N, diminutalis, Snell. S.E. Borneo. 

N. villidalis, Walk. (unilinealis, Snell.) S.E. Borneo. 


OLIGERNIS, g. n. 


Forehead rounded ; ocelli present ; tongue developed. Antenne 
3, in ¢@ serrate, minutely ciliated. Labial palpi long, curved, 
ascending, second joint loosely scaled beneath, terminal joint nearly 
as long, acute. Maxillary palpi moderate, with loose spreading 
scales. Posterior tibize with outer spurs almost: as long as inner. 
Forewings with 3, 4, 5 approximated at base, 10 out of 9, 11 
absent. Hindwings with 3 and 4 froma point, 7 out of 6 near 
origin, anastomosing with 8 to near middle. 

A specialised development of Nymphula. Type, O. 

endophthalma. 


O. endophthalma, sp. n. 


&@. 11 mm. Forewings elongate-triangular, apex rectangular, 
hindmargin bowed, little oblique ; white ; basal area suffused with 
light fuscous ; first line white, indistinct, edged on costa with 
ochreous-yellow, elsewhere with a cloudy dark fuscous suffusion ; 
an ochreous-yellow streak from middle of first line to 3 of disc, 
thence angulated to costa at ¢; a similar ochreous-yellow streak- 
beneath and parallel to this, but more curved, not reaching costa. 


Pyralidina from the Malay Archipelago. 471 


Hindwings white ; first line white, indistinct, edged with a cloudy 
dark fuscous suffusion ; posterior half mostly occupied by three 
ochreous-yellow blotches, apical, central, and anal. 


S.E. Borneo ; one specimen. 


O. leucochrysa, sp. n. 


?. 12 mm. Forewings elongate-triangular, apex rectangular, 
hindmargin bowed, little oblique ; white; markings ochreous- 
yellow ; margins of first line formed by two interrupted streaks ; 
two parallel streaks from middle of disc to 3, thence angulated to 
costa before apex; a submarginal streak, finely edged with 
blackish anteriorly. Hindwings white ; three ochreous-yellow 
rather irregular transverse fasciz, second dilated in disc ; sub- 
marginal streak as in forewings. 


S.E. Borneo; one specimen. A third species of the 
same genus is in the collection from the same locality, 
but not in condition to describe. 


Oligostigma ceratucha, sp. n. 


36 @. 19—24mm. Antenne in ¢ with eight lowest joints of 
stalk clothed with long suberect scales above. Forewings very 
elongate-triangular, hindmargin very oblique, rounded, in ¢ with 
costal edge very narrowly folded over beneath on basal 2, and 
roughly hairy towards base ; 10 and 11 out of 9; shining white ; 
a fuscous costal streak from base to middle; a broad ochreous- 
yellow dorsal streak from base to anal angle, enclosing a narrow 
fuscous dorsal streak towards middle, tending to be connected by 
an indistinct fuscous suffusion with costa ; a fuscous triangular 
blotch on costa beyond middle, subcostal vein in this finely white ; 
a yellow-ochreous streak, suffused with fuscous posteriorly, from 
costa at + to apex of costal blotch ; an ochreous-yellow finely 
black-edged hindmarginal streak, enclosing a hindmarginal series of 
fine white lunulate marks. Hindwings abruptly sinuate below 
apex, in ¢ with strong rounded anal prominence, and deep sub- 
dorsal furrow beneath ; ochreous-yellow ; a white fuscous-edged 
fascia before middle ; a white fascia beyond middle, not reaching 
margins, edged anteriorly with fuscous and posteriorly with black ; 
an interrupted black hindmarginal line, forming two small spots 
below subapical sinuation, upper preceded by a silvery-white black- 
edged dot. 


Pulo Laut ; two specimens. 


472 Mr. HE. Meyrick on 


O. endosaris, sp. n. 

@. 18 mm. Forewings very elongate-triangular, hindmargin 
very oblique, rounded, posterior end of cell forming a subhyaline 
impressed patch, inner margin with abnormally long cilia ; 10 out 
of 9 ; light fuscous; a small dark fuscous spot on middle of costa, 
preceded by a yellowish spot; a deep yellow transverse spot in 
disc beyond middle ; two shining white posterior streaks from 
costa, first reaching to 2, second to near dorsum, space between 
them deep yellow anteriorly, second followed by a deep yellow 
anteriorly blackish-edged hindmarginal fascia; a hindmarginal 
row of blackish dots. Hindwings abruptly sinuate below apex, 
in ¢ at anal angle with a large rough tuft of mixed black and 
whitish hairs; deep yellow ; a white fuscous-edged fascia before 
middle; apex white, with black marginal and submarginal lines ; 
an interrupted black hindmarginal line, forming two or three 
spots below subapical indentation, upper preceded by a silvery- 
white black-edged spot. 


Pulo Laut ; one specimen. 


O. orthoteles, sp. n. 

4¢@. 20—23 mm. Posterior femora in ¢ witha large dense 
rough tuft of whitish and black hairs beneath. Forewings very 
elongate-triangular, hindmargin straight, rather strongly oblique ; 
10 out of 9 ; shining white; a deep ochreous-yellow streak along 
costa from base to middle, thence to # of disc; a black mark on 
middle of costa; a cloudy blackish median line from base to 
beyond middle of disc, thence bent downwards to near anal angle, 
and again acutely angulated to costa at 3, where it forms a yellow 
spot ; a deep ochreous-yellow dorsal streak from before middle to 
anal angle; a deep ochreous-yellow posteriorly fuscous-edged 
fascia at 5, finely attenuated downwards, not reaching anal angle ; 
a deep ochreous-yellow black-edged hindmarginal fascia. Hind- 
wings angularly indented below apex; deep ochreous-yellow ; a 
shining white black-edged median fascia ; black submarginal and 
marginal lines, latter interrupted to form three black spots below 
subapical indentation, uppermost preceded by a white black-edged 
spot. 

Sambawa ; two specimens. 


O. aureolalis, Snell. Sambawa. 

O. ochreipicta, Moore. Celebes. 

O. idiotis, sp. n. 

@. 18mm. Forewings elongate-triangular, hindmargin bowed, 
oblique; 10 out of 9; fuscous mixed with orange, costa and 


Pyralidina from the Malay Archipelago. 473 


inner margin suffused with orange ; a white subterminal streak, 
not reaching inner margin, preceded by a rather darker fascia, and 
followed by an orange black-edged hindmarginal fascia. Hind- 
wings abruptly sinuate below apex ; brownish-orange ; a grey, 
darker-sprinkled patch occupying anterior half, except extreme 
base and inner margin; a slender whitish subterminal streak on 
dorsal half, edged anteriorly with fuscous, and posteriorly with 
black ; a whitish, fuscous-edged anteapical dot; a black hind- 
marginal line on lower half, forming three black spots below sub- 
apical sinuation, two upper quadrate and edged anteriorly by 
white black-edged spots. 


Celebes; one specimen. Very like O. ochreipicta, but 
entirely without the strongly-developed costal fold and 
dorsal protuberance of that species. 


O. gibbosalis, Gn. ( plicatalis, Walk. ; hamalis, Snell.) 
Celebes, Sambawa; the Celebes form has the tri- 
angular costal postmedian blotch in ¢ wholly 
brown, the Sambawa form partly white; in the 
g of both forms the blotch is similar, mostly 
white. 


O. tripunctalis, Snell. Sambawa. 

O. melanodes, sp. n. 

Q. 12 mm. Forewings very elongate-triangular, hindmargin 
rounded, rather strongly oblique ; 10 out of 9 ; dark fuscous, 
whitish-sprinkled ; a whitish streak from costa at 3, reaching half 
across wing; a subterminal white streak, not reaching inner 
margin, followed by a brownish-orange, blackish-edged hind- 
marginal streak. Hindwings abruptly sinuate below apex ; dark 
fuscous; three black orange-edged hindmarginal spots below 
sinuation, preceded by white spots. 


S.E. Borneo; one specimen. 


O. argyropis, sp. 0. 

o2¢. 16—20 mm. Middle femora in ¢ beneath with tuft of 
long white hairs. Forewings very elongate-triangular, hindmargin 
rounded, oblique, inner margin in ¢ forming a strong triangular 
prominence at 2; 10 out of 9, 11 closely approximated ; shining 
white ; a dark fuscous costal streak from base to 4, lower edge 
forming a triangular projection beyond middle ; a light yellowish 
curved streak from 1 of inner margin to beneath this projection ; 
a rather broad orange hindmarginal fascia, edged with black marks 
or dots, enclosing a fine silvery blackish-edged subterminal streak. 


TRANS. ENT. SOC. LOND. 1894,—parT lI. (SEPT.) 24 


474 Mr. E. Meyrick on 


Hindwings abruptly sinuate below apex ; shining white, posteriorly 
suffused with light yellow-cchreous, becoming orange on hind- 
margin; a silvery submarginal streak before apex; five black 
hindmarginal dots below sinuation, preceded by silvery-metallic 
blackish-edged dots. 


Celebes ; two specimens. 


O. basilissa, sp. n. 

g. 16 mm. Posterior tibia with two spreading tufts of 
hairs at origin of spurs beneath. Forewings elongate-triangular, 
hindmargin bowed, rather strongly oblique ; inner margin forming 
a strong triangular prominence at +; 10 out of 9, 11 closely 
approximated ; shining white ; an orange costal streak from base 
to +, lower edge forming a strong triangular posteriorly blackish- 
margined projection beyond middle; a broad ochreous-yellow 
dorsal streak from base to 3, enclosing a blackish dorsal streak 
from 1 to 2; a rather broad orange hindmarginal fascia, edged 
anteriorly by a black streak not reaching inner margin, and en- 
closing a silvery-white black-edged subterminal streak ; an inter- 
rupted black hindmarginal line, forming a small spot at apex. 
Hindwings abruptly sinuate below apex ; shining white, posteriorly 
suffused with light ochreous-yellow, anterior edge of suffusion 
marked with two black spots in disc; an orange hindmarginal 
streak, at apex preceded by a silvery-metallic line, below sinuation 
marked with five round black spots anteriorly edged with silvery- 
metallic. 


Sambawa; one specimen. 
Margarosticha bimaculalis, Snell. Celebes. 


Anydraula xanthobathra, sp. n. 


9. 14 mm. Forewings whitish; a basal spot and inwardly 
oblique fascia beyond it deep ochreous-yellow, fuscous-edged, on 
costa suffused with fuscous; median area densely irrorated with 
blackish, on costa suffused with fuscous; three posterior deep 
ochreous-yellow fuscous-edged fascize (third hindmarginal) con- 
verging from costa to anal angle, where they form a blotch 
containing a metallic grey spot, lower half of space between 
second and third shining grey ; hindmargin sinuate above anal 
angle, sinuation more marked in dark basal portion of cilia, 
Hindwings whitish ; an ochreous-yellow fuscous-edged fascia about 
*, connected below middle by a slender metallic-edged streak with 
anal angle ; median area irrorated with blackish, limited posteriorly 


Pyralidina from the Malay Archipelago. 475 


by a double blackish line; four large irregular black hind- 
marginal spots, separated by narrow ochreous-yellow marks, and 
containing pale bluish-golden-metallic dots. 

Pulo Laut; one specimen. Near the Indian A. june- 
talis, Hamps., but the yellow basal spot of forewings is a 
sufficient distinction. 


Cataclysta nyctopis, sp. n. 


g. 14 mm. Forewings rather dark fuscous ; first line and 
costal portion of second suffusedly darker ; dull brownish-orange 
subterminal and hindmarginal streaks from costa reaching half 
across wing, lower extremities separated by a shining prismatic- 
grey spot ; a brownish-orange hindmarginal mark above anal angle, 
preceded by a metallic-grey spot. Hindwings with 5 and 8 present ; 
rather dark fuscous ; a whitish dark-edged sinuate transverse discal 
mark ; second line obscurely whitish, edged with dark fuscous ; an 
irregular black hindmarginal fascia, not reaching margins, obscurely 
edged with ochreous-whitish anteriorly, and containing five ill- 
defined bluish-leaden-metallic spots. 


S.H. Borneo; one specimen. 
Schoenobius punctellus, Z. Pulo Laut, $.H. Borneo. 


S. celidias, sp. n. 


g. 20—27 mm. Antennal ciliations 1. Forewings elongate- 
triangular, hindmargin bowed, oblique; brown; a longitudinal 
darker suffusion above middle of disc ; three dark fuscous dots on 
fold, towards base and before and beyond middle ; a fourth in dise 
beyond middle ; an ill-defined irregular dark fuscous line from 
costa before apex to third dot on fold. Hindwings white at base, 
changing gradually to light brown at apex. 


S.E. Borneo ; two specimens. 


Scirpophaga mnesidora, sp. n. 

g. 24 mm. Labial palpi very long. Abdomen with a yellow- 
ochreous subbasal band. Forewings with hindmargin nearly 
straight, rather strongly oblique; white; a deep yellow costal 
streak from base to apex, enclosing some metallic and blackish 
scales towards base, elsewhere indistinctly margined with dark 
fuscous beneath; lines dark metallic-fuscous, first somewhat 
inwards-curved, preceded by a short projection from costal streak, 
second from beyond 3 of costa to near anal angle, thence sharply 
bent to middle of disc, and again angulated to 3 of inner 


476 Mr. HE. Meyrick on 


margin; a triangular projection from middle of costal streak, 
edged with dark fuscous, touching angle of second line; anal 
space enclosed by second line ochreous-yellow ; a metallic-grey 
hindmarginal streak, partly edged anteriorly with dark fuscous ; 
cilia deep yellow, tips whitish. Hindwings white. 


Sambawa; one specimen. Although so differently 
marked from an ordinary Scirpophaga, there is no 
difference of structure except that the palpi are longer 
than usual. 


PYRALIDIDA. 


Endotricha decessalis, Walk. Sambawa. 

HH. orthotis, sp. n. 

$- 15—17mm. Forewings with hindmargin straight, oblique ; 
4 and 5 stalked; yellowish-crimson, posteriorly finely black- 
sprinkled ; first and second lines straight, parallel, blackish, first 
very indistinct: cilia crimson, with a black line, beyond this pale 
yellowish from below apex to anal angle, at apex mixed with 
black. Hindwings with hindmargin almost straight ; colour and 
lines as in forewings: cilia pale yellowish, basal third crimson, 


Sambawa; two specimens. Peculiarly distinguished 
by the straight hindmargin. 


Pyralis oenoalis, Walk. (— ealis). S.E. Borneo. 
P. fuscicostalis, Snell. S.E. Borneo. — 


P. manihotalis, Gn. Sambawa. 


Balanotis euryptera, sp. n. 

$- 16mm. Forewings triangular, hindmargin slightly rounded, 
little oblique ; pale greyish-ochreous, suffused with greenish- 
ochreous towards base ; a median series formed by a black dot on 
costa, three in disc, and one above inner margin ; second line 
fuscous, indistinct, marked witha black dot on costa and about 
four in disc ; a fuscous hindmarginal band on costa sprinkled with 
black. Hindwings fuscous-whitish ; a pale fuscous hindmarginal 
fascia and faint line before it. 


Sambawa ; one specimen. 


CRASIGENES, n. g. 
Face rounded ; ocelli present ; tongue developed. Antenne 2, 
in ¢ minutely ciliated, basal joint with a small apical tubercle in 
front. Labial palpi moderate, porrected, second joint thickened 


Pyralidina from the Malay Archipelago. 477 


with rough scales above and beneath, terminal joint very short, 
concealed. Maxillary palpi rather long, porrected, towards apex 
with rough spreading hairs. Posterior tibiz with outer spurs as 
long as inner. Forewings with 3 and 4 short-stalked, 5 approxi- 
mated, 7 and 8 out of 9, 10 closely approximated. Hindwings 
with 3 and 4 stalked, 5 approximated, 7 out of 6 near origin, 
anastomosing with 8 to about middle. 


Somewhat nearly allied to Trichophysetis. 


C. microspila, sp. n. 

&@. 12 mm. Labial palpi deep yellow, tip blackish. Fore- 
wings elongate-triangular, costa arched, apex very obtuse, hind- 
margin much rounded, little oblique; ochreous-white, more 
ochreous posteriorly ; first and second lines faintly ochreous, 
hardly traceable, curved-angulated, first marked on costa with a 
black dot, second double towards ccsta; a small black spot on 
costa at 3, and a black dot on hindmargin above anal angle, con- 
nected by a bent series of three indistinct blackish dots (con- 
spicuously marked on lower surface) ; hindmarginal line and two 
lines in cilia fuscous. Hindwings ochreous-white, becoming pale 
brownish-ochreous posteriorly ; first and second lines fine, straight, 
light brownish, on inner margin black ; hindmarginal line and two 
lines in cilia fuscous. 


Sambawa ; one specimen. 


Diplopseustis heniophthalma, Meyr. Pulo Laut. 


SICULODID. 


Addza trimeronalis, Walk. Sambawa. 

A, probolopis, sp. n. 

2¢@. 16—18 mm. Antenne whitish-ochreous, ringed with 
black. Forewings with hindmargin sinuate below subacute apex, 
bowed, oblique ; reddish-ochreous, strigulated with reddish-brown ; 
basal half much suffused with reddish-brown, limited by an an- 
teriorly ill-defined narrow dark reddish-fuscous fascia somewhat 
beyond middle, parallel to hindmargin, but rather angularly pro- 
jecting in middle, becoming obsolete towards inner margin ; costa 
with blackish scales and whitish dots ; a narrow brownish hind- 
marginal suffusion. Hindwings with hindmargin nearly straight ; 
reddish-ochreous, strigulated with reddish-fuscous ; a slightly 
darker median fascia; a reddish-ochreous-brown suffused hind- 
marginal fascia, strigulated with dark fuscous, 


Pulo Laut, 8.E. Borneo; three specimens, 


478 Mr. KE. Meyrick on 


A. syndesma, sp. n. 

@. 15 mm. Forewings with apex rectangular, hindmargin 
bowed, oblique ; whitish-ochreous ; costal edge black, interrupted 
with whitish ; an elongate-triangular brown patch, its base ex- 
tending along anterior half of inner margin, its apex resting on 
apical extremity of costa, including a blackish discal dot ; brown 
subterminal and hindmarginal streaks, confluent above middle and 
not reaching apex. Hindwings with hindmargin nearly straight ; 
whitish-ochreous ; five brownish dark-edged straight parallel 
transverse streaks ; a fuscous submarginal stria. 


Pulo Laut; one specimen. 


Stculodes emblicalis, Moore. Pulo Laut. 


S. micacealis, Walk. Pulo Laut, Celebes. 

SS. ancylosema, sp. n. 

é. 22mm. Forewings with hindmargin bowed, oblique ; pale 
brownish, indistinctly strigulated with darker brown ; about seven 
round white costal spots, containing some fuscous scales, costal 
interspaces dotted with black ; four pairs of whitish transverse 
striez connected with costal spots, posteriorly only distinct towards 
inner margin; between second and third a dark brown median 
fascia, not reaching costa, its upper extremity forming a triangular 
projection posteriorly ; a semicircular white mark below apex, 
beneath which is a dark brown spot. Hindwings with hindmargin 
irregularly bowed ; brownish-whitish, strigulated with brown ; a 
straight dark brown fascia before middle ; some dark brown marks 
towards anal angle ; a brownish transverse fascia before upper half 
of hindmargin. 


Pulo Laut; two specimens. 


S. subrosealis, Leech. Pulo Laut. 
S. imbutalis, Walk. Pulo Laut. 
S. chalcosidera, sp. n. 


$- 21—23mm. Anal prensors very long and densely haired. 
Forewings with hindmargin bowed, rather strongly oblique ; deen 
coppery-brown, strigulated with dark fuscous, more or less suffused 
with whitish in disc and towards costa ; indications of about eight 
slender darker fasciz, one in middle broader and conspicuous on 
upper half ; subterminal and marginal series of metallic-grey spots, 
and some others towards inner margin. Hindwings with hind- 
margin rounded ; coppery-brown, with dark fuscous strigule ; 
numerous scattered metallic-grey spots ; a suffused costally-broad 
median, and narrower hindmarginal fascia dark fuscous. 


Pyralidina from the Malay Archipelago. 479 


S. platyntis, sp. n. 

¢.- 15mm. Thorax with unusually broad scales. Forewings 
broad-triangular, hindmargin rounded, rather oblique; brown, 
strigulated with dark fuscous; a basal patch, partly obsolete 
towards costa, an irregular median fascia, and a hindmarginal fascia 
dark purplish-fuscous. Hindwings with hindmargin rounded ; 
brown, strigulated with dark fuscous; subbasal, median, and hind- 
marginal fascie dark purplish-fuscous. 


Pulo Laut; one specimen. 


S. pudicula, Gn. Pulo Laut, S.E. Borneo. 

S. argentalis, Walk. Sambawa, Pulo Laut. The con- 
spicuous dark blotch of the forewings is not 
nearly so broad in the Sambawa form. 

S. mochlias, sp. n. 

$2. 18—22 mm. Forewings with hindmargin rounded ; 

shining white, with scattered dark fuscous strigule; costal edge 
black, marked with white dots; anterior half of costa suffused 
with fuscous; a dark-fuscous narrow transverse mark beyond 
middle of inner margin, reaching 4 across wing ; three black sub- 
terminal dots below apex. Hindwings with hindmargin rounded ; 
shining white, with scattered dark-fuscous strigule ; a subterminal 
series of black dots, sometimes obsolete except near apex. 


Pulo Laut; two specimens. 


S. anticalis, Walk. Pulo Laut. 
S. bastialis, Walk. (?) Pulo Laut; one rather doubtful 
specimen. 


PHYCITIDA. 


Myelois pulchva, Butl. Sambawa. 
Sulebria apotomella, Meyr. Pulo Laut. 
Ephestia desuetella, Walk. Sambawa. 


GALLERIADA. 
PRASINOXENA, 0. &. 

Antenne with basal joint swollen, in ¢ above with broad concave 
horny projection, forming an eye-cap. Labial palpi in ¢ short; 
curved, in ? moderately long, porrected. Forewings with 7 and 8 
out of 9, 10 absent. Hindwingsin ¢ with inner margin narrowly 


480 Mr. E. Meyrick on Pyralidina from Malay. 


folded beneath, enclosing a hair-pencil ; 3 and 4 stalked or coinci- 
dent, 5 absent, 7 out of 6 near origin, anastomosing with 8 to 
beyond middle. 

A curious genus; the specimens are rather imperfect, 
hence the above description is not full, but I think 
sufficient. ‘l'ype P. monospila. 


P. monospila, sp. n. 

49. 15—16 mm. Forewings elongate-triangular, costa arched, 
hindmargin slightly rounded, oblique; bright green; a fuscous or 
dark fuscous dot in middle of disc: cilia light brown, spotted 
with blackish. Hindwings pale ochreous-yellowish. 


S.E.. Borneo ; two specimens. 


P. hemisema, sp. n. 

@. 18 mm. Forewings more elongate thar in P. monospila, 
costa less arched, hindmargin more oblique: bright green; a 
fuscous, almost interrupted bar from inner margin before middle, 
reaching half across wing: cilia yellow-brownish, spotted indis- 
tinctly with blackish. Hindwings white. 

Sambawa; one specimen. 


CRAMBIDA. 


Crambus malacellus, Dup. Sambawa. 


ash | 


XIX. Supplemental List of the Longicorn Coleoptera 
obtained by Mr. J. J. Walker, R.N., F.LS., 
during the voyage of H.M.S. “ Penguin,’ wnder 
the command of Captain Moore, R.N. By Caar.es 
J. Ganan, M.A., FES. 


[Read May 2nd, 1894.] 


Tue Longicornia collected by Mr. Walker in Australia 
and 'asmania are enumerated in a list which was 
published in the Transactions cf this Society for 1893. 
In order to complete the account of the beetles of this 
family which were obtained by Mr. Walker during the 
recent voyage of H.M.S. ‘‘ Penguin,” I now give a 
list of the species taken at the other localities which 
were visited. A few species, taken at Damma Island, 
are, however, omitted from the list, and will be described 
in a separate paper. Full references are only given 
for those species which have been described since the 
publication of the Munich Catalogue, and for one old 
Fabrician species which has been omitted from that 
work. ‘lhree species are described as new. 


PRIONIDH. 
1. Hurypoda antennata, Saund. 
One example; taken at Nimrod Sound, Chusan 
Archipelago. 
2. Algosoma marginale, Fabr. 
Hong Kong and Kowloon (8. B. T. Skertchley) ; two 


examples. 
3. Algosoma sinicum, White. 
Shanghai; two specimens. 
4, Philus antennatus, Gyll. 
Hong Kong; three examples. 


CERAMBYCIDH. 
5. Xystrocera globosa, Oliv. 
Shanghai; one example. Haining; one example. 
TRANS. ENT. SOC. LOND. 1894,—PaRT III. (SEPT), 


482 Mr. C. J. Gahan’s List of Mr. Walker’s 
6. Cerambyx cantori, Hope. 
Shanghai; one example. 


7. Aolesthes aurifaber, White. 
Silam in North Borneo (Skertchley) ; one example. 


ee 


Afolesthes sinensis, Gahan, Ann. and Mag. Nat. Hist., 
ser. 6, vol. vi., p. 255 (1890). 


Hong Kong; one example. 
9. Rhytidodera bowringi, White. 
Hong Kong; one example. 


10. Gnatholea subnuda, Lacord. 
Silam, N. Borneo (Skertchley) ; one example. 


11. Ceresium raripilum, Newm. 
Silam, N. Borneo (Skertchley) ; one exampie. 
12. Oeresiwm simplex, Gyll. 
Koepang, Timor; one example. 


13. Pyresthes hematica, Pase. 
Hong Kong; two examples. 


14. Hrythrus fortunei, White. 

Luhwang I., Chusan Archipelago; one exampie. 
15. Aromia bungii, Fald. 

Shanghai; one example. 

16. Polyzonus bicinetus, Pall. 
Nimrod Sound, Chusan Archipelago ; six examples. 

17. Clytus chinensis, Chevr. 
Shanghai; one example. 
18. COlytanthus signaticollis, L. and G. 


Chusan Island ; one example. 


19. Dere thoracica, White. 
Tamason Island, Chusan Archipelago. 


Longicorn Coleoptera. 483 


20. Sternoplistes temmincki, Guér. 
One example; taken on the mainland opposite 
Tygosan Island, Chusan Archipelago. 
LaMiD%. 
21. Dorcadida bilocularis, White. 
Tasmania. ‘This species was accidentally omitted from 
the first list. 
22. Hpepeotes plorator, Newm. 
Koepang, Timor ; one female example. 
23. Psacothea hilaris, Pasc. 
Haining. 
24. Monohammus tesserula, White. 


Shanghai, and Luhwang Island, Chusan Archipelago. 


25. Monohammus luridus, Pasc. 


Mainland opposite Tygosan Island, Chusan Archi- 
pelago. 


26. Cyriocrates horsfieldi, Hope. 
Chusan Island. 
27. Melanauster chinensis, Forst. 


Hong Kong, Shanghai, and Pwanche Island, Chusan 
Archipelago. 


28. Melanauster glabripennis, Motsch. 
Haining; one example. 
29. Aristobia hispida, Saund. 
Tygosan Island, Chusan Archipelago. 
30. Blepephaeus succinctor, Chevr. 
Hong Kong. 
31. Xenolea tomentosa, Pasce. 
Samboangan. 
32. Xenolea tomentosa, Pasc., var. 


Hong Kong. The subglabrous patches on the elytra 
are blackish in colour. 


33. Batocera lineolata, Chevr, 
Shanghai. 


484. Mr. C. J. Gahan’s List of Mr. Walker’s 


34. Batocera davidis, Deyr., Ann. Soc. Ent. Fr., 1878, 
p: lot 


Pwanche Island, Chusan Archipelago; two examples 
(¢ and ¢). ‘This species has previously been recorded 
only from the interior of China. 

35. Batocera thomsoni, Javet, var. 

Silam, N. Borneo (Skertchley). In the single example 
taken there is only one spot on each elytron, but it does 
not seem to be otherwise distinct from DB. thomsoni. 

36. Apriona rugicollis, Chevr. 
Chinhae. 
37. Rosenbergia megalocephala, v. de Poll, Notes from 
the Leyden Museum, vii., p. 32, pl. 1, figs. 5 
and 5a (¢). Id., vol. ix., p. 184 (g). 
Port Darwin, North Australia ; one example. 


38. Himantocera plumosa, Oliv. 
Silam, N. Borneo (Skertchley). 
39. Gnoma propinqua, Pasce. 
Ternate, Malay Archipelago; g and 9? examples. 
40. Coptops auguralis, Pasc. 
Damma Island, Malay Archipelago ; one example. 
41. Olenecamptus bilobus, Fabr. 
Damma Island and Hong Kong. 
42. Gerania bosci, Fabr. 


Koepang, Timor. 


43. Hestima floccosa, Pase. 
Ternate. 


44, Pterolophia illicita, Pase. 
Ternate; one example. 


45. Pterolophia costalis, Pasc. 
Ternate. 


46. Pterolophia obdueta, Pasc. 
Koepang in Timor, 


Longicorn Coleoptera. 485 


47. Pterolophia, sp. 
Manila (J. Green) ; one example. 


48. Pterolophia rigida, Bates. 
China, the mainland opposite Tygosan Island; one 
example. 
49, Ropica squamosa, sp. n. 
Samboangan. 
50. Ropica, sp. 
_ Samboangan. 
51. Ropica indigna, Pase. 
Ternate. 


52. Sybra umbratica, Pase. 
Ternate. 
53. Tetraglenes insignis, Newm. 
Chusan Island. 
54, Hvxocentrus, sp. 
Shanghai; one example. 


55. Agapanthia amurensis, Kraatz, Deutsche Ent. Zeit. 
xxi. (1879), p. 115. 
Shanghai. 
56. Glenea elegans, Oliv. 
Amboyna. 


57. Glenea melia, Pasc. 
Silam, N. Borneo (Skertchley). 
58. Glenea fortunei, Saund. 


Tamaon Island, Chusan Archipelago. 


59. Glenea cantor, Fabr. 
Lamia cantor, Fabr., Mant. Insect., vol. i., p- 142. 
Hong Kong (Skertchley). 


60. Phytcecia ventralis, Bates. 
Tamaon Island, Chusan Archipelago. 


486 Mr. C. J. Gahan’s List of Mr. Walker's 


61. Oberea nigriceps, White. 
Hong Kong. 


62. Oberea inclusa, Pase. 


Tygosan Island. 


63. Oberea walkeri, sp. n. 
Hong Kong. 


64. Linda fraterna, Chevr. 


sub Hemilophus fraternus, Chevr., in Cat. Gemminger 
and Harold. = Oberea seminigra, Fairm., Ann. 
Soc. Ent. Belge, 1887, p. 134. 


Tygosan Island, Chusan Archipelago. 


65. Chreonoma basalis, sp. n. 


Hong Kong; one example. 


Ropica squamosa, sp. 1. 


Squamositate fulva et grisea induta; capite prothoraceque 
minute haud dense punctatis, hoc lateraliter paullo rotundato ; 
elytris dense sat fortiterque punctatis, utrisque postice lineolis 
duabus fuscis et maculis minutis albis duabus vel tribus signatis ; 
antennis quam corpore fere sesqui-longioribus, articulo 1° punc- 
tato, fulvescente-pubescente, ceteris fuscis, cinereo-maculatis. 
Long. 53—7 mm. 

Hab. Samboangan. 


Dark brown, clothed with minute scales, which are, for the most 
part, fulvous-brown in colour, bat are of a greyish tint along the 
sutural area on the anterior half of the elytra. The head and 
prothorax are minutely and rather sparsely punctured. The elytra 
are thickly and more strongly punctured, with the punctures partly 
arranged in longitudinal rows, and each bearing a minute white 
seta stretched across its opening. Some whitish scales are aggre- 
gated to form two or three small spots a little beyond the middle 
of each elytron, and from these spots there pass backwards two 
very short dark-brown lines. Minute dark-brown spots appear at 
intervals along the suture and on other parts of the elytra. 


Ropica didyma, Pasc., seems to approach the present 
species most nearly in general shape and colour ; but its 
covering is of a different character, being in the nature 


Longicorn Coleoptera. A87 


of short hairs, as in the majority of the species of the 
genus, and not in the form of minute flattened scales, as 
in the present species and in R. servilis, Pasc. 


Oberea walkeri, sp. n. 

Fulva ; capite, antennis, elytris pygidioque nigris ; tibiis apice, 
tarsisque et (in mare) segmentis intermediis abdominis infuscatis : 
capite distincte sat denseque punctato ; prothorace quam longiori 
evidenter latiori, dense punctato, lateraliter medio obtuse rotun- 
dato ; elytris fortiter denseque et seriatim punctatis, per medium 
fusco-testaceo-dilutis, griseo tenuiter pubescentibus, area minima 
circum scutellum et macula sub humerum testaceis. Long. 
16—19 mm. 


Hab. Hong Kong (J. J. Walker). 


Head, antenne, elytra, and pigidium black ; thorax and under- 
side of body fulvous, with the sides of the three intermediate 
abdominal segments slightly infuscate in the male ; tarsi and distal 
portion of tibize blackish. Head distinctly punctured, with the 
punctures more thickly spread on the vertex than in front. Pro- 
thorax distinctly broader than long, thickly punctured ; slightly 
gibbous in the middle above, a little bulged out on each side, 
reaching its greatest width just behind the middle. Elytra 
strongly punctured, with the punctures arranged in closely 
approximated rows; a slight testaceous tint appears along the 
middle of each elytron, and there is a distinct, but very small 
testaceous space around the scutellum ; a faint greyish pubescence 
covers almost the whole of the dorsal surface ; the apices are each 
slightly emarginate, and feebly dentate at each of the four angles. 
The sides of the breast are strongly but very sparsely punctured. 

The last ventral segment, black, with the exception of a narrow 
anterior border, bears in the male a rather broad longitudinal 
channel which narrows anteriorly ; in the female this segment has 
near the apex a narrow triangular depression, which is continued 
anteriorly as an impressed line. The male is further distinguished 
by its slightly longer antenne, and by having the front of its head 
narrower, and covered with a more distinct greyish pubescence. 


Appears to be most nearly allied to O. japonica, Bates, 


Chreonoma basalis, sp. n. 


Fulvo-testacea, elytris (limbo basali excepto) violaceis aut 
cyaneis ; capite sparsim punctato, medio frontis in mare breviter 
sed prominente carinato: prothorace sat dense punctato, supra con- 
vexo, lateraliter medio rotundato; elytris sat dense punctatis ; 


488 Mr. C.J. Gahan on Longicorn Coleoptera. 


antennis articulis 4—6 ultimis nigro-fuscis. Long. 10—11. Lat. 
3}—4 mm (4 @). 
Hab. Hong Kong (J. J. Walker and J. C. Bowring). 


Fulvous ; the elytra violet or deep blue, with a very narrow 
fulvous band at the extreme base reaching from the scutellum 
to the outer margin. Antenne with the last four to six joints 
blackish ; those of the male about as long as the body, those 
of the female somewhat shorter. 


The front of the head in the male bears in the middle 
a short tuberculiform carina. In this and other charac- 
ters the species seems to come nearest to C. frontalis, 
Gahan. 


Glenea cantor, Fab. 
Lamia cantor, Fab., Mant. Ins., vol. i., p. 142. 


Hab. Hong Kong (Skertchley), and China (? Hong 
Kong) (Bowring). 


This species is omitted from the Catalogue of Gem- 
minger and Harold. It is nearly allied to Glenea spilota, 
Thoms., from which it may be distinguished by the 
following characters :— 


The elytra are usually of a pale yellowish colour, only excep- 
tionally offering a slight pinkish tint. The shoulders are pitchy- 
black, slightly margined behind with red. The foremost of the 
two black spots placed near the apex of each elytron extends in an 
almost directly transverse direction from the outer margin to near 
the suture. In spélota there are two spots (sometimes united) 
occupying a similar position, but placed more obliquely. In G. 
cantor the entire hind legs and the tarsi of the two anterior pairs 
are blackish in colour. 


( 489 ) 


XX. Descriptions of a new species of Raruuia, L., and of 
three new species of ‘lrichoptera from the Balkan 
Peninsula, with critical remarks on Panorpa 
gibberosa, McLach. By Professor Franz 
Krapitsk, F.E.S. 


[Read June 6th, 1894.] 
PuaTE X. 


In the second half of July and the first half of August, 
1893, I made a journey in Bulgaria and Hast Roumelia, 
the results of which are published in the Magazine for 
Art and Science, edited by the Bulgarian Ministerium for 
Cultus and Education. ‘To make the descriptions of the 
new species also accessible to Non-Slavic Naturalists, 
I have the permission of the editors of that magazine to 
publish them simultaneously in a foreign periodical, 
and having the honour to be a member of the Entomo- 


logical Society of London, I take the liberty to present 
the English text. 


Raphidia rhodopica, n. sp. 

Head black, oval, narrowed gradually to the collum, deeply 
punctate, the punctures having an appearance of transversal ridges. 
Front black, shining. Ocelli distinct. Antenne at the base tes- 
taceous, darker to the apex, which is wholly black. The smooth 
median longitudinal space on the collum and occiput black, with a 
distinct longitudinal impressed line. Labrum fuscous, testaceous 
on the margins. Mandibule testaceous, blackish at the tips. 
Palpi maxillares fuscous, gale testaceous. Labium fuscous, with 
exception of two very distinct yellow, triangular spots. Palpi 
labiales shining black. The underside of the head black, more 
sparingly punctured than above, with a longitudinal groove in the 
middle, which becomes broader towards the mouth. Collum 
moderately long, a little narrower behind. 

Prothorax ith shorter than the head anc collum, narrowed in the 
first third, broader behind. Pronotum punctured, and transversely 
rugose, with short pubescence, black, the anterior edge narrowly 
yellowish and the hind angles yellow. Mesothorax black, on the 
-notum anteriorly with a distinct rhombic yellow spot; on the sides 

TRANS. ENT. SOC. LOND. 1894.—parRT ly. (DEC.) 21 


490 Prof. F.. Klapalek’s descriptions 


before the base of the wings is a small oblong yellow spot ; 
the scutellum has in the middle a yellow point, hinder edge nar- 
rowly reddish-brown. 

Coxe and trochanters black ; on the anterior and middle femora 
the base and outside, except a short space on the apex, is black, 
the innerside, except the base and the apex, yellowish-brown. 
Posterior femora with exception of the yellowish-brown apex 
black. Tibie yellowish-brown, tarsi yellowish-brown, fuscescent 
at the apex. Abdomen shining black, with a narrow yellow lateral 
line ; the intermediate segments narrowly margined with yellow 
posteriorly at the sides, and on the middle of the posterior margin 
beneath. The 7th abdominal segment short, the 8th divided into 
a dorsal and ventral valve ; the first of them is longer than the 
foregoing segment, quadrangular if viewed from the side, under 
the thickened side margins swollen on either side into a shining 
black smooth hemispheric lobe ; the ventral valve, directed down- 
ward, has the apical margin notched, thickened and fuscous. 

On the innerside of this valve lie two testaceous crotchets 
directed against the apical margin and slightly curved inwardly. 
Under these crotchets lies the penis, which is canaliculated beneath. 
The 9th segment is widely opened and slightly notched on the 
hindmargin, which is yellow ; the hinder angles strongly thickened 
and beset with black spines. 

Anterior wings three times as long as broad, with elliptical apex, 
pellucid. Two cells along the radius from the origin of the 
sector and three discoidal cells. Costal area dilated in the first 
third, with 11-12 simple veinlets. Subcosta joins the costa at a 
distance from pterostigma equal to two-thirds of its anterior mar- 
gin. Pterostigma trapezoid, dark-brown, with a single simple 
veinlet parallel with its outer margin ; its inferior margin scarcely 
half the length of the superior. First discoidal cell begins at the 
same distance before the pterostigma as is the termination of the 
subcosta, and extends very little beyond its hindmargin. Four 
cubital cells. First apical vein simply furcated ; the second thrice 
furcated ; the third divided into two branches ; the fourth furcated 
twice ; and the two next following are divided into three branches. 

Posterior wings in the costal area with 8-10 transverse veinlets. 
Cubitus with one branch anteriorly, and in the area between the 


sector radii and the anterior branch of cubitus there is only one 
transverse veinlet. 


2 unknown. 
Exp, 23°5-25 mm., long. 11°5 mm. 


By the structure of wings R. rhodopica is near to R. 


of new species of Raphidia and Trichoptera, Sc. 491 


schneider’, Ratzb. and R. sericea, Albarda, from which it 
differs by the dark brown pterostigma, by the relative 
length of the first discoidai cell, and by the form of the 
genital parts. These are similar to those of L’. major, 
Burm., from which it differs by the single cross veinlet in 
the pterostigma and by four cubital cells. 

I collected two ¢ on bushes along the road from 
Philipopol to the monastery Bela Cerkva, 30th July. 


Polycentropus excisus, n. sp. 

Similar to P. flavomaculatus, Pict., in general appearance, but 
the antennz are yellow, the apex only annulated indistinctly with 
brownish. Palpi and legs yellow, and the markings on the anterior 
wings are pale yellow; the footstalk of the apical fork No. 3 is at 
least half so long as the fork. 

In the ¢ the dorsal plate is almost quadrangular, slightly 
excised on the apex. Superior appendages a little longer than the 
dorsal plate, broad. Viewed from the side their upper margin is 
rounded and the lower almost straight. From above they are quad- 
rangular, with rounded exterior apical angle. Their base is covered 
above by a large triangular lobe joined to the base of the dorsal 
plate. Intermediate appendages short, reddish-brown, escaping 
from beneath the apical angles of the dorsal plate, curved down, 
but their tip turned again upward. Inferior appendages a little 
shorter than the superior, concave inwardly and convex outwardly 
with deeply excised apical margin. Both the upperand lower edge 
are so far turned inwardly that they touch each other, forming, in 
this way, a tube opened at the apex. 

¢ unknown. 

Exp. 15 mm., length 5 mm. 

I would not have dared to describe this species from 
the single ¢ collected in Bojana, near Sophia, 20th July, 
had not the form of the appendages been so very cha- 
racteristic. 


Tinodes unidentata, n. sp. 


Body black, clothing black mingled sparingly with testaceous. 
Antenne fuscous, very indistinctly and narrowly annulated. Legs 
brown or fuscous, trochanters and knees testaceous. Anterior 
wings blackish, sparingly covered with black pubescence. Neura- 
tion distinct, similar to that of TY. dives, Pict. ; posterior wings 
similar in colour to the anterior, not iridescent. Fringes blackish 


grey. 


492 Prof. F. Klapalek’s descriptions 


In the ¢ the dorsal plate from side is elongate, slightly convex 
above at the base, with parallel upper and lower margins, 
the lower being obliquely arched upward at the apex. Superior 
appendages 22 as long as the dorsal plate, slender, slightly dilated 
towards the base, slightly curved downward and covered with 
blackish hairs. Intermediate appendages curved downward, 
bearing on the apex short thin spines and furnished on the 
underside before the apex with few slender long backward spines 
and with a feeble spine arising from the underside and curved 
downward, but with the apex again turned upward in form of the 
letter S. Inferior appendages two-jointed, 1st joint short, 
quadrangular, the 2nd much broader, elongately quadrangular, 
bearing on the middle of the apical edge a strong upwardly curved 
tooth. The exterior of these appendices, except the dorsal and 
apical margin of the second joint, is covered with black fine hairs, 
the apex of the second joint is furnished with dense straight strong 
sete, which almost conceal the apical tooth. Between the base 
of these appendages arises the shining, reddish-brown, spiniform 
penis, which is arcuate and with its apex turned downward 
between the appendices. The colour of all appendices in speci- 
mens preserved in alcohol is testaceous except the basal half of the 
inferior appendages, which is fuscous; in dry specimens the colour 
is fuscous, the apex of inferior appendages only being shining 
testaceous. 

? similar to that of J. dives, Pict. Exp. ¢ 9°5—11 mm.; 
Q 11—12 mm. 


This species, most resembling 7’. dives, Pict., is easily 
distinguishable by the uniform blackish wings, the 
slender superior appendages, the simple spiniform penis 
and the single tooth on the apical edge of the inferior 
appendages of the ¢. . 

frequents the rapid streams on the slopes of the 
Vitosa Mountain near Sophia, Bojana, 20th July. 


Rhyacophila obtusa, n. sp. 

Fuscous, head and pronotum clothed with golden yellow hairs. 
Antennw uniformly fuscous, with dense short pubescence. Palpi 
fuscous. In the legs the coxe are fuscous, the rest testaceous, 
tarsi fuscescent to the apex ; spurs brown and therefore very dis- 
tinct. Wings short and broad, anterior with an almost semi- 
circular apex. Membrane smoky-grey, uniformly densely covered 
with golden-brown pubescence, mingled only very sparingly with 
blackish hairs; on the posterior the pubescence is thinner and 


of new species of Raphidia and Trichoptera, Sc. 498 


black. Pterostizma of all four wings distinct greenish brown. 
Neuration distinct, dark brown; radius not furcated before the 
apex ; second apical fork extending only very little further in- 
wardly than No. 1 ; costal margin of the anterior wings furnished, 
at its base only, with plumose hairs amongst the simple. Fringes 
dark grey. 

In the ¢ the 9th abdominal segment on the dorsal and ventral 
side distinctly developed, though its ventral part is considerably 
narrower. The external lobes are very large, reaching to the 
middle of the inferior appendages, viewed from side rhomboid, 
with the lower part bent round outwardly. If viewed from above, 
the external margin appears to form a raised arcuate edge dis- 
appearing at the apex and forming an axis round which the lower 
part of the lobe is bent outwardly. Both lobes are united at the 
base, but separated for more than the apical half of their length ; 
they are sparingly furnished with stiff hairs. First joint of the 
inferior appendages oblong, convex on its lower edge; second 
joint much shorter, very obliquely truncate at the apex, so that 
the upper edge is much shorter than the lower, which is concave 
at the base, convex at the considerably produced apex ; the apical 
edge straight. Lower penis-cover not exserted, large, boat-shaped, 
fuscous, and slightly excised at the apex. Exp. 16 mm., length 
5°5 mm. 

@ unknown. 


Three ¢ at a brook in Dragalevci flowing from the 
Vitosa Mountains near Sophia, 22nd July. 

When living this species has the appearance of a pale 
Silo or Inthax, and was found sitting on the underside 
of leaves of plants on the bank. Though its radius 
is not divided at the apex, we must put it into the Section 
C. of the genus Rhyacophila (McLach., Rev. and Syn., 
p- 158), from its short and broad wings and the form of 
the anal appendices. Though the external lobes appear 
very similar to those of Rh. eatont, McLach., the lower 
penis-cover and the uniform colour of the wings show 
its relation to the group of Rh. levis, Pict., from which 
it differs especially in the form of the external lobes and 
by the presence of the plumose hairs on the costal 
margin of the anterior wings. 


Panorpa gibberosa, McLach. 
This highly interesting species was described by Mr. 


McLachlan in his Synopsis of the species of Panorpa 
occurring in Europe and the adjoining countries (Trans. 


494, Prof. F. Klapalek’s descriptions 


Ent. Soc. 1869, Part J.) from one g and 9? ; but since 
that time, as the author has kindly informed me, it has 
not been seen by him. I hope, therefore, it will not be 
out of place to give some critical remarks on it, as I have 
been so fortunate as to collect a series of specimens in 
two separate localities in Bulgaria and Hastern Roumelia. 
The most important characters in which this species 
differs from P. germanica, L., to which it is very nearly 
allied, are in the furca (“appendices segmenti 9,” 
MclLach.), whose branches are longer and have their 
apex obliquely truncate and in the larger process of the 
third dorsal segment. The wing markings are similar to 
those of P. germ mica, var. apicalis, St. Two of my 
specimens are quite typical, agreeing with the description 
and figures in all but in the dorsal process of the 3rd 
abdominal segment, which does not appear larger than 
in germanica. From this species they differ, besides the 
other characters namedabove, by the shining black occiput, 
the brown colour which covers regularly the whole occiput 
in P. germanica is limited to the hinder eye margins. 
Abdominal segments 7th and 8th marked on either side 
by a large black shining spot; furca fuscous, almost 
black at the apex, with dense strong black setz ; on the 
rostrum are two very distinct shining black lines. Another 
specimen agrees in all respects with the first two, but the 
pterostigma is only yellowish without the usual dark- 
brown spot, instead of which it has a little point at the 
base of the sector and two similar below the pterostigma. 
Two other specimens agree with the last, but have in the 
pterostigma a dark brown spot, which is smaller than 
in typical examples. A series of specimens are in the 
markings similar to P. germanica, but the anal parts are 
those of gibberosa. One most aberrant example is 
smaller, very dark, having the 7th and 8th abdominal 
segments wholly shining black ; the wings are transparent, 
very shining, with a slight greenish and brownish tinge ; 
the spots are conspicuous, and, besides the above- 
named, we find two dark brown points below the middle 
of the anterior margin of the wings. A series of ? 
show no difference from those of P. germanica collected on 
the mountains of the Bohemian frontier. They are very 
dark, and with conspicuously marked wings. I do not 
hesitate to identify these examples, at least the first named 
five, with P. gibberosa, McLach., and I am inclined to 


of new species of Raphidia and Trichoptera, Sc. 495 


hold this species distinct from P. germanica, but to have 
absolute certainty, we must wait for more materials ; 
it would be especially desirable to ascertain whether 
the typical P. germanica occurs in those regions ; the 
season when I visited them was rather advanced. The 
difference between P. germanica and gibberosa is indeed 
very slight, but yet greater than between P. communis, 
L., and P. vulgaris, Imhoff, which, after all, are only 
two forms of one species. 


EXPLANATION OF PLATE X. 


Fics. 1—4. Raphidia rhodopica, n. sp. 1. Wings. 2. Head. 
3. Apex of the ¢ abdomen from side. 4. The same from 
behind. 


Fras. 5 & 6. Panorpa gibberosa, McLach. 5. Terminal segments 
beneath. 6. Apex of abdomen from side. 


Fias. 7 & 8. Polycentropus excisus, n. sp.7. Apex of the abdomen of 
¢ from above. 8. The same from side. 


Fic. 11. Tinodes unidentata, n. sp. 9. Apex of abdomen of the ¢ 
from above. 10. The same from side. 11. Apex of 
abdomen of the 9. 


Figs. 12 & 13. Rhyacophila obtusa, n. sp. Apex of abdomen of the 
dé. 12. From above. 13. From below. 


[Figs. 1, 2, enlarged *; Figs, 3, 4, 12; Fig. 5, 4° ; Fig. 6, 2; Figs. 
7, 9, 10; 32 = Figs, 8, 11; 12; aud 13, 40. All ques with 
camera lucida. ] 


GUA Ee) 


SOT, A Monograph of British Braconide. Part V. By 
the Rev. Tuomas A. Marssatt, M.A., F.E.S., and 
Member of the Société Entomologique de France. 


[Read March 28th, 1894 ] 
Plates XI. and XII. 


V. EXODONTES. 
XXIV. ALYSIIDES. 


Heap large, with dilated cheeks ; occiput concave, not margined. 
Mandibles broad, flattened, tridentate, or quadridentate, usually 
wide open after death, and often carried in that position by the 
living insect, even in repose ; they project on each side of the head 
when inspected from above. Antenne multiarticulate, often very 
long. Maxillary palpi generally with 6, labial with 3, joints, but 
the number is variable. Thorax oval, convex; prothorax short; 
furrows of the mesonotum usually inchoate, denoted by two 
humeral impressions ; often obsolete ; less frequently complete. 
Three cubital areolets in the forewings, the first sometimes con- 
founded with the second, or with the first discoidal ; stigma 
variable in form, oval or subtriangular, more or less elongate and 
linear, or disappearing altogether in the genus Aspilota, where it 
is confounded with the metacarp; posterior wings often furnished 
with a pobrachial transverse nervure. Both sexes are apterous 
in the genus Chasmodon; the @ of Panerema has rudimentary 
wings, the ¢ is unknown ; in Allwa the wings of the ¢ are very 
short and narrow, with all the nervures and areolets contracted ; 
those of the ? are generally, but not always, complete. Abdomen 
sessile, subsessile, or subpetiolate, longer than the thorax, more or 
less depressed, seldom compressed, as in Panerema, Mesocrina, and 
Aspilota ; Ist segment rugulose, the rest almost always smooth 
(except Trachyusa) ; suturiform articulation effaced, or barely 
visible ; segments 2-3 taken together longer than all the following. 
Terebra exserted, of variable length, short, and almost concealed 
only in Adelura. 


The above characters are equally applicable to the 
next tribe, Dacnusides, except that these have only two 
TRANS, ENT. 80C. LOND. 1894,—ParTIv. (DEC.) 


498 Rev. T. A. Marshall’s Monograph of 


cubital areolets in the forewings, thus standing in the 
relation of secondary forms, like Apanteles as compared 
with Microgaster. The analogy of the two tribes is so 
close that their sepsration, as divisions of equal value to 
that of preceding tribes, is merely a matter of taste. The 
older writers down to «and including Haliday, regarded 
Dacnusa as a subsection of Alysia. The latter genus was 
first distinguished by Latreille in 1805, having for its 
type A. stercoraria, Latr., previously named Ichnewmon 
manducator by Panzer. Only two subsequent authors 
have attempted to describe the numerous allied species ; 
Nees v. Hsenbeck published 41 in six sections, and his 
monograph was greatly extended and improved by 
Haliday in the fifth volume of the “ Entomological 
Magazine,” and in a separate tract entitled, ‘ Alysia, 
Fasciculus alter,” published in 1839, and containing the 
Dacnuside. Haliday’s memoirs are so exhaustive, as far 
as relates to Great Britain and Ireland, that very few 
fresh discoveries have since been made, and his descrip- 
tions render most of the insects unmistakable. On the 
Continent nothing has appeared since the date of Nees v. 
Esenbeck’s work, 1834, except a few scattered notices, 
and Forster’s ‘Synopsis of Genera.” This last work is 
valuable as giving names to all the sections pointed 
out by Haliday, which have now become genera, in 
accordance with the modern conception of the term 
genus. But Forster’s peculiar penchant for multiplying 
genera, carried him, as usual, too far, and many of his 
divisions, especially those adapted for one species only, 
may be conveniently suppressed. The writings of 
Wesmael, which have contributed so much to the sub- 
ject of the present series of papers, unfortunately stop 
short at the end of the Optides. His collection contains 
two or three hundred Alysiids, which he would doubt- 
less have published if his health had permitted; these 
I have seen, but as every specimen required an elaborate 
process of cleaning, which I had no permission to apply, 
with small chance of good results, I was obliged to 
content myself with taking a few notes. 

The Alysiides seem to be almost exclusively parasites 
of Diptera, and especially of the fungivorous Tipulide 
and Muscidz ; the few observations which connect them 
with other orders of insects will be mentioned in their 
places ; they are all liable to a suspicion of error. 


British Braconide. 499 


The determination of species is in general not difficult, 
owing to the obvious characters exhibited by the wing- 
veins ; in other respects there is great uniformity. The 
colours are black and piceous brown, relieved in some 
cases with red or testaceous, and are tolerably constant 
in the same species. As in the case of the Opvides, 
good setting is a necessity, especially in the smaller 
species; the characters are mostly derived from the 
upper surface, but the face and the mesopleurz should 
be open to examination. Pinned specimens, and all 
those whose wings cannot be clearly seen, are generally 
valueless. 


TABLE OF GENERA. 


1. Both sexes wingless... co sks — ... 1, CHASMODON. 
2. Both sexes winged; the wings rarely rudi- 
mentary (Panerema), or shortened (Allea). 
(4) 38. Wings of the 9 very short, rudimentary, almost 
without nervures ( unknown) iil, PANEREMA. 
4. Wings ample, except in Alla@a, where they a are 
liable to be more or less shor' ened, the neura- 
tion always remaining complete. 
5. Second cubital areolet confounded with the first. xviii. ASPILOTA. 
6. Second cubital areolet separated from the first. 
(22) 7. First intercubital nervure as long as, or longer 
than, the 2nd abscissa of the radial nervure. 
8 
9 


(9) 8. First cubital areolet confounded with the Ist 
’ discoidal ... iii. SYNCRASIS. 
(8) 9. First cubital areolet separated from the Ist 
discoidal 


(11) 10. Fourth joint of theantennz longer thanthe 3rd_ x. IDIAsTA. 
(10) 11. Fourth joint of the antenne not longer than 

the 3rd. 
(18) 12. Second abdominal segment punctate, not 

shining, marked with a medial transverse 

impression .. iv. TRACHYUSA. 
(12) 18. Second abdominal “segment smooth, shining, 

without a transverse impression. 
(15) 14. Metathorax longitudinally carinated in the 

middle; wings often more or less abbreviated 

bee : ses ee sae .. V. ALLG@A. 

(14) 15. Metathorax not carinated; wings always fully 

developed. 
(17) 16. Furrow of the mesopleurz smooth or obsolete. ix. PENTAPLEURA. 
(16) 17. Furrow of the mesopleure crenate or rugose. 
(19) 18. Anal nervure interstitial ... vi. CRATOSPILA. 
(18) 19. Anal nervure springing from the middle, or 

below the middle, of the extremity of the "ond 

discoidal areolet. 
(21) 20. Stigma short, emitting the radial nervure from 


beyond the ‘middle 3 . vil. ALYSIA. 
(20) 21. Stigma elongate, emitting the ‘radial nervure 
before the middle s . Vill. TANYCARPA. 


(7) 22. First intercubital nervure shorter than the 2nd 
abscissa of the radial nervure (except in 
Adelura dictynna). 


500 Rev. T. A. Marshall’s monograph of 


(24) 23. First cubital areolet confounded with the 1st 

discoidal ... . xi APHERETA. 
(23) 24. First cubital areolet separated. ‘from the Ist 

discoidal. 
(26) 25. Fourth joint of the antenne longer than the 3rd; 

pobrachial areolet of the hindwings less than 

half as long as the preebrachial ... Xi PHENOCARPA. 
(25) 26. Fourth joint of the antenne not longer than th 

8rd; pobrachial areolet of the hindwings half 

as long as the prebrachial, or longer. 
(28) 27. Stigma obsolete, or nearly so, not distinguished 

from the metacarp ; 2nd cubital areolet some- 


times confounded with the Ist.. . xviii. ASPILOTA. 
(27) 28. Stigma distinct, evidently stouter than the 
metacarp. 


(82) 29. Stigma cuneiform or oval; not linear, 7.e., not 
of equal breadth throughout. 
(31) 30. Stigma elongate, cuneiform, emitting the radial 


nervure before the middle aa xy. PROSAPHA. 
(30) 31. Stigma oval-lanceolate, short, emitting the 
radial nervure from the middle.. ... xvi. MESOCRINA. 


(29) 82. Stigma linear, of equal breadth throughout. 
(84) 33. Radial nervure originating at the extreme base 

of the stigma ses xiv. ANISOCYRTA. 
(83) 35. Radial nervure originating at any other point 

of the stigma. 
(86) 34. Anal nervure interstitial or nearly so; terebra 

very short, almost concealed... xiil. ADELURA. 
(35) 36. Anal nervure springing from the middle of the 

extremity of the 2nd discoidal areolet ; terebra 

exserted, falcate ... “oc ae Bs . XVil. OrTHosTIGMA. 


1. Caasmonon, Hal. 
Hal., Ent. Mag., i., 486 (1833). 


Apterous in both sexes. Head flattened ; eyes minutely pubes- 
cent ; maxillary palpi 6-, labial 4-jointed. Second joint of the 
flagellum longer than the first (as in Genus xii.. Phanocarpa). 
Thorax much narrower than the head, compressed, subcylindric ; 
sutures of the mesonotum distinct, complete, crenulate. Abdomen 
@ oval, as long as and broader than the thorax ; belly compressed, 
obliquely truncate at the extremity ; abdomen ¢ narrower, some- 
what depressed. Terebra exserted. 


1. Chasmodon apterus, Nees. 


Bassus apterus, Nees, a Ges. Berl., 1814, p. 207. 
Alysia aptera, Nees, Mon., .» 264, d. 
CO. apterus, Hal., Ent. Maes Ve; 214, Or: 


Smooth, shining ; ferruginous, with the head and extremity of 
the abdomen blackish. Mandibles tridentate. Palpi ferruginous. 


British Braconide. 501 


Antennz ferruginous, darker towards the tips, 17-21-jointed, as 
long as the body in the 9, longer and more slender in the ¢. 
Scutellum minute, tuberculiform ; mesopleure with a rugose 
furrow ; metathorax punctato-rugose, subcarinate, rather darker 
than the rest of the thorax, truncate posteriorly. Beneath the 
tegule two small membranous lobes represent the wings. Legs 
ferruginous. First abdominal segment obconic, finely striolate, 
subcarinate at the base. Terebra straight, issuing from the lower 
end of the apical truncature of the abdomen, and when measured 
from its base, half as long as the abdomen. Length 3-14 lin. 

Var. g deep brown, with the anterior part of the mesothorax and 
the base of the abdomen paler. Head black ; oral parts, base of 
antenne, and legs, ferruginous. 


Rare; I possess but two examples. The insect has 
occurred in England, Ireland, and Scotland; on the 
Continent it has only been noticed in Germany and 


Holland. 


il, Panerema, Forst. 
Forst., Verh. pr. Rheinl., 1862, p. 263. 


Male unknown. Wings rudimentary, showing but few of the 
nervures. Head somewhat hemispherical; eyes bare; palpi 
broken, in my specimens. Second joint of the flagellum shorter 
than the 1st. Thorax narrower than the head, short, sub- 
cylindric, compressed ; furrows of the mesonotum obsolete ; the 
middle of its. disk marked with a depression. Abdomen com- 
pressed, linear when viewed from above, longer and narrower 
than the thorax, almost vertically truncate at the extremity. 
Terebra exserted. 


1. Panerema inops, Forst. (Pl. XI, fig. 1.) 
P, inops, Forst., l. . 


Q. Black, smooth, and shining; abdomen reddish, except the 
first segment which is black, and the apex which is blackish. Man- 
dibles tridentate, black ; clypeus and palpi dusky. Antenne red, 
each joint narrowly dusky at the extremity ; rather stout, a little 
longer than the body, 24—25-jointed ; the joints are progressively 
shorter from the base, and become moniliform towards the 
extremity. Furrow of the mesopleure smooth; scutellum very 
small, preceded by a fovea larger than itself ; metathorax coarsely 


502 Rey. T. A. Marshall’s monograph of 


rugose, without a medial carina. Forewings as long as the meta- 
thorax, subtriangular, furnished with a stigma near the extremity, 
3 nervures and 4 areolets (see Plate) ; hindwings nerveless, except 
a portion of the radial nervure visible towards the extremity. Legs 
red, rather stout and long. First abdominal segment nearly linear, 
hardly narrower at the base, deeply striate, the spiracles placed in 
the middle ; the following segments smooth and shining ; 2nd 
suture effaced ; apical segments very short. Terebra short, falcate, 
ascending. Length, 14 lin. 


This species was discovered to be British by Dr. 
Capron, who took several specimens in a sand-pit at 
Shiere in Surrey: the description is made from two of 
these, which he kindly gave me. 


ii. Syncrasis, Forst. 
Forst., Verh. pr. Rheinl., 1862, p. 264. 


Palpi short, the number of joints differing in the two species. 
First joint of the flagellum longer than the second. Metathorax 
carinated. Radial areolet lanceolate, not reaching the tip of the 
wing ; radial nervure curved ; 2nd abscissa shorter than the 1st 
intercubital nervure ; 2nd cubital areolet not broader than long, 
measured along tbe cubital nervure ; anal nervure interstitial. 


The species are unknown to me; they are thus dis- 
tinguished by Haliday :— 


Maxillary palpi 4-, labial, 3-jointed me ~.- L. fuctcola, Hal: 
Maxillary palpi 2-, labial 2-jointed a3 .; 2. Halidan, Forst. 


1. Syncrasis fucicola, Hal. 


Alysia fucicola, Hal., Ent. Mag.,v., 217, 3 2, pl. xvii., 
f. 9 (wing). 

Maxillary palpi short, dusky. Body deep black, shining ; man- 
dibles reddish. 9 Antenne 17-jointed, scarcely as long as the body. 
No punctiform impression before the scutellum; furrow of the meso- 
pleurz faintly rugulose ; metathorax short, uneven, rugulose, with 
two smooth dorsal spaces. Wings dull hyaline ; squamule fuscous ; 
stigma blackish, narrow, emitting the radial nervure from its middle ; 
2nd discoidal areolet entirely open at the end ; hindwings very 
narrow, without a transverse vein or axillary areolet. Legs dull 
testaceous with the femora and tibie dusky, except the base of 
the latter. Abdomen oboval; 1st segment striolate, obconic. 
Terebra as long as 2 of the abdomen, ¢ Similar ; the antennz 


3 


longer, 19-jointed. Length, 1 ; wings, 14-2 lin. 


British Braconide. 503 


It has only been found in Ireland; common, according 
to Haliday, in dry sea-weed on the coast. 


2. Syncrasis Halidaii, Forst. 


Alysia fuscipes, Hal., Ent. Mag., v., 217, 2 (not of 
Nees). 

Phenolyta fuscipes, Forst., Verh. pr. Rhainl., 1862, 
p- 264. 


@. Palpi shorter than in the preceding, as also are the antenne 
and the general form. Legs blackish. Abdomen suborbicular, 
depressed. Terebra hardly visible. g unknown. Length, 2 
wings, J} lin. 


No more is said of this species, except that it resembles 
fucicola, with which it was taken rarely ; found also once 
in the London district by Walker. The synonymy given 
by Haliday in the Ent. Mag. is erroneous ; Alysia fuscipes, 
Nees, belongs to Genus x1. Aphereta. To avoid confusion, 
Forster changed the name of the present insect to 
Halidaii, and also founded upon it his uncharacterised 
genus Phexnolyta. 


iv. Tracayusa, Ruthe. 
Ruthe, Stett. Zeit. 1854, p. 352. 


Palpi of ordinary length, maxillary with 6, labial with 4 joints. 
Two first joints of the flagellum of nearly equal length. Meta- 
thorax not carinated. Radial areolet cultriform, narrow, reaching 
the tip of the wing; radial nervure straight ; 2nd abscissa shorter 
than the 1st intercubital nervure ; 2nd cubital areolet twice as 
broad as its length, measured along the cubital nervure ; anal 
nervure not interstitial ; stigma large, oblong, obtuse at both ends, 
emitting the radial nervure from its outer half. Second abdo- 
minal segment punctulate, dull, with a transverse impression in the 
middle. .'lerebra hardly exserted. 


The elegant species which forms this genus seems 
rightly separated from all others; its form is more 
elongate, and resembles, as Haliday has remarked, that 
of the Cyclostomous genus Colastes; his observation 
refers, no doubt, to Xenarcha lustrator (Ent. Tr., 1885, 
p- 50). The name Trachyusa appeared to Forster too 


504 Rev. T. A. Marshall’s monograph of 


much like Trachusa, Jur., a genus of bees, and he 
therefore invented a new name, Cosmiocarpa. Trachusa, 
however, is not adopted by hymenopterists, as far as I 
know, and, besides, it is not the same as T'rachyusa. 
For these reasons I have restored the name given by 
Ruthe. 


1. Trachyusa aurora, Hal. (PI. XI., fig. 2, 3). 


Alysia aurora, Hal., Ent. Mag., v., 217, $2). 
Trachyusa nigriceps, Ruthe, Stett. Zeit., 1854, p. 352. 


Variable, usually testaceous, with the head, the metathorax, and 
the base of the abdomen, black. @ Head shining ; mandibles tes- 
taceous ; palpi very pale. Antenne slender, longer than the body, 
blackish with the base testaceous, 30-34-jointed. Furrows of the 
mesonotum converging to a fovea situated before the scutellum ; 
pleuree somewhat dusky ; metathorax rugulose. Wings hyaline ; 
squamule and stigma testaceous, the latter almost orange-coloured ; 
posterior angle of the 2nd cubital areolet produced ; 2nd discoidal 
areolet incompletely closed at the end ; recurrent nervure hardly 
rejected. Legs testaceous. Abdomen linear, slightly clavate, 
depressed ; Ist segment short, not much narrowed at the base, 
rugulose, blackish or fuscous, either at the base or entirely; 
2nd thickly punctulate and dull, except at the extremity, trans- 
versely bisected by an impressed line; the following segments 
punctulate at the base. ‘Terebra very short. ¢ Stigma black ; 
posterior abdominal segments blackish. Length, 14-1? ; wings, 
23-33 lin, 

Var. 1. Mesonotum fuscous. 

Var. 2. ¢ Black; underside of scape, mandibles, legs, and abdo- 
men, testaceous ; Ist abdominal segment blackish. Intermediate 
varieties also occur, 


A rare species; taken in England, Ireland, Germany, 
and Holland. I have only once met with it in England, 
but Dr. Capron obtained several at Shiere. 


v. Atna@a, Hal. 
Hal., Ent. Mag., i., 265 (1833). 


Maxillary palpi 6-, labial 4-jointed. First joint of the flagellum 
longer than the 2nd. Metathorax carinated. Wings of the ? 
usually as long as the body, but sometimes shortened more or less ; 
radial areolet large, cultriform, reaching the tip of the wing ; radial 


British Braconide. 505 


nervure straight, its 2nd abscissa shorter than the Ist intercubital 
nervure ; 2nd cubital areolet twice as broad as long, measured on 
the cubital nervure; anal nervure interstitial ; stigma narrow, 
lanceolate, emitting the radial nervure from the middle ; 2nd dis- 
coidal areolet contracted, indistinct ; recurrent nervure hardly 
rejected. Wings of the ¢ not longer than the thorax, very 
narrow, with the stigma much enlarged, the nervures thickened, 
the areolets deformed and contracted (see Plate). Second abdo- 
minal segment smooth. Terebra almost concealed. 


Imperfection of the organs of flight, among the 
Hymenoptera, usually shows itself in the ?, but here 
the rule is reversed; the ¢ has always shortened wings, 
unfit for flight, the 2 is but rarely in that condition. We 
are not able to assign any probable cause for this pecu- 
liarity : if $s with perfect wings exist anywhere, at least 
they have never been found. Férster named this genns 
Diaspasta, having apparently overlooked the name pro- 
posed long before by Haliday. 


1. Alleea contracta, Hal. (Pl. XI., fig. 3, %; fig. 3a, ¢.) 


Alysia contracta, Hal., Ent. Mag., v., 218, $ ¢, pl. xvii, 
fala oi LS: 6. (wise). 


@ Black ; head smaller than in most of this tribe, somewhat 
shining ; eyes prominent ; face rugulose ; mandibles narrow, tri- 
dentate, the two lateral denticulations minute, the middle one 
acutely prominent; clypeus, mandibles, and palpi ferruginous. 
Antenne as long as the body, ferruginous, dark at the tips, 
21-jointed, the 3rd joint elongate. Thorax rugose, with the 
middle of the mesonotum and of the scutellum smoother and 
rather shining ; furrows of the mesonotum distinct, 3 in number, 
the two lateral shortened ; metathorax dentiform behind, as viewed 
laterally. Wings hyaline; squamule testaceous; stigma and 
nervures fuscous; hindwings narrow. Legs ferruginous, Abdo- 
men depressed, spatulate, shining; Ist segment sublinear, irre- 
gularly rugose, the rest smooth, Terebra very short. g Antenne 
blackish, narrowly rufous at the base. Nervures and stigma of the 
contracted wings much darker than those of the @? ; legs rufescent, 
with the base of the hind coxze fuscous. Length, about 1 line ; 
wings 9 2$, g¢ 1} lin. 

Var. 1. 2 Mesothorax chestnut-brown in front. 

Var. 2. 2 Wings shortened more or less, the nervulation re- 
maining complete. 

TRANS. ENT. SOC. LOND, 1894.—PaRTIV. (DEC.) 2k 


506 Rev. T’. A. Marshall’s monograph of 


Only noticed in Hngland and Ireland; I find them 
in a marshy meadow near the river Lynher in Corn- 


wall, and formerly obtained them in Yorkshire and 
Wiltshire. 


vi. Cratosriua, Forst. 
Forst., Verh. pr. Rheinl., 1862, p. 265. 


Maxillary palpi 6-, labial 4-jointed. First joint of the flagellum 
longer than the Ynd. Metathorax not carinated; furrow of the 
mesopleure crenulate. Radial areolet cultriform, reaching the tip 
of the wing; radial nervure straight, its 2nd abscissa one half 
shorter than the 1st intercubital nervure; 2nd cubital areolet 
somewhat broader than long; anal nervure interstitial ; stigma 
oblong, rather stout, emitting the radial nervure beyond the 
middle ; recurrent nervure hardly rejected ; pobrachial areolet of 
the hindwings shorter than half the preebrachial. Second segment 
of the abdomen smooth. Terebra very short. 


1. Oratospila circe, Hal. 


Alysia circe, Hal., Ent. Mag., v, 219, 69, pl. xvi, 
f. 6 (wing). 


Blackish, or dark brown, with the front of the head and base of 
the 2nd abdominal segment ferruginous. @ Head shining, ferru- 
ginous, the border of the occiput and middle or greater part of the 
vertex blackish ; mandibles paler fuscous ; palpi long, whitish. 
Antenne slender, longer than the body, 30-34-jointed, pale rufous 
near the base, blackish towards the tips; 3rd joint very long. 
Thorax dark chestnut-brown, shining ; prothorax rufous ; furrows 
of the mesonotum punctulate, converging to a small fovea in front 
of the scutellum ; metathorax blackish, punctato-rugulose, Wings 
hyaline ; squamulze ochreous ; stigma and nervures pale fuscous, 
the latter becoming obsolete towards the extremity of the wing. 
Legs slender, pale rufous. Abdomen spatulate, subpetiolate, 
blackish ; 1st segment narrow, obconic, faintly striolate, with 
scarcely visible tubercles; the following segments smooth ; 2nd 
ferruginous or chestnut-coloured anteriorly ; apical segments and 
belly pale. Terebra very slightly exserted. ¢ First abdominal 
segment almost linear. Length, 14; wings, 22 lin. 


Taken by Walker in the London district, but very 
rare ; it has not occurred since the time of Haliday. 


British Braconide. 507 


vii. Atysia, Latreille. 


Latr.. H. N., xiii, 177 (1805) ; Alysia, Strophea, and 
Goniarcha, Forst., Verh. Pr. Rheinl., 1862, p. 265. 


Second joint of the flagellum not longer than the Ist, which is 
also short; face short, usually rugulose; clypeus very small ; 
mandibles broad. Furrows of the mesonotum abbreviated in most 
of the species, the point of their convergence indicated by an 
oblong fovea, which is not to be confounded with the ordinary 
antescutellar fovea ; this latter is deeply sunk, crenate, or striate ; 
metathorax flattened, sometimes carinated in the middle, and 
rugulose ; furrow of the mesopleure crenate or rugose. Stigma 
distinct, oblong, emitting the radial nervure generally from its 
posterior or outer half; anal nervure not interstitial ;.2nd cubital 
areolet never larger than the 1st; 1st intercubital nervure as long 
as the 2nd abscissa of the radial nervure, or longer ; Ist cubital 
areolet separated from the 1st discoidal ; pobrachial areolet of the 
hindwings half as long as the prebrachial. Abdomen somewhat 
depressed ; Ist segment oblong, striolate, with median or sub- 
median tubercles, from which to the base it is gradually narrowed; 
this segment is generally about one-half longer than its apical 
breadth ; if longer still, and more slender, it makes the abdomen 
subpetiolate. Terebra slightly exserted. 


TABLE OF SPECIES. 


(6) 1. Antenne rather stout, sometimes shorter 
than the body; anal nervure springing 
from the lower angle of the 2nd dis- 
coidal areolet; recurrent nervure re- 
jected; radial areolet somewhat lanceo- 
late or acuminate, not reaching the 
extremity of the wing (Alysia and 
Strophea, Forst.). 

. Spiracles of the metathorax large, distinct, 

and margined nee ee ate Bae 

. Spiracles minute, concealed by the rugosity 
of the metathorax, not margined. 

. Antenne @ 40-jointed (g unknown) ... 2. soror, Marsh. 

. Antenne ¢ 9 22-24-jointed ... 9 rufidens, Nees. 

. Antenne slender, elongate ; anal nervure 
springing from the middle of the ex- 
tremity of the 2nd discoidal areolet ; 
recurrent nervure nearly interstitial, 
rarely evected ; radialareolet cultriform 
with sulparatlel sides, alwost reuching 
tie extremity of the wing (Goniarcha, 
F Grst. ) 

(8) 7. Hind femora sinvated at the base, incras- 
sated, compressed ... Bk are a 

7) 8. Hind femora of the usual form. 
(10) 9. Wingsinfumated ... tre “Bi ... 9d. fuscipennis, Hal. 
(9) 10. Wings hyaline. 


1. manducator, Panz. 


— 
or 
— 

aom>F w bp 


4. loripes, Hal. 


508 - Rev. 'T. A. Marshall’s monograph of 


(24) 11. Abdomen sessile, base of the 1st segment 

about half as w de as the metathorax. 
(17) 12. Legs dull red. with some black s're.sks. 
(14) 18, Antenne Q 25-joiuted(g unknown) ... 6. similis, Nees. 
(13) 14. Antennz Q with more than 25 joints. 
(16) 15. Mesoth: racic furrows inchvate, but effaced 

on the disk .. : 7. mandibulutor, Nees. 
(15) 16. Mesothoracic furrows complete, very finely 

traced, punctulate, couverging to a 

linear fovea in tront of the scute lum... 8. atra, Hal. 
(12) 17. Legs testaceous or red, without black 

streaks. 
(21) 18. Abdowen entirely black. 
(20) 19. Antenne & 31-jointed ; terebraas long as 


$ of the abdomen ( ¢ unknown)... 9. truncator, Nees. 
(19) 20. Antennze of 34-85-jointed ; terebra as 
long as 3 of the a domen és ... 10. tipule, Scop. 


(18) 21. Abdomen ‘partly red or testaceous. 
(23) 22. Abdomen, after the lst segment, testa- 
ceous; antenne 38-jointed (fg un- 
known) 11. Sophia, Hal. 
(22) 23. Abdomen black, with the base of the 2nd 
segment reddish, or with all the seg- 
meuts, except the Ist, red banded with 
black.. . 10. tipule, varr. 1, 2. 
(11) 24. Abd« men subpetiola'¢ e, Ist ‘segment nar- 
row, not widened poste: iorly. 
(26) 25. Abdomen entirely black; Ist segment 
twice as long as its ap cal breadth ... 12. incongrua, Nees. 
(25) 26. Abdomen red, except the Ist segment, 
which is three times as lony as its apical 
breadth au es cn Sac ... 18. lucicola, Hal. 


1. Alysia manducator, Panz. (Pl. XI., fig. 4, ?.) 


Ichneumon manducator, Panz., F. G., lxxii., 4. 

Cryptus manducator, Fab., Piez., 87. 

Bassus manducator, Panz., Krit. Rev., ii., 75; Nees, 
Mag. Ges. Berl., 1814, p. 202.. 

A. manducator, Latr., Gen. Crust. et Ins., iv., 15; 
Nees, Mon.,i., 289; Hal., Ent. Mag., v., 220, ¢ 9, pl. xvii., 
ae | (forewing), fig. 28 (hindwing) ; Cur., Farm ee 

44, 

A. stercoraria, Latr., H. N., xiii, 177. 

A. apicalis and similis, Cur., B. E., 141. 

A. truncator, Nees, Mon., i., 240, No. 2 (not i., 248, 
No. 7, also named truncatur). 


Spirac'es of the metathorax conspicuous, margined. The largest 
species, of r bust appearance. @ black, shining ; head very large; 
face punctato-rugulose ; mandibles large, often reddish. whol’y or 
in part; palpifuscous. Antenne shorter than in the other species, 
not much exceeding the length of the head and thorax, black, often 


British Braconide. 509 


reddish at the base beneath, 22-35-jointed, flagellum stout, bristly, 
composed of short cyathiform articulations, which are striolate. 
Mesothoracic sutures inchoate, broad, punctate ; middle lobe of 
the mesonotum elevated anteriorly ; an oblong fovea in front of 
the scutellum ; mesopleure rugose, with a broad furrow; meta- 
thorax short, almost truncate behind, covered with confused 
rugosities, carinate in the middle. Wings hyaline, often with an 
indeterminate brownish stain in the disk; squamule rufous ; 
stigma and nervures blackish brown ; stigma stout, triangular, 
emitting the radial nervure a little beyond the middle ; Ist inter- 
cubital nervure straight, a little longer than the 2nd abscissa ; 
pobrachial areolet of the hindwings half aslong as the prebrachial. 
Legs stout, rufous; tarsi dusky, hairy. Abdomen oboval, de- 
pressed, very shining ; Ist segment twice as broad at the extremity 
as at the base, rugulose, obtusely carinate in the middle, excavated 
at the base ; tubercles prominent, obtuse. Terebra very slightly 
exserted. dg similar; antenne longer than the body, subsetiform, 
33-50-jointed. Length, 13-31; wings, 34-7 lin. 

Var. @. Second abdominal segment rufous, black at the hind 
margin; legs paler red; wings hyaline ; terebra longer. Length, 
21; wings, 5 lin. Taken once only, in the London district. 
Haliday. 

This species is common throughout Europe, and pro- 
bably better known than most of the tribe; it was chosen 
by Latreille as the type of his genus Alysia, and is very 
indifferently figured by Panzer. The females may be 
observed, sometimes in considerable numbers, on carrion, 
excrements, and vegetable refuse, in search of dipter- 
ous maggots, in which to deposit their eggs. The sexual 

instinct attracts thither also the males; but both sexes 
likewise frequent flowers for the purpose of feeding. 
They scent the aroma of carrion at a surprising distance, 
as Lonce had occasion to observe in watching the re- 
mains of a dead rook, upon which they descended in 
constant succession, apparently from the sky, like vul- 
tures. ‘The males generally alighted on blades of grass 
close to the attractive object, as if to wait for their 
partners, without interrupting them in their unsavoury 
occupation. ‘l'hey have been reared from various mag- 
gots, as Lurilia cesar, L., Cyrtoneua stabulans, Fall., 
Hydrotza dentipes, Fab., and on one occasion recorded 
in the Ent. Monthly Mag., from the ferocious larve of 
the coleopterous Creophilus mavwillosus, L., living habitu- 


510 Rev. T, A. Marshall’s monograph of 


ally in carrion. Bouché has described the larva of this 
Alysia as: Oblong, fleshy, white, soft, glabrous, and 
semitransparent ; head somewhat rounded; oral parts 
indistinct ; dorsal segments elevated, the anal segment 
narrowed. Cocoon covered with a yellowish tissue of 
great tenuity. Length, 2 lines. 


2. Alysva soror, Marshall. 


A. soror, Marsh., Hymén. d’Europe et d’Algérie, 
Braconides, vol. ii., 377. 


Spiracles of the metathorax inconspicuous, punctiform, not 
margined. Antenne ? 40-jointed ; stigma triangular. (¢@ un- 
known.) Very like the preceding, but differing in the following 
particulars, which have obliged me to keep it separate. Form 
more elongate. Antennz subsetaceous, almost as long as the body, 
and with five additional joints, Wings hyaline, longer in propor- 
tion; 2nd cubital areolet differently shaped, produced in an acute 
angle towards the base of the wing ; 1stintercubital nervure twice 
as long as the 2nd ; in manducator the areolet is nearly square, 
the internal angle is hardly produced, and the 1st intercubital 
nervure is very little longer than the 2nd. First abdominal seg- 
ment longer, more narrowed at the base, not elevated in the 
middle, and without visible tubercles. Other differences are not 
apparent, but the structure of the antenne and wings forbids me 
to consider this a variety of manducator. Length, 2}; wings, 
53 Jin. 


I captured the only specimen in the London district. 


3. Alysia rufidens, Nees. 


A. rufidens, Nees, Mon., 1., 241; Hal., Ent. Mag., v., 
221, 3 9, pl. xvii, fig. 2 (wing). 

Antenne ¢ 9 22-24-jointed. Stigma elliptic. 9 black, pubes- 
cent; face finely granulated, dull; mandibles reddish ; palpi 
fuscous. Antenne black, filiform, shorter than the body. Meso- 
thoracic sutures three, the two lateral shortened ; metathorax very 
finely punctato-rugulose. Wings hyaline; nervures fuscous; 
squamula and stigma piceous, the latter oblong, elliptic, emitting 
the radial nervure from its exterior third ; radial areolet lanceo- 
late, not reaching the extremity of the wing ; 3rd abscissa slightly 
curved towards the end ; Ist intercubital nervure curved, almost 
twice as long as the 2nd abscissa ; Ist abscissa rather long, directed 


British Braconide. Sirk 


obliquely. Legs blackish, tips of the trochanters and of the femora, 
fore tibie entirely, and base of the four posterior, brownish. 
Abdomen shining ; Ist segment finely striated, subcarinate at the 
base. Terebra as long as half the abdomen. ¢ Antenne almost 
as long as the body ; wings narrower ; stigma large, elliptic, black, 
obliterating the 1st abscissa ; Ist cubital areolet very narrow ; legs 
paler than those of the 2. Length, 11-1? ; wings, 2-3 lin. 


This is probably a parasite of Hnsina sonchi, L., the 
larva of which inhabits the buds of Aster tripoliwm. 
Haliday captured it frequently in Ireland, on the sea- 
coast, and I have met with both sexes in England; it is 
also recorded as found in Germany and Holland. 


4, Alysia loripes, Hal. 
A. loripes, Hal., Ent. Mag., v., 222, d¢. 


Hind femora sinuated at the base, incrassated, compressed. 9 
Deep black, shining ; mandibles piceous. Mesothoracic sutures 
finely traced, punctulate, converging to a fovea before the scutel- 
lum. Wings hyaline ; squamula and stigma piceous brown, the 
latter narrow, emitting the radial nervure near its extremity ; 1st 
intercubital nervure somewhat longer than the 2nd abscissa. Legs 
piceous brown; tibiz and lower edge of femora dull testaceous. 
Terebra as long as half the abdomen. ¢ unknown. Length, 2 ; 
wings, 4 lin. 

The only specimen was taken by Walker near 
Windsor; the antennz were mutilated. 


5. Alysia fuscipennis, Hal. 


A. mandibulator, var. 8, Nees, Mon., i, 243, 6°. 
A. fuscipennis, Hal., Ent. Mag., v., 224, 3°. 


Wings infumated. @ Black, shining ; mandibles brown ; palpi 
blackish ; face hardly punctulate. Antennz longer than the body 
28-37-jointed. Metathorax rugose. Stigma and nervures brown 
or blackish, the former sometimes rather pale, emitting the radial 
nervure from its outer half ; radial areolet cultriform, not quite 
reaching the extremity of the wing ; 1st abscissa shorter than the 
thickness of the stigma ; 2nd at least as long as the 1st intercubital 
nervure ; 2nd cubital areolet contracted towards the extremity. 
Legs blackish brown, apex of the trochanters and base of the tibiz 
paler. First abdominal segment twice as broad at the extremity 


512 Rev. T. A. Marshall’s monograph of 


as at the base, sessile, striolate, sometimes smooth at the extremity, 
or (in small specimeus) almost wholly smooth; the following 
segments smooth and shining. Terebra as long as the abdomen; 
valves stout, hairy. ¢ Antenne longer, 37—-4(-juinted ; 1st abdo- 
minal segment scarcely widened posteriorly, faintly striolate, or 
almost smooth. Length, 12-2; wings, 31-44 lin. 


Var. Abdomen piceous with the Ist segment rufescent and 
nearly smooth; legs paler piceous. 


Taken in England by Walker; in Ireland, rarely, by 
Haliday. Billups has captured it at Oxshott, and I have 
two which I captured near Abergavenny, and a third 
which I found at Nantua last summer, in the Jura. I 
have seen others from the Continent, taken in Italy and 
Germany. 


6. Alysia similis, Nees. 


(?) Bassus similis, Nees, Mag., Ges. Berl., 1814, 
p. 203. 

(?) A. similis, Nees, Mon.,i., 240,¢2; Hal., Ent. 
Mag., v., 223, ?. 


Antennz 9? 25-jointed. Black, shining; mandibles pitchy red. 
Antenne stout, shorter than the body. Metathorax punctato- 
rugulose. Wings hyaline; squamula blackish-brown ; stigma 
blackish, stouter than in the allied species, confounded at the ex- 
tremity with the metacarp, emitting the radial nervure from near 
its middle; radial areolet not quite reaching the extremity of the 
wing ; recurrent nervure interstitial. Legs dull red; fore femora 
streaked with black at the base, the 4 posterior femora streaked 
throughout their whole length; tarsi, coxe, and trochanters 
blackish. First abdominal segment regularly striated. Terebra, 
according to Nees, half as long, according to Haliday a quarter as 
long, as the abdomen. ¢ Similar, according to Nees, but not 
described ; unknown to Haliday. Length, 14 ; wings, 3 lin. 


This species is unknown to me. MHaliday expresses a 
doubt whether his insect was the same species as the 
similis of Nees, on account of the difference in size, and 
in the length of the terebra. In other respects the 
descriptions of these authors coincide. 

Taken once in the London district, with A. man- 
ducator. . 


British Braconide. 5138 


7. Alysia mandibulator, Nees. 


Bassus mandibulator, Nees, Mag., Ges. Berl., 1814, 
p. 204. . 
A. mandibulator, Nees, Mon., i., 242 (not the var.) ; 
Hal., Ent. Mag., v., 224, 6 ¢. 

Mesothoracic sutures inchoate, but effaced on the disk. 9 Deep 
lack, shining ; mandibles pitchy red. Head and thorax vaguely 
punctulate, thinly beset with whitish hairs. Antenne a little 
longer than the body, filiform, rather stout, 32-34-jointed. The 
effaced sutures of the mesonotum are indicated by rows of very 
fine punctures ; before the scutellum is a median line of similar 
punctures ; metathorax rugose. Wings dingy byaline ; squamula 
reddish ; nervures brown ; stigma reddish, subelliptic, emitting the 
radial nervure from its exterior half; radial areolet (measured 
along the costa) hardly longer than the stigma, lanceolate, not 
reaching the extremity of the wing ; 2nd abscissa a little shorter 
than the Ist intercubital nervure; @rd abscissa slightly curved at 
the extremity ; recurrent nervure interstitial ; pobrachial areolet 
of the hind wing half as long as the prebrachial. Legs as in the 
preceding species ; hind tibiz sometimes dark at the extremity. 
Abdomen elongate-oval, deep black, brilliant; lst segment some- 
what narrowed towards the base, regularly striolate. Terebra 
about 2 of the length of the abdomen. ¢ unknown. 


Haliday’s only specimen had a dark stigma, but in my 
four specimens the stigma is of a reddish tinge. Walker 
first detected this species in England, and I have siuce 
found it in some numbers in an osier-bed at Nunton, 
Wilts, but females only: more I believe are in Dr. 
Capron’s collection. 


8. Alysia atra, Hal. 


A. atra, Hal., Ent. Mag., v., 223, d¢, pl. xvu, £3 
(wing). 

Mesothoracie sutures finely traced, complete, punctulate, con- 
verging to a linear fovea before the scutellum. Very like the 
preceding, but more hairy. ¢ deep black, shining, with the ex- 
tremity of the abdomen pale piceous ; mandibles reddish. Antennz 
a little shorter than the body, 29-81-jointed. Wings hyaline ; 
squamula pale brown; stigma dull testaceous. Legs reddish ; 
coxe, upper side of femora, and tarsi, blackish. Terebra as long 
as 2 of the abdomen. ¢ antenne longer, 35-jointed ; stigma black ; 


514 Rev. T. A. Marshall’s monograph of 


1st cubital areolet narrower ; 2nd smaller ; radial hardly attaining 
the tip of the wing; recurrent nervure somewhat rejected. This 
d much resembles rujidens (No. 3), but differs in the length of 
the antenne, and the neuration. Length 14; wings, 34 lin. 


Very rare in North Ireland, according to Haliday. 
My collection contains two ¢?s taken respectively in 
Yorkshire (Lastingham) and Wiltshire (Nunton), and 
one 6 captured at Plumstead by Mr. Billups. It was 
also found by Mr. Walker in Finmark. 


9. Alysia truncator, Nees. 


Bassus truncator, Nees, Mag. Ges. Berl., 1814, p. 204. 

A. truncator, Nees, Mon.,i., 243, No. 7 (not i., 240, 
No. 2, also named truncator, which is a synonym 
of manducator) ; Hal., Ent. Mag., v., 222, ¢ ?. 


Antenne ? 31-jointed ; terebra as long as 1 of the abdomen 
(3S unknown). Deep black, shining ; mandibles ferruginous, black 
at the points, on the sides, obtusely denticulated ; palpi ferru- 
ginous ; face shining, nearly smooth. Antenne rather stout, almost 
as long as the body ; 1st joint ferruginous, fuscous underneath ; 
2nd ferruginous. Metathorax finely punctulate. Wings hyaline ; 
squamula ferruginous ; stigma obscurely reddish ; nervures brown ; 
radial areolet acuminate, hardly reaching the tip of the wing ; 1st 
intercubital nervure longer than the 2nd abscissa ; interior angle 
of the 2nd cubital areolet considerably produced. Legs rufo- 
testaceous, tarsi darkened at the tips. Abdomen compressed at 
the extremity ; Ist segment very slightly narrowed towards the 
base, depressed, rugulose. The g¢ mentioned by Nees is doubt- 
ful :—Similar to the ? ; antenne longer than the body ; hind 
femora and hind tibie black at the extremity ; their tarsi also 
dark. Length barely 2 lin, 


I have not seen this species, which was found in 
England by Curtis. According to Goureau, it is a para- 
site of Agromyza macquarti, Rob. Desv., the larva of 
which mines the leaves of Verbascum thapsus; and of 
Anthomyia platyura, Meig., the larva of which inhabits 
the bulbs of Cepa ascalonica, the eschalot. 


10. Alysia tipulz, Scop. (PI. XL. fig. 1) 2.) 


Ichneumon tipulx, Scop., Ent. Car., 288 (1763). 
Bassus abdominator, Nees, Mag. Ges, Berl., 1814, 
p. 205. 


British Braconide. 51S 


A, abdominator, Nees, Mon., i., 245, ¢ 2. 
A. tipule, Hal., Ent. Mag., v., 224, d 2, pl. xvi, f. 4 
(wing). 

Antenne @ 34~-35-jointed; terebra as long as 2 of the ab- 
domen. Black, shining ; abdomen often partiy red, in varieties ; 
mandibles testaceous ; palpi rufous, Antenne rather stout, as 
long as the body, black, with the two first joints testaceous ; cheeks 
often more or less rufescent. Prothorax piceous ; mesothoracic 
sutures inchoate, effaced posteriorly ; before the scutellum is an 
oval fovea ; metathorax rugose. Wings hyaline ; squamula rufous ; 
nervures and stigma testaceous, the latter emitting the radial ner- 
vure from its outer half; radial areolet cultriform, not quite 
reaching the tip of the wing; 2nd discoidal areolet emitting 
the anal nervure from the middle of its angular extremity ; Ist 
intercubital nervure somewhat shorter than the 2nd abscissa ; re- 
current nervure interstitial, Legs testaceous, with dark tarsi ; an 
indistinct dark spot on the upper side of the hind coxe. Abdomen 
oblong-oval ; 1st segment finely rugulose, hardly twice as wide 
at the extremity as at the base; ist suture deeply impressed ; 
2nd and following segments very smooth, generally more or 
less brownish-black, with the 3 or 4 apical segments paler brown. 
6 antennz longer than the body, filiform, 43-jointed ; stigma fus- 
cous ; abdomen entirely black. Length 13-2 ; wings, 32-43 lin. 

Var. 1. ¢ 2. Second abdominal segment rufescent at the base. 

Var. 2. 9. Abdomen, after the Ist segment, rufous, each 
segment banded with black ; apical segments pale. This variety, 
at least in England, is more common than the typical form, and 
most of my specimens belong to it. 


Common throughout Europe, frequenting fungi in 
woods, and parasitic probably in the larve of Myceto- 
phila, and its allies. 

11. Alysia Sophia, Hal. 

A. Sophia, Hal., Hunt. Mag., v., 225, ¢. 

Q. Antenne 38-jointed, flavo-rufous with the extremity fus- 
cous (¢ unknown). Black, shining, abdomen testaceous, except 
the first segment. Clypeus brown ; mandibles rufous ; palpi the 
same, but paler. Antenne a little longer than the body. Wings 
hyaline; squamula and stigma flavo-rufous. Legs flavo-rufous, 
with paler coxe. Second abdominal segment testaceous, but 
darker than the rest. Terebra as long as the 5 or 4 apical seg- 
ments. Length, 13 ; wings, 4 lin. 


Taken, once only, by Haliday, in Ireland. 


516 Rey. T. A. Marshall’s monograph of 


12. Alysia incongrua, Nees. 


A. incongrua, Nees, Mon., i1., 244 (?); Hal., Ent. 
Mar. ¥., 220 ee 

A. Iucit, Hal., Ent. Mag., v., 226, 6. 

Abdomen black; 1st segment sublinear, twice as long as its 
apical breadth. @ Black, shining ; mandibles rufous, Antennz 
with more than 29 joints (broken), as long as the body; 2 first 
joints rufous beneath, or wholly rufous. Metathorax rugose, 
reticulated Wings hyaline; squamula flavo-rufous ; stigma stout, 
blackish. Legs pale testaceous; apex of hind tibix, and their 
tarsi, fuscous, Abdomen subpetiolate or subsessile, black, brownish 
towards the extremity ; 1st segment of equal breadth throughout, 
longitudinally rugulose. ‘Terebra as long as 2 of the abdomen. 
$ Antennez slender, longer than the body, 40-joiuted ; second 
cubital areolet somewhat Jonger than in other species ; legs flaves- 
cent, apex of hind tibiz, and their tarsi, fuscous ; 2nd abdominal 
segment obscurely rufescent at the base. Length, 9 2}; wings, 5; 
6 length, 14 ; wings, 44 lin. 


The doubt attaching to the identity of the incongrua 
of Nees arises from that author’s omission of the form of 
the first abdominal segment. I possess only the ¢, 
which is undoubtedly the Lucia of Haliday, as he him- 
self conjectured. It was taken near Plymouth by Mr. 
Bignell. The d described in the Ent. Mag. was from 
Edinburgh. 


13. Alysia lucicola, Hal. 

A. lucicola, Hal., Ent. Mag., v., 226, 6 9. 

Abdomen, after the Ist segment, rufo-testaceous ; Ist segment 
linear, three times as long as its apical breadth. @ Black, shining ; 
mandibles rufous ; palpi ferruginous. Antennw a little longer than 
the body, 32-jointed ; the two first joints testaceous. Mesothoracic 
sutures complete; a circular fovea before the scutellum ; meta- 
thorax rugose. Wings hyaline with a faint dusky tinge ; squamula 
pale ferruginous; nervures and stigma fuscous; radial areolet 
cultriform, reaching the tip of the wing ; Ist intercubital nervure 
very obliquely placed, a little longer than the 2nd abscissa ; recur- 
rent nervure evected ; 2nd discoidal arcolet emitting the anal 
nervure from the middle of its angular extremity ; 2nd cubital 
areolet a little narrowed outwardly. Legs testaceous; coxe paler ; 
tips of the hind tibiz, and their tarsi, infuseated. Abdomen pyri- 
form, somewhat compressed ; Ist segment black, striolate, making 


British Braconidx. Eyilng 


one-third of the total length of the abdomen; the following 
segments smooth, testaceous. Terebra a little longer than half the 
abdomen. ¢ Antenne one-half longer than the body, slender, 
setaceous, 38-jointed ; abdomen somewhat clavate, depressed, of a 
more dingy testaceous than that of the ?, the apical segments 
infuscated. Length, 2; wings, 43 lin. 


The 2 is not unlike A. tipule (No. 10), but imme- 
diately distinguishable by the form of the Ist abdominal 
segment. Taken first by Walker in England; in Ireland, 
rarely, by Haliday, amongst fungi. I have three speci- 
mens, 2 6, 1 ?, which I captured in Devonshire. 


viii. Tanycarpa, Forster. 
Forst., Verh. Pr. Rheinl., 1862, p. 265. 

Fourth joint of the antenne shorter than the 3rd. Metathorax 
not carinated ; furrow of the mesopleure crenate. Stigma elongate, 
emitting the radial nervure before the middle; radial areolet 
reaching the tip of the wing ; recurrent nervure interstitial ; 1st 
intercubital nervure longer than the 2nd abscissa; Ist cubital 
areolet separated from the Ist discoidal ; 2nd complete, separated 
from the first ; anal nervure springing from the middle of the ex- 
tremity of the 2nd discoidal areolet. First abdominal segment 
somewhat elongate; 2nd segment smooth, shining. ‘Terebra 
considerably exserted. 


TABLE OF SPECIES. 


. Abdomen red and black, or red and f fuscous. 

Abdomen red, with thes Ist segment black, or 

bla-k with segments 2 2-3 red ... 2. rufinotata, Hal. 
. Abdomen blackish or fuscous, with ‘the Ist 

segment and base of the 2nd reddish, or 

impure testaceous alee noe ; . 3. ancilla, Hal. 


(4) 1. Abdomen entirely black. 

(3) 2. Antennze 22-25-jointed .. 305 306 . 3. ancilla, var. 

(2 . Antennz 33-37-jointed .. .. 1. gracivicornis, Nees. 
(a 

(6 


_— ———~r 
o enca 


1. Tanycarpa gracilicornis, Nees. 
Bassus qracilicornis, Nees, Mag., Ges. Berl., 1814, 
p. 206. 
Alysia eee Nees, Mon., i, 247; Hal., Ent. 
Mag., v., 228, ¢ 9, pl. xvii, f. 11 (wing). 

2. Black, steht face minutely punctulate ; clypeus and 
mandibles ferruginous, palpi paler. Antenne slender, nearly one- 
half longer than the body, black with the two first joints rufous, 
33-37-jointed. Metathorax rugulose, sometimes smooth at the 


518 Rey. I’. A. Marshall’s monograph of 


base. Wings hyaline ; squamula pale ferruginous; nervures 
slender ; stigma dull testaceous, very long, linear, emitting the 
radial nervure at the end of the 1st third ; radial areolet cultriform; 
2nd cubital areolet narrowed outwardly. Legs flavo-rufous with 
the coxee paler ; tips of 4 anterior tarsi, hind tarsi entirely, and tips 
of hind tibie, infuscated. Abdomen subpetiolate ; Ist segment 
elongate, almost linear, hardly widened behind, with two dorsal 
carine vanishing posteriorly, very finely rugulose. Terebra as 
long as 2 of the abdomen. @ Similar; antennz about twice as 
long as the body, 39-42-jointed ; 2nd abdominal segment sometimes 
piceous. Length, 12-2 ; wings, 32-43 lin. 


Parasite of Agromyza cicerine, Rondani. It has oc- 
curred in Germany, Italy, and Ireland. 


2. Tanycarpa rufinotata, Hal. 


Alysia rufinotata, Hal., Ent. Mag., v., 227, 39, pl. 
xvii, f. 13 (wing). 


Abdomen rufous with the Ist segment black, or black with 
segments 2-3 rufous. @ Black; head very large shining, the face 
somewhat duller ; clypeus, mandibles, and palpi rufous. Antenne 
hardly longer than the body, 35-jointed, black with the two basal 
joints rufous. Mesothoracic sutures inchoate ; a fovea in front of 
the scutellum ; metathorax rugulose. Wings hyaline; squamula 
rufous ; nervures and stigma fuscous, the latter linear-lanceolate, 
emitting the radial nervure a little before the middle ; 2nd cubital 
areolet narrowed externally ; lower exterior angle of the 2nd 
discoidal areolet produced. Legs rufous; hind tarsi and tips of 
hind tibiz infuscated. Abdomen clear red, or more or less 
tinged with brown; Ist segment obconic, attenuated towards the 
base, rugulose. Terebra as long as the abdomen. g¢ Antenne 
longer, 39-jointed. Length, 13-2 ; wings, 34-4 lin. 


England and Ireland; somewhat rare. Found in 
autumn, amongst fungi. 


3. Tanycarpa ancilla, Hal. 


Alysia ancilla, Hal., Ent. Mag., v., 227, d ¢. 

Abdomen pitchy black, with the first segment and base of the 
2nd rufescent, or dull testaceous. @ Brownish-black, shining ; 
clypeus and mandibles rufous; palpi paler. Antenne hardly 
longer than the body, 21-25-jointed, blackish with the two first 


joints and base of the 3rd flavo-rufous. Discal fovea of the 


British Braconide. 519 


mesonotum very small ; furrow of the mesopleure finely traced, 
crenulate. Wings hyaline ; squamula and stigma pale ochreous ; 
nervures brownish, arranged as in the preceding ; hind wings 
narrower, without a transverse nervure. Legs pale :flavo-rufous, 
hind tarsi and tips of hind tibiz scarcely darker. First abdominal 
segment elongate, finely rugulose. Terebra as long as 2 of the ab- 
domen. ¢ Antenne somewhat longer than the body, with the same 
number of joints as those of the @ ; stigma fuscous, Length, 1; 
wings, 24 lin. 
Var. Abdomen entirely pitchy black, concolorous. 


England and Ireland ; the males, according to Haliday, 
more common than the females. 


ix. PenvarLeura, Forster. 
Férst., Verh. Pr. Rheinl., 1862, p. 264. 

Fourth joint of the antenne shorter than the 3rd. Maxillary 
palpi 6-, labial 3- or 4-jointed. Metathorax not carinated ; furrow 
of the mesopleurze smooth or obsolete. Stigma small, sometimes 
indeterminate at the extremity where it is confounded with the 
metacarp, and sometimes subobsolete; radial areolet ample, cultri- 
form, attaining the tip of the wing; 1st intercubital nervure longer 
than the 2nd abscissa ; Ist cubital areolet separated from the Ist 
discoidal ; 2nd cubital areolet complete, pentagonal; recurrent 
nervure plainly evected; 2nd discoidal areolet imperfectly closed ; 
anal nervure subinterstitial. 'Terebra considerably exserted. 


The small black species of this genus may be recog- 
nized at once by their wings; the 2nd cubital areolet is 
produced angularly to meet the recurrent nervure, thus 
forming a 5th side to the areolet; hence the name of the 
genus. ‘This structure, however, must be considered 
ideal, for the nervures, as usual, lose their colour on 
approaching the anastomosis. ‘The antennz exhibit a 
diminished number of joints, and the labial palpi are 
sometimes triarticulate. 

The species are not easy to tabulate, but perhaps the 
following attempt may conduce to their determination :— 


(2) 1. Stigma much attenuated, subobsolete, con- 

founded with the metacarp. Antenne 

9 17, g 22-jointed... aR oe .. 1, angustula, Hal. 
(1) 2. Stigma of ordinary form, or only moderately 

narrowed. 
(4) 3. Stigma of ordinary form. Antenne @ 24-, 

& 27-jointed ... “58 a: - ... li. fuliginosa, Hal. 
(8) 4, Stigma narrowed, but distinct, confounded 

with the metacarp only at its extremity. 

Antenne 92 19-, g 25-jointed ... .. li. pumilio, Nees. 


bo 


520 Rey. ‘I’. A. Marshall’s monograph of 


I. Pentaplewra angustula, Hal. 


Alysia angustula, Hal., Ent. Mag., v., 229. 9; pl. 
xvii, f. 8 (wing). 


Stigma subobsolete, confounded -with the costa and metacarp. 
Antenne @ 17-, ¢ 22-jointed. @ Black or pitchy ; mandibles 
dull testaceous ; labial palpi 3-joimted. Antenne rather shorter 
than the body. Mesopleurze smooth, with no trace of a furrow ; 
metathorax nearly smooth. Wings hyaline ; nervures fuscescent ; 
radial areolet very long ; 1st intercubital nervure hardly longer 
than the 2nd abscissa ; 1st abscissa longer than the greatest breadth 
of the stigma ; hindwings narrow; pobrachial areolet short ; only 
one transverse nervure. Legs dull testaceous. Abdomen sessile, 
somewhat compressed ; Ist segment smooth, obtusely carinated in 
the middle. Terebra as long as the abdomen, falcate, ascending: 
¢ Antenne twiceas long as the body, slender, black with the two 
basal joints testaceous ; abdomen oval, depressed. Length, 3; 
wings, 13-2 lin. 


Found in England and Ireland, but seldom; I have 
one specimen, captured in Northants. 


2. Pentapleura fuliginosa, Hal. 

Alysia fuliginosa, Hal., Ent. Mag., v., 228, 5; 519,9. 

Stigma oblong, well defined at both ends. Antenne 9? 24-, 
¢ 27-jointed. @ Black, shining; mandibles dull testaceous ; 
labial palpi 4-jointed. Antennz as long as the body. Mesopleure 
with no trace of a furrow ; metathorax short, rugulose. Wings 
subhyaline ; nervures and stigma fuscous, the former disposed as 
in angustula ; 1st abscissa only half as long as the thickness of the 
stigma. Legs dull testaceous, Abdomen compressed near the apex ; 
1st segment striolate. Terebra as long as 2 of the abdomen. ¢ An- 
tenn one-half longer than the body. Length, 1 ; wings, 2+ lin. 


Less common than the following species ; Haliday took 
his specimen in Kent; I have a d from Abergavenny, 
and a ? from Bishop’s Teignton. 


3. Pentaplewra pumilio, Nees. 
Bassus pumilio, Nees, Mag., Ges. Berl., 1814, 
p. 205. 
Alysia pumilio, Nees, Mon.,i., 244, 2; Hal., Ent. 
Mag., v., 228, d ¢ ; pl. xvii, f. 7 (wing). 


British Braconide. PA 


Stigma attenuated, confounded with the metacarp at its outer 
end. Antenne 9? 19-, g 25-jointed. @ Black, shining ; man- 
dibles reddish ; labial palpi 3-jointed. Antenne shorter than the 
body, black ; 3rd joint elongate. Mesothoracic sutures inchoate, 
almost complete in large specimens ; a dorsal fovea before the 
scutellum ; furrow of the mesopleure barely indicated, un- 
punctured ; metathorax indistinctly rugulose. Wings hyaline ; 
squamula fuscous ; nervures and stigma pale brown or testaceous ; 
Ist abscissa a little shorter than the greatest thickness of the 
stigma; pobrachial areolet of the hindwings somewhat less than 
half as long as the prebrachial. Legs dull testaceous ; cox, and 
tips of the femora and tibis, infuscated. Abdomen oblong-oval ; 
Ist segment obconic, finely striolate. Terebra as long as 
% Of the abdomen. ¢ Antenne one-half longer than the body; 
legs longer, more slender, and darker ; wings slightly infumated. 
Length, 3 ae ; wings, 22 lin. 


Common in England and Ireland ; Nees was acquainted 
with it in Germany, and Walker obtained it in his tour 
through Finmark. 


x. Iprasta, Forster. 
Forst., Verh. Pr. Rheinl., 1862, p. 265. 


Fourth joint of the antenne longer than the 3rd. Maxillary 
palpi 6-, labial 4-jointed. Mesothoracie sutures deeply impressed, 
complete, crenulate or merely inchoate ; metathorax carinated at 
the base. Stigma large, determinate ; Ist intercubital nervure 
longer than the 2nd abscissa; 1st cubital areolet separated from 
the 1st discoidal ; 2nd quadrangular ; radial areolet lanceolate, not 
reaching the tip of the wing; 3rd abscissa straight ; recurrent 
nervure not evected ; 2nd discoidal areolet completely closed ; anal 
nervure not interstitial. Terebra considerably exserted. 


(2) 1. Black with a tint of bronze, submetallic ; 
wings clouded with obscure patches; anal 


nervure not interstitial . ... 1. maritima, Hal, 
(1) 2. Black without a metallic tinge; wings infumated, 

without darker spots; anal nervure ens 

interstitial ee ae ye E ... li, Nephele, Hal. 


1. Idiasta maritima, Hal. (PI. XII, fig. 2, ¢.) 
Alysia maritima, Hal., Ent. Mag., v., 230, ¢?; pl. 
xvi, f. 14 (wing). 
? Black, with a tint of bronze, submetallic, shining, beset with 
whitish pubescence ; face and margin of the vertex very finely 
TRANS. ENT. SOC. LOND. 1894.—rarriv. (pEC.) 21 


522 Rev. T. A. Marshall’s monograph of 


rugulose ; mandibles reddish ; palpi dusky. Antenne filiform, as 
long as the body, 24-25-jointed, black with the base of the 
flagellum dull testaceous. Mesothoracic sutures crenate-punctured, 
separated posteriorly by a rugose space traversed by an oblong 
fovea ; metathorax rugose with two smooth basal are, separated 
by acarina. Wings yellowish, with all the transverse nervures, 
and a blotch near the tip of the radial areolet, clouded with 
fuscous ; squamula rufous ; nervures and stigma fuscous, the latter 
large, semi-oval, cutting off the greater part of the Ist abscissa 
recurrent nervure conspicuously rejected ; anal nervure springing 
below the middle of the extremity of the 2nd discoidal areolet ; 
pobrachial areolet of the hindwings almost as long as half the 
prebrachial. Legs dull testaceous; cox blackish. Abdomen 
depressed, shining, rounded at the extremity ; Ist segment one- 
half longer than broad, narrowed towards the base, longitudinally 
striolate, elevated in the middle at the base. Terebra as long as 4% 
of the abdomen. ¢ Similar; antennz 29-jointed. Length, 13-2 ; 
wings, 3-4 lin. 

This species occurs sparingly amongst decaying sea- 
weeds on the coast: the localities named are Hebrides, 
Yorkshire, Hampshire. I have taken it on Lymington 
Salterns, and Mr. Billups found it at Dulwich. 


2. Idiasta Nephele, Hal. 
Alysia Nephele, Hal., Ent. Mag., v., 231, ¢. 


Black without metallic tinge ; wings infumated without darker 
blotches ; anal nervure almost interstitial. 9 Mandibles reddish ; 
eyes minutely pubescent. Antenne as long as the body, filiform, 
20-jointed, blackish, pitchy at the base beneath. Mesothoracic 
sutures inchoate; a small dorsal fovea before the scutellum; metatho- 
rax punctato-rugose. Wings narrow; squamula brownish ; stigma 
fuscous, not so stout as in the preceding; recurrent nervure not much 
rejected; pobrachial areolet of the hindwings shorter. Terebra about 
as long as the abdomen. ¢ unknown. Length, 1 ; wings, 23 lin. 


One example taken by Haliday in the Hebrides, and 
another in Finmark by Walker. 


xi. APHHRETA, Horster. 
Forst., Verh. pr. Rheinl., 1864, p. 264. 


Fourth joint of the antennz longer than the Srd. Maxillary 
palpi 6-, labial 4-jointed. First cubital areolet confounded with the 


British Braconide. 523 


1st discoidal (as in gen. iii., Syncrasés) ; 2nd cubital areolet separated 
from the Ist; 1st intercubital nervure shorter than the 2nd 
abscissa ; 2nd discoidal areolet obsolete ; anal nervure interstitial ; 
hindwings very narrow, pobrachial areolet and transverse nervure 
obsolete. Terebra considerably exserted. 


With this genus commences the second artificial 
section of the Alystids, comprising those which have an 
elongated 2nd cubital areolet ; in other words, the Ist 
intercubital nervure is shorter than the 2nd abscissa 
of the radial nervure. There is only one species, 
variable in size and appearance, but with characters that 
cannot be mistaken. 


1. Aphxreta cephalotes, Hal. 


Stephanus minutus, Nees, Mag. Ges. Berl., 1811, 
Boye. 

Alysia fuscipes, Nees, Mon., i., 254, 2 (not of Hal.). 

Bassus minutus, var. Nees, Mag. Ges. Berl., 1814, 
p- 212. 

Alysia minuta, var. y, Nees, Mon. 1., 252, ?. 

Alysia cephalotes, Hal., Ent. Mag., v., 231, d 2, pl. 

xvu, f. 18 (wing). 

Alysia confluens, Ratz., Ichn. d. Forst., 1., 55. 

Black or pitchy-black, shining ; head large ; mandibles and palpi 
rufous. ¢ Antenne a little longer than the body, blackish, with 
the two first joints rufous, 19-25-jointed. Prothorax often 
piceous ; mesothoracic sutures usually distinct, smooth, less visible 
in small specimens ; no dorsal fovea before the scutellum ; furrow 
of the mesopleure rugose ; metathorax rugulose, smoother in the 
middle and towards the base. Wings subhyaline; squamula 
brown ; nervures and stigma testaceous, more or less dusky, the 
latter much attenuated, confounded posteriorly with the metacarp ; 
radial areolet cultriform, elongate, reaching the tip of the wing ; 
2nd cubital areolet elongate, narrowed towards the end; 2nd 
abscissa almost twice as long as the 1st intercubital nervure ; anal 
nervure springing from the cubital; no 2nd discoidal areolet. Legs 
dull rufous. First abdominal segment obconic, finely rugulose, 
sometimes rufescent. Terebra as long as2of the abdomen. ¢@ 
similar ; antenne hardly twice as long as the body, 27-jointed. 
Length, 1-1; ; wings, 23-3 lin. 

Var. 1. 9. Form more slender, Antenne one-half longer than 
the body, blackish, broadly testaceous at the base, 25-jointed ; 
mesonotum with a fovea before the scutellum ; furrow of the 


524. Rev. 'T. A. Marshall’s monograph of 


mesopleurze very finely traced, crenulate; metathorax nearly 
smooth ; legs slender, elongate, flavo-rufous; wings hyaline. 
Length, 1; wings, 2 lin. This may be, as suggested by Haliday, 
a distinct species. I possess a single specimen. 

Var. 2. ¢. Slender; entirely black ; antenne slender (muti- 
lated) ; mesothoracic sutures effaced; no dorsal fovea; wings 
hyaline ; nervures and stigma pale testaceous, the latter almost 
obsolete ; legs dark brown with the knees and 2nd joint of the 
trochanters rufescent. Length, 2; wings nearly 2 lin. This 
variety, communicated by Dr. Capron, seems as doubtful as the 
preceding ; it may be the fuscipes of Nees, too briefly described, 
but which, without further information, I must regard as a 
synonym of cephalotes, differing only in the darker colour of the 
legs. 

Var. 3. @. Length, 14; wings, 3£ lin. Antenne longer than 
the body, 24-jointed ; mesothoracic sutures complete ; the dividing 
nervure between the Ist cubital and the 1st discoidal areolets 
inchoate. The large size of this specimen, taken in Cornwall, gives 
it a different aspect from the others, yet its essential characters are 
the same. 


This species is rather common: it has been noticed in 
England, Ireland, Germany, Holland, and Russia, fre- 
quenting sea-weed on the coasts, as well as inland 
situations. Ratzeburg believed his Alysia conjflwens to 
be a parasite of the lepidopteron Amphidasys betularius, 
L.; but it came most probably from some dipterous larva 
which had escaped his observation. 


xii. Pamnocarra, Forster. 
Forst., Verh. Pr. Rheinl., 1862, p. 267. 


Fourth joint of the antenne longer than the 3rd. Maxillary 
palpi 6-, labial 4-jointed. First cubital areolet complete ; 1st inter- 
cubital nervure shorter than the 2nd abscissa ; 2nd discoidal areolet 
complete; anal nervure subinterstitial, rarely interstitial ; pobra- 
chial areolet of the hindwings not half as long as the prebrachial. 
With few exceptions, the antenne are long and slender, the 4th 
and 5th joints both longer than the 3rd; the rugosities of the 
metathorax and Ist abdominal segment are less distinct than in 
Alysia ; the clypeus larger; face convex and nearly smooth; 
stigma oblong, emitting the radial nervure beyond the middle, 
often attenuated and indeterminate towards the tip of the wing. 
The species much resemble Alysia, but must be distinguished by 
the longer 2nd cubital areolet (see table of genera). 


British Braconide. Be 


I have included here three more of Forster’s genera, 
Homophyla, Asobara, and Idiolewis, each adapted to a 
single species. ‘lhe ‘European fauna contains 17 known 
species of Phenocarpa, of which 14 are British. The 
number would, no doubt, be greatly increased if foreign 
entomologists were to turn their attention to this tribe ; 
but, in our country, few species, if any, still remain 
undetected. 


TABLE OF SPECIES. 


(2) 1. Radial areolet not reaching the tip of the 
wing (Homophyla, Foérst.) . 1. pullata, Hal. 
(1) 2. Radial areolet reaching the tip of the wing. 
(4) 8. Pobrachialareolet produced beyond the prex- 
brachial; 2nd discoidal areolet small, oval 
instead of quadrate ; radial areolet of the 
hindwings almost bisected by an accessory 
transverse nervure (Idiolexis, Forst.) ... 2. punctigera, Hal. 
(3) 4. Pobrachial areolet not longer than the pre- 
brachial ; 2nd discoidal areolet of ordinary 
size and form, rarely obsolete or sub- 
obsolete ; radial areolet of the hindwings 
simple. 
. Second discoidal areolet obsolete ; stigma 
much attenuated, indeterminate eee 
Forst.) .. 3. tabida, Nees. 
. Second discoidal. areolet complete, or nearly 
so ; stigma more or less conspicuous. 

. Transverse nervures bordered with brown... 4 picinervis, Hal. 

. Transverse nervures not so bi rdered. 

. Metathorax transversely subcarinate near 

the extremity; head generally rufous, 
body black ; or both head and body testa- 
ceous; stigma semioval, rather stout ... 5. rujiceps, Nees. 
(9) 10. Metathorax not subcarinate near the ex- 
tremity ; head concolorous with the body, 
black or testaceous; stigma less stout, 
often much attenuated. 

(14) 11. Antenne 9 hardly as long as the body, 19- 
21-jointed; g unknown. 

(18) 12. Second cubital areolet rather short, the 2nd 
abscissa being very little longer than the 
lst intercubital nervure... 6. Maria, Hal. 

(12) 18. Second cubital areolet elongate, the 2nd 
abscissa twice as long as the 1st inter- 
cubital nervure 

(11) 14. Antenne # 2 much longer than the body, 
consisting of more than 21 joints. 

(16) 15. Body rufous or testaceous, with a spot on 
the vertex, the greater part of the thorax, 
and the 1st abdominal segment, fuscous... 8. Eugenia, Hal. 

(15) 16. Body almost entirely black, or pitchy black. 

(20) 17. Mesothoracic sutures complete, meeting just 
before the scutellum. 

(19) 18. Antennze 24-32-jointed ; terebra as long as % 
of the abdomen aa ee ... 9. conspurcator, Hal. 

(18) 19. Antennze 33- 41-jointed ; "terebra longer than 
the whole body hic 450 nite ... 10. pratelle, Curtis. 


— 
or [=r 
—— = 
a Or 


—_~— 
6 CONT 


“TI 


Galatea, Hal. 


526 Rev. T. A. Marshall’s monograph of 


(17) 29. Mesothoracic sutures inchoate, effaced pos- 
teriorly. 

(24) 21. Wings infumated. 

(23) 22. Terebra shorter than the abdomen; ¢ 


unknown oe .... 11. Nina, Hal. 
(22) a Terebra longer than the abdomen... ... 12. Hunice, Hal. 
(21) 24. Wings hyaline. 
(26) 25. Abdomen black or pitchy black oe . 18. flavipes, Hal. 
(25) 26. Abdomen, after the Ist seni medidiahe or 

testaceous oe are Bae . 14. livida, Hal. 


1. Phenocarpa pullata, Hal. (PI. XITI., fig. 5, wing.) 


Alysia pullata, Hal., Ent. Mag., v., 232, 3 9, pl. xvii, 
f. 15 (wing). 

Radial areolet not reaching the tip of the wing. @ Deep black, 
shining ; mandibles brownish; palpi dusky. Antenne as long as 
the body, 34-jointed. Mesothoracic sutures complete, crenulate, 
meeting in an acute angle near the scutellum ; metathorax rugu- 
lose. Wings subhyaline : squamula rufous ; nervures and stigma 
fuscous, the latter oblong, emitting the radial nervure beyond the 
middle ; radial areolet short, lanceolate ; radial nervure curved ; 
2nd abscissa very little longer than the 1st intercubital nervure ; 
recurrent nervure interstitial ; pobrachial areolet of the hindwings 
shorter than half the prebrachial. Legs blackish or brown, the 
forelegs lighter, as well as the underside of all the femora, and 
the base of the tibiz. Abdomen depressed, shining ; 1st segment 
hardly longer than broad, striolate. Terebra not surpassing the 
apex of the abdomen, directed upwards. ¢ Similar; antenne 
longer than the body, 38-jointed. Length, 2; wings, 4 lin. 


I have taken this species in S. Wales, Wiltshire, and 
Devonshire. According to Haliday, it is rare in Ireland. 


2. Phenocarpa punctigera, Hal. 


Alysia punctigera, Hal., Ent. Mag., v., 238, 2; pl. xvii, 
f. 27 (forewing), f. 30 (hindwing). 

Fusco-castaneous with the hesd black, shining ; mandibles and 
palpi rufous. Antennz hardly as long as the body, pubescent, 
blackish, with the two basal joints rufous, 4th joint hardly elongate. 
Mesothoracic sutures incomplete ; a punctiform fovea before the 
scutellum ; furrow of the mesopleure finely traced, crenulate. 
Wings obscure ; squamula rufous ; stigma pale fuscous, oblong ; 
radial areolet cultriform ; 2nd abscissa curved at the base, hardly 
longer than the 1st intercubital nervure ; 3rd abscissa straight ; 
pobrachial areolet much longer than the przbrachial, being pro- 


British Braconide. ane 


duced in form of an obtuse lobe ; 2nd discoidal areolet small, 
nearly obsolete owing to the shortness of the two transverse 
nervures ; anal nervure interstitial ; pobrachial areolet of the hind- 
wings not half as long as the prebrachial; radial areolet almost 
bisected by an accessory transverse nervure. Legs rufous. 
First abdominal segment subbicarinate, reddish castaneous. 
Terebra as long as 3 of the abdomen. ¢ unknown. Length, 1}; 
wings, 23 lin. 

One specimen only was taken by Haliday in north 
Ireland, and given by him to Curtis; it is therefore pro- 
bably now in Australia. 


3. Phexnocarpa tabida, Nees, (Pl. XIL., fig. 3, 2). 


Alysia tabida, Nees, Mon., i., 252, ?; Hal., Ent. Mag., 
Vi oileOrk s 

Second discoidal areolet obsolete ; stigma much attenuated. @ 
Piceous, shining ; head and posterior part of the abdomen blackish, 
Ist segment rufo-testaceous ; mandibles and palpi rufous. Antenne 
one-half longer than the body, fuscous with the 4-5 basal joints 
testaceous, 19-21-jointed, all the joints long, the 4th very long. 
Prothorax rufous; mesothorax castaneous or rufescent ; sutures 
incomplete ; metathorax sometimes testaceous, nearly smooth ; a 
circular fovea before the scutellum. Wings hyaline; squamula 
rufous ; nervures and stigma dull testaceous; radial areolet cultn- 
form, attaining the tip of the wing ; 2nd cubital areolet elongate, 
much narrowed towards the outer extremity; anal nervure 
interstitial; hindwings narrow, pobrachial areolet very small. 
Legs testaceous. First abdominal segment uneven, hardly 
rugulose, bicarinate, testaceous, the following segments rufescent, 
becoming gradually darker to the apex, which is black. Terebra 
as long as 2 of the abdomen. ¢ Similar; antenne longer, 20-22- 
jointed. Length, 1; wings, 2} lin. 


Found very sparingly in Germany and England; I 
have taken a single example. 


4, Phenocarpa picinervis, Hal. 
Alysia picinervis, Hal., Ent. Mag., v., 233, 62%; pl. 
xvi, f. 29 (wing). 
Transverse nervures bordered with brown. @ Shining black, 


with a bronzed tint. Face punctulate, subcarinate ; mandibles and 
palpi rufous, the Jatter more obscure. Antenne slender black, 


628 Rey. ‘I’. A. Marshall’s monograph of 


rufescent at the base beneath, longer than the body, 27-jointed. 
Mesothoracic sutures crenulate, complete, meeting in a curve 
posteriorly ; between them is an impressed longitudinal line ; 
metathorax rugulose, smoother towards the base. Wings fusces- 
cent : squamula testaceous, nervures and stigma fuscous, the latter 
conspicuous, emitting the radial nervure from its posterior half; 
radial areolet cultriform, reaching the tip of the wing; 2nd cubital 
areolet elongate, very slightly narrowed externally ; 2nd abscissa 
twice as long as the 1st intercubital nervure; 2nd intercubital 
nervure more distinctly clouded with brown than the other 
nervures; pobrachial areolet of the hindwings very small. Legs 
dull rufous. Abdomen depressed ; Ist segment rather long, twice 
as wide at the extremity as at the base, finely striolate. Terebra, 
measured from the base, as long as the abdomen. ¢ Similar; 
antenue one-half longer than the body, or still longer, 30-32- 
jointed. Length, 13-1? ; wings, 3-4 lin. 

Rather common in damp shady places, in England and 
Ireland ; I have often captured it. 


5. Phxnocarpa ruficeps, Nees. 


Bassus ruficeps, Nees, Mag. Ges. Berl., 1814, p. 205. 

Alysia ruficeps, Nees, Mon., 1., 246; Hal., Ent. Mag., 
v., 233, 6%; pl. xvi, f 16 (wing); Bouché, 
Naturg, 1834, p. 147. 

Var. Bassus testaceus, Nees, Mag. Ges. Berl., 1814, 
p- 206. 

A. testacea, Nees, Mon., i., 246 3. 

A, gracilis and pallida, Cur., B. E., 141. 

A, oculator, Ratz., Ichn. d. Forst., ii., 71. 

Metathorax transversely subcarinate near the extremity, variable, 
head generally rufous, body black, or (in var. 3) body mostly 
testaceous; stigma rather stout, oval. Q Shining; stemmaticum 
black ; face punctulate, Antenne longer than the body, 22-27- 
jointed, two basal joints rufous, 4th very long. Mesothoracic 
sutures punctate anteriorly, less distinct behind, converging to a 
deep fovea before the scutellum; metathorax finely rugulose, 
smooth near the base, sometimes with a medial area more or less 
distinct. Wings hyaline ; squamula dull rufous; nervures and 
stigma fuscous, the latter determinately rounded at each end, 
stouter than in the allied spp. pécinervés and conspurcator (Nos. 4, 9); 
pobrachial areolet of the hindwings longer. Legs rufous or testa- 
ceous; tips of the tarsi obscure. Abdomen somewhat piceous, 
subcompressed ; Ist segment obconie, striolate. Terebra as long as 


British Braconide. . 529 


the abdomen, the valves ciliated. ¢ Similar; antenne much 
longer than the body, 25-32-jointed ; head more broadly rufous 
than in the 9. Length, 11-2; wings, 24-5 lin. 

Var. 1. ¢ Wings with a yellowish tinge, the nervures decolor- 
ous towards the tip. 

Var. 2. ¢@. Red colour of the head more or less extended on 
the thorax. 

Var. 3. ¢ 9 Head pale rufoas or testaceous; thorax and Ist 
abdominal segment the same, but more obscure; the rest of the 
abdomen fuscous or fuscescent. Alysia testacea, Nees. Two Qs 
which I received from Scotland belong to this variety ; their 
metathorax is almost smooth ; the mesothoracic sutures and the 
mesopleure fuscous. 

Var. 4. Head black. This variety is not uncommon ; it must 
be recognised by the subcarinate metathorax, and the thickness 
of the stigma. 

This very common species is known to be parasitic in 
the larve of Anthomyia radicum, L., Lonchea vaginalis, 
Fall., and Piophila casei, L. Ratzeburg must have 
been mistaken in supposing that his Alysia oculator 
came from the caterpillar of Tortrix rosana, L. 


6, Phenocarpa Maria, Hal. 
Alysia Maria, Hal., Ent. Mag., v., 237, ¢ 


Second cubital areolet short (for the genus), the 2nd abscissa 
being not much longer than the Ist intercubital nervure. ¢ 
Shining black ; mandibles rufescent. Antenne rather shorter than 
the body, filiform, 19-21-jointed. Mesothoracic sutures complete, 
punctulate, meeting posteriorly in an acute angle ; furrow of the 
mesopleure finely traced, crenulate ; metathorax smooth and 
shining. Wings hyaline, somewhat whitish ; squamula testaceous ; 
nervures and stigma fuscous, the latter small, acute at both ends ; 
Ist abscissa as long as the width of the stigma; radial areolet 
cultriform, attaining the tip of the wing ; recurrent nervure inter- 
stitial ; anal nervure subinterstitial ; pobrachial areolet of the hind- 
wings + as long as the prebrachial. Legs fuscous ; base of tibize 
rufescent. Abdomen attenuated at base and apex, wide in the 
middle ; 1st segment a little longer than its apical breadth, faintly 
striolate, longitudinally elevated in the middle. Terebra as long as 
Lof the abdomen. ¢ unknown. Length, 1}; wings, 4-3 lin, 


England and Ireland; very rare. I captured a ? at 
Sandwich, and another in an osier-bed in Wiltshire. 


530 Rey. T. A. Marshall’s monograph of 


7. Phenocarpa Galatea, Hal. 


Alysia Galatea, Hal., Ent. Mag., v., 238, ? pl. xvu, 
f. 17 (wing). 

Second cubital areolet elongate ; 2nd abscissa twice as long as 
the Ist intercubital nervure. @ Black or piceous, shining ; man- 
dibles rufescent. Antenne hardly as long as the body, 21-jointed. 
Mesothoracic sutures incomplete, scarcely inchoate ; a punctiform 
fovea before the scutellum; furrow of the mesopleure sub- 
obsolete, punctulate. Wings hyaline, whitish; squamula dull 
rufous ; stigma pale fuscous, narrower than in the last species ; 
2nd cubital areolet with parallel sides ; 1st cubital and 1st discoidal 
areolets indistinctly separated, the dividing nervure being nearly 
colourless ; recurrent nervure interstitial ; anal nervure not inter- 
stitial. Legs fuscous ; tibia, trochanters, and apex of fore femora, 
rufescent, Terebra exserted, short. d Unknown. Length, 12; 
wings, 23 lin. 


Taken by Haliday once only on Salix argentea, 
growing on a sandy shore near Dublin; otherwise 
unknown. 


8. Phexnocarpa Hugenia, Hal. (Pl. XII, fig. 4, 2.) 
Alysia Hugenia, Hal., Ent. Mag., v., 234, 6 2. 


Rufo-testaceous, shining ; stemmaticum, greater part of thorax, 
and first abdominal segment, fuscous. 9 Antenne testaceous, 
fuscescent towards the apex, twice as long as the body, 41-42- 
jointed, 4th joint a little longer than the 8rd, which is equal to the 
5th. Prothorax and anterior part of the pleurz rufo-testaceous ; 
mesothoracie sutures complete, smooth, meeting before the scutel- 
lum in an acute angle, which is punctate ; mesonotum impressed 
with some scattered punctures ; scutellum convex, smooth ; meta- 
thorax punctato-rugulose, smooth at the base. Wings hyaline 
with an obscure tinge ; squamula rufous; nervures brown; stigma 
yellow, rather narrow, attenuated outwards ; 2nd abscissa longer 
by a half than the first intercubital nervure ; recurrent nervure 
interstitial ; anal nervure not interstitial. Legs stout, rufous. 
Abdomen, after the Ist segment, rufo-testaceous; ist segment 
black, as long as its apical width, striolate, elevated in the middle. 
Terebra rather longer than half the abdomen, the valves bristly, 
broader towards the extremity. ¢ Similar ; antenne 49-jointed ; 
stigma fuscous, Length, 2; wings, 4 lin. 

Var. @ Head almost entirely pitchy black ; antenne somewhat 
shorter ; prothorax dark, except the pronotum ; sides and apex of 


British Braconide. 5381 


the abdomen fuscescent. Wings infumated; stigma fuscous. 
Length, 11; wings, 3 lin. Taken once in the London district by 
Walker. 


Extremely rare in north Ireland, according to Hali- 
day. <A single ? occurred in Cornwall, not far from 
my house. The size and colours render this species 
remarkable. 


9. Phenocarpa conspurcator, Hal. 

Alysia conspurcator, Hal., Ent. Mag., v., 236, 6 ?. 

¢ Black ; mandibles dull rufous ; palpi paler. Antenne some- 
what longer than the body, 24-30-jointed, the two basal joints dull 
rufous. Mesothoracic sutures complete, finely punctate, meeting 
in an acute angle posteriorly ; metathorax uneven, irregularly 
marked with a few elevated lines. Wings subhyaline ; squamula 
rufescent ; nervures and stigma fuscous, the latter very narrow, 
much attenuated outwards, where it is indeterminately confounded 
with the metacarp ; 2nd cubital areolet slightly narrowed exter- 
nally, the upper angle of its base often somewhat obtuse ; anal 
nervure not interstitial ; recurrent nervure interstitial. Legs dull 
rufous. First abdominal segment rather longer than its apical 
width, not shining, hardly rugulose, faintly carinate in the middle, 
the carina bifurcate near the base. ¢ Similar ; antenne longer 
and more slender, 32-jointed. Length, 14-24 ; wings, 3-5 lin. 

This may be known from its nearest allies by the 
narrowness of the stigma. It is, perhaps, the commonest 
species of the genus, noticed in England, Ireland, France, 
Holland, and probably occurring throughout Europe. 
The ? may be often seen on dunghills, and the droppings 
of cattle, in the fields, seeking for the larvee of Scato- 
phaga and other flies. 


10. Phenocarpa pratelle, Curtis (Pl. XII., fig. 6, 2). 


Alysia pratellz, Cur., B. H., 141 ; Hal., Ent. Mag., v., 
230, ¢. 


¢ Shining, pitchy black, the middie of the mesonotum some- 
times rufescent. Head large, deep black; clypeus and mandibles 
rufous. Antenne very slender, filiform, almost twice as long as 
the body, 33-35-jointed, blackish with the base rufous ; 4th joint 
elongate. Mesothoracic sutures complete, smooth; metathorax 
very short, uneven, slightly rugulose. Wings hyaline ; squamula 
testaceous ; nervures fuscescent ; stigma yellow, narrow, abruptly 


532 Rev. T. A. Marshall’s monograph of 


attenuated at the outer end; 2nd cubital areolet narrowed exter- 
nally ; recurrent nervure interstitial ; anal nervure not interstitial. 
Legs testaceous. First abdominal segment not much narrowed at 
the base, one-half longer than broad, very finely striolate, some- 
times rufous with the upper surface black. 'Terebra as long as the 
antenne, and much longer than the body. ¢ Similar; antenne 
41-jointed ; middle of the mesonotum (in my specimen), scutellum, 
and 1st abdominal segment rufous, the last blackish above. 
Length, 2 ; wings, 43 lin. 


Found ainongst fungi, in England and Scotland, but 


rarely; I captured a pair in Groveley Wood, near 
Salisbury. 


11. Phenocarpa Nina, Hal. 
Alysia Nina, Hal., Ent. Mag., v., 236, 2. 


2? Black ; mandibles dull rufous. Antenne as long as the body, 
22-jointed, blackish, brownish beneath at the base. Mesothoracic 
sutures incomplete ; discal fovea minute. Resembles flavipes (No. 
13), and conspurcator (No. 9). Haliday gives no further description 


of this species, except :—Length, 11; wings, 3 lin. 


In a note in the Ent. Maz., v., 236, Haliday speaks of 
the three species “anne Nina, and flavipes as 
requiring further examination, in order to separate them : 
we may conclude therefore that they are closely allied. 


This species has only been taken in the Hebrides, by 
Haliday. 


12. Phexnocarpa Eunice, Hal. 
Alysia Eunice, Hal., Ent. Mag., v., 235, ¢. 


? Black ; face punctulate ; eyes slightly pubescent ; mandibles 
rufous ; calpi dusky. Antenne as long as the body, brown at the 
base underneath, 25-jointed, 4th joint not elongate, 3rd a little longer 
than the 5th. Mesothoracic sutures inchoate ; a short fovea before 
the scutelluam. Wings infumated; squamula pale testaceous ; stigma 
narrow ; pobrachial areolet of the hindwings very small. Legs dull 
rufous. Terebra longer than the abdomen. 

The above is Haliday’s description, to which I add the diagnosis 
of an insect which I regard as the ¢, although in such a case there 
can be no certainty. Black ; antennz almost twice as long as the 
body, 30-jointed, fuscous, with the two basal joints testaceous. 
Mesothoracic sutures inchoate ; a shallow circular fovea before the 


British Braconidex. 53 i) 


scutellum. Wings infumated ; stigma narrow, etc., as in the ?. 
Length, 2 ; wings, 43 lin. 


The ¢ was found rarely in Ireland by Haliday ; I cap- 
tured the ¢ near Salisbury. 


13. Phenocarpa flavipes, Hal. 
Alysia flavipes, Hal., Ent. Mag., v., 286, ?. 


Q Pitchy black, shining ; clypeus and mandibles rufous ; palpi 
yellowish. Antenne longer than the body, with the first 9 or 10 
joints testaceous, 29-32-jointed. Mesothoracic sutures very faint 
or effaced posteriorly; a fovea before the scutellum situated in a 
bilobed rufous spot ;metathorax uneven, shining. Wings hyaline ; 
squamula yellowish ; nervures and stigma fusco-testaceous, the 
latter distinct, cutting off the greater part of the Istabscissa ; 2nd 
cubital areolet narrowed externally ; recurrent and anal nervures 
interstitial. Legs flavo-testaceous. Abdomen depressed; Ist 
segment a little longer than its apical breadth, not much narrowed 
towards the base, faintly striolate; 2nd segment sometimes with a 
transverse yellowish band at the base. Terebra as long as 3 of the 
abdomen. ¢ Similar; antenne twice as long as the body, 30- 
jointed in my specimen ; stigma much attenuated outwards, con- 
founded with the metacarp; 1st abscissa short, but visible. 
Length, 14-13; wing, 2}-3? lin. 

Noticed in Ireland, England, and Holland; I took the 
3 at Bugbrooke near Northampton. 


14, Phenocarpa livida, Hal. 
Alysia livida, Hal., Ent. Mag., v., 237, de. 


Abdomen, after the 1st segment, castaneous red or rufotes- 
taceous. @ Pitchy black; oral parts rufous. Antenne almost 
twice as long as the body, 27-28-jointed, blackish with the 3 or 4 
basal joints testaceous ; 4th joint not elongate, 3rd and 5th sub- 
equal. Prothorax testaceous ; mesothoracic sutures inchoate ; an 
oval fovea before the scutellum ; metathorax dull, finely punctato- 
rugulose. Wings hyaline; squamula yellowish ; nervures and 
stigma pale fuscous or testaceous, the latter narrow, attenuated 
outwards and confounded with the metacarp; 2nd cubital areolet 
narrowed towards the extremity ; recurrent nervure interstitial ; 
anal nervure almost interstitial; pobrachial areolet of the hind- 
wings obsolete. Legs yellowish, Abdomen depressed; Ist seg- 
ment blackish, dull, very finely punctato-rugulose. Terebra as 


534 Rev.'T’. A. Marshall’s monograph of Brit. Braconide. 
long as 2 of the abdomen. € Similar; antenne three times as 
long as the body, 3l-jointed ; a bilobed rufous patch on the 
mesonotum (in my specimen); abdomen, after the 1st segment, 
testaceous with the apical segments infuscated. Length, 1-14; 
wings, 24-3 lin. 

Rare in England and Ireland, according to Haliday ; I 
possess a d from the New Forest, and a ¢ captured 
near Salisbury. 


(To be continued.) 


EXPLANATION oF Pirates XI. & XII. 


PLATE XI. 


Fic. 1. Panerema inops, First. ¢ 
1 a. Abdomen viewed sideways. 
1 b. Wings. 


. Trachyusu Aurora, Hal. g 
3. Allea contructa, Hal. ?. 
3a. Male. 
3b. Wings of ¢. 
4, Alysia manducator, Panz.°?. 
4a. Antenna (after Curtis). 


4 b. Mandible. 
4c. Abdomen viewed sideways. 


bo 


PLATE XII. 


Fig. 1. Alysia tipule, Scop. 9. 
. Idiasta maritima, Hal. ¢. 
. Phenocarpa tabida, Nees, 9. 
. Phenocarpa Eugenia, Hal. 9. 
4a. Wing. 
5. Wing of Phenocarpa pullata, Hal. 
6. Phonocarpa pratelle, Hal. 9. 


moh 


€, 535: 4) 


XXI1. Catalogue of the Pterophoride, Tortricide, and 
Tineide of the Madeira Islands, with notes and 
descriptions of new species. By the Right Honble. 
Lorp Watsincnay, M.A., LL.D., F.R.S. 


[Read Oct. 3rd, 1894.] 


A sMALL collection of micro-lepidoptera made by the late 
T. Vernon Wollaston in Madeira, has been submitted 
to me by Mr. Bethune-Baker, together with some 
drawings by the late Professor Westwood of the types 
described by Wollaston and Stainton now in the British 
Museum. ‘lhe specimens, unfortunately, are not in good 
condition, and the majority of the species have been 
already dealt with in two papers published by those 
authors [Wollaston, T. V., “ Brief Diagnostic Characters 
of Undescribed Madeiran Insects,” Ann. and Mag. N.H. 
(3s.), I, 118—124; London, 1858. Stainton, H.'T., “ Notes 
on Lepidoptera collected in Madeira by T. V. Wollaston, 
Hsq., with Descriptions of some New Species,” Ann. and 
Mag. N.H. (8 s.), IL., 211—214; London, 1859]. 

Such as are new are described below. 

The opportunity seems to be favourable for compiling 
a complete list of the species known to occur in the 
Madeiras, which will, perhaps, form a useful parallel to 
the catalogues of micro-lepidoptera of the Canaries, lately 
published by Dr. Rebel [Rebel, H., “ Beitrag zur Micro- 
lepidopterenfauna des canarischen Archipels,’ Ann. 
K. K. Hofmus. VII., Heft., 3, 241—284, Taf. XVII., 
Wien, 1892. “Zur Lepidopterenfauna der Canaren,” 
Ann. K. K. Hofmus. IX., Heft. 1, 1—96, Taf.. L, 
Wien, 1894]. 

Mrs. Holt White has lately published a catalogue of 
the lepidoptera of Teneriffe [White, A. H. Holt, edited 
by White, R. Holt, “The Butterflies and Moths of 
Teneriffe,” pp. i— xii. and 1—107, Pl. I—IV.; London, 
1894], which, though dealing specially with the larger 
lepidoptera, gives lists of the few micros known to occur 
in that island. 

TRANS. ENT. soc. LOND. 1894,—pPapRr iv. (DEC.) 


536 Lord Walsingham’s catalogue of the Pterophoride, 


Mrs. Wollaston’s collections from St. Helena (also in 
the British Museum), of which descriptions were pub- 
lished by herself and the late Francis Walker [Walker, 
F., in Meliss’ Saint Helena; London, 1875. Wollaston, 
Mrs. T. V., “Notes on the Lepidoptera of St. Helena, 
with Descriptions of New Species,” Ann. and Mag., 
N.H. (5 s.), 111.; London, 1879: this was republished in 
pamphlet form], are also important contributions to the 
study of the subject as connected with these groups of 
Atlantic islands which he nearest to the African coast. 

The present paper shows that the genus Blastobasis 
and its allies are more than proportionately represented 
in the Madeiras, and the line of their distribution over 
the European and American continents is thus connected 
and emphasized. The genera represented are, for the 
most part, common to both Kurope and America, but the 
majority cf the species recugnized are Kuropean. One 
notable exception may be found in Cosmopteryx pul- 
cherrimella, Chamb., hitherto recorded only from the 
United States, this has been very carefully compared 
with American specimens in my collection. 

I have only to express my thanks to Mr. Bethune- 
Baker for kindly supplying me with so much interesting 
material, by which he has reinspired me with the hope 
of some day making a more complete and representative 
collection either personally or by deputy, in the Madeiran 
group of islands which I have long been anxious to 
visit. 

By the list given it will be seen that sixty-six species 
of micro-lepidoptera are now known to occur in the 
Madeiras (four are not described); of these, thirty are 
peculiar to these islands, twelve are common to the 
Madeiras and Canaries (of which two do not occur else- 
where), and one extends its range only to North Africa. 
One species is peculiar to the Madeiras and to the United 
States. Thiuty-two species occur in Europe, and of 
these, nine extend to the Canaries, thirteen to North 
America, and four to North Africa. 

Over thirty species are added to the list, one new genus, 
seven new species, and two new varieties being here 
described. 


Tortricide, and Tineidx of the Madeira Islands. 537 


mo 


PTEROPHORID. 
Oxyptilus, sp 
Platyptilia searanodbietalt “Hb. 
Alucita monodactyla, L. ... : 
TORTRICIDA. 
TORTRICINA. 
Tortrix subcostana, Stn. 
»  retiferana, ‘Stn. 


= subjunctana, Wistn. 
», reticulata, Stn. 


CONCHYLIDINA. 
Carposina atlanticella, Rbl. 


GRAPHOLITHINA. 


Bactra lanceolana, Hb. 
Peedisca carduana, Gn... 
Grapholitha madere, Wistn. 
Carpocapsa grossana, Hw.... 
splendana, Hb.. 
Steganoptycha signatana, Dgl 
” Sp. ». 
TINEID A. 
TINEINA. 
Trichophaga abruptella, hg 
Tinea pellionella, L... 
s,  nigripunctella, ‘Hw.. 
»,  irrorella, Wlstn. 


ad BD - eet ie 
Tineola biselliella, Humm... 
ae allutella, Rbl. 


Oenophila V-flavum, Hw. ... 
Bedellia somnulentella, Z 


HYPONOMEUTINA. 
Hyponomeuta bakeri, Wlsm. 
PLUTELLINA. 
Plutella cruciferarum, Z. ... 


GELECHIANA. 
Lita pulchra, Wlstn. 


5, submissella, Stn. ... iy 


», ocellatella, Stn. 

»,  portosanctana, Stn. ... 
Bryotropha domestica, Hw. 
Sitotroga cerealella, Oliv. ... 
Anacampsis albipalpella, H.S. 

PA elachistella, Stn. 
a wollastoni, Wlsm. 


GLYPHIPTERYGINA. 


Choreutis bjerkandrella, Thnb. 
Simaethis nemorana, Hb. . 

Fe oxyacanthella, Ee 
Glyphipteryx, sp. . 


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TRANS. ENT. SOC. LOND, 1894.—PART IV. 


10 


11 


12 


538 Lord Walsingham’s catalogue of the Pterophoride, 


3 ° 
a Se hee 4 
a fi/<af|4 53 
6|ala|a | ax 
DEPRESSARIAN A. cena (S| He 
40. Depressaria conciliatella, Rbl. 10 | 22] ... ¢ 
41. a applana, F. ... | 23 | 10 ; 
42. Siganorosis heracliana, D. eG 24 | 11 os 
(COPHORINA. 
43. C£cophora pseudospretella, Stn. ... 25 | 12 
BLASTOBASINA, 
44, Endrosis lacteella, Schiff. ... 26 | 13 ood 
45. Blastobasis lavernella, Wlsm. eta ere 16 
46. AS decolorella, Wlstn. rea oer ee 17 
47. e fuscomaculella, Rag... es ee | er ae 
48. 3 desertarum, Wistn. : Se ees 18 
48a. i desertarum + radiata, Wlsm. 18A 
49. an lignea, Wlsm. . ee 19 
49a. A, lignea + adustella, Wism. 194 
50. 5 ochreopalpella, Wistn. 20 
51. as marmorosella, Wlstn... 21 
52. 55 nigromaculata, Wlstn. ... | wee 22 
53. aurantiaca. sWAStn acc. 0 eeermune maeel eines .| 28 
54. Epistetus divisus, Wlsm. .| 24 
5b. by pica, Wlsm. 225 
56. Hypatima fasciata, Stn. .| 26 
LAVERNINA. 
57.4 lbaverna vittabaAWilstms «hs..¢) - ea tebhn ee. eterearin ets ae 97 
58. rebeli, ’Wism. Ae 28 
59. Chauliodus daucellus, nae 28 ie 
60. Asychna insularis, Wlstn.. oe 29 
COSMOPTERYGIN A. 
61. Cosmopteryx pulcherrimella, Chamb. 14 
GRACILARIANA. 
62. Gracilaria staintoni, Wlstn. dl retail pecs use AN Sti) 
63. Re roscipennella, Hb. .| 12 | 29 neo || od 
64. scalariella, Z. ... .. | 30 ate . 
65. Lithocolletis messaniella, Z. 31 5 
LYONETIANA. 
66. Phyllobrostis daphneella, Stgr. ... 32 
66. 12 | 32] 14/ 5 | 380 
PTEROPHORID 4. 
Oxyetitus, Z. 
Oxyptilus, sp. 
Madeira; one specimen. 
Puatyptittia, Hb. 
Platyptilia acanthodactyla, Hb. 
Pterophorus acanthodactylus, Stn., Ann. and Mag. 


N. H. (3:s.), IID., 214 (1859). 


Madeira ;' Funchal, eight specimens (Baker). 


Tortricidx, and Tineidx of the Madeira Islands. 539 


Auveira, L. 
Alucita monodactyla, L. 


Pterophorus pterodactylus, Stn., Ann. and Mag. N. H. 
(3 s.), IIL., 214 (1859).1 

Pterophorus monodactylus, Rbl., Ann. K. K. Hofmus. 
VII., 263, 282 (1892). 

Madeira ;' five specimens (Baker). 


TORTRICID. 
TORTRICINA:. 


Tortrix, L. 
Tortriv subcostana, Stn. 


Tortrie subcostana, Stn., Ann. and Mag. N. H. (8 8.), 
TS SLL (1859) Wikes Cat. lapelins. Bs Ms 
XXVIII, 826 (1863). 

Madeira ;! San Antonio da Serra, five specimens 
(Baker). 

This species is mentioned by Dr. Rebel as occurring 
in the Canary Islands [Ann. K. K. Hofmus. IX., 16, 81-82 
(1894)] on the evidence of specimens received by me 
from Teneriffe, collected by Mr. Leech in April, 1885, 
but a careful comparison leads me to regard these as 
distinct from the true Madeiran type. They are some- 
what smaller, and have invariably a strong indentation 
on the inner side of the transverse fascia above the 
middle of the wing which I am unable to find in any 
specimens from Madeira; moreover, in the absence of a 
costal fold in the 3, it cannot be rightly placed in the 
genus Cacoecia. 


Tortrix retiferana, Stn. 
Tortria retiferana, Stn., Ann. and Mag. N. H. (8 s.) 


III., 211-12 (1859) ;! Wkr. Cat. Lip. Ins. B. M. 
XXVIIT., 326 (1863). 


Madeira ;' The Mount, five specimens (Baker). 


Tortria subjunctana, W\stn. 


Tortriz subjunctana, Wistn., Ann. and Mag. N. H. 
(diss)j To -E201(k858)s* Wr, Cat: Ip. Tuas Be M., 
XXX., 985 (1864), 


Madeira,! 


540 Lord Walsingham’s catalogue f the Pterophoridz, 


Tortria (?) reticulata, Stn. 


Tortria (?) reticulata, Stn., Ann and Mag. N. H. (3s.), 


II., 211 (1859) ;! Wkr. Cat. Lp. Ins. B 
XXVIIT., 326 (1863). 


1 ae 
Madeira;+ San Antonio da Serra, one specimen 


(Baker). 


CONCHYLIDIN. 
Carposina, H.S. 
Carposina atlanticella, Rb). 


Carposina atlanticella, Rbl., Ann. K. K. Hofmus. IX., 
92 (1894). . 


San Antonio da Serra; three specimens (Baker). 
Madeira, May; six specimens. (Leech, Mus. 
Wlsm.) 

GRAPHOLITHINA. 
Bacrra, Stph. 


Bactra lanceolana, Hb. 
San Antonio da Serra; four specimens. 


Papisca, Ld. 


Pedisca carduana, Gn. 


Madeira; three specimens (Baker). Mr. Leech also 
met with this in Madeira in May. 


GRAPHOLITHA, T'r. 
Grapholitha madere, Wlstn. 


Ephippiphora maderx, Wlstn., Ann. and Mag. N. H. 
(3 s.), I., 120 (1858).1 


Grapholita madere, Wkr. Cat. Lp. Ins. B. M., XXX., 
990 (1864).? 


Grapholitha madere, Rb]., Ann. K. K. Hofmus. IX., 
17, 87-88 (1894).8 
Madeira ;! Funchal and The Mount, eleven specimens 
(Baker). Canaries.? I have specimens taken by Mr. Leech 
at Madeira in April, and at Teneriffe in May. 
Carpocapsa, Tr. 


Carpocapsa grossana, Hw. 
Madeira; one specimen. 


Tortricide, and Tineide of the Madeira Islands. 541 


Carpocapsa splendana, Hb. 
Madeira ; two specimens. 


SreGANoPTYCHA, Stph. 
Steganoptycha signatana, Del. 
Madeira ; The Mount, one specimen. 


Steganoptycha, sp. 
Madeira; one specimen. 


TINEID&. 
TINEINAL. 
TricHopHaGca, Rag. 
Trichophaga abruptella, W1stn. 

N. syn. = Tinea bipartitella, Rag. 

Tinea abruptella, Wistn., Ann. and Mag. N. H. (3s.), 
I., 120 (1858) ;' Wkr., Cat. Lp. Ins., B. M., XXX., 
1003 (1864). 

Tinea bipartitella, Rag. Bull. Soc. Ent. Fr, 1892, 
Ixxxill. 

Tinea tapetzella, Rbl., Ann. K. K. Hofmus. VII., 268-9, 
283 (1893) : IX., 17 (1894). 

Trichophaga bipartitella, Rag. Ann. Soc. Ent. Fr. 
LXIIL., 122-4: (1894). 

Porto Santo.' 


The synonymy is verified by comparison with Ragonot’s 
type from Tunis; I have also a specimen from Hgypt. 

‘This species differs from tapetzella, with which Dr. 
Rebel appears to have wrongly confounded it, in the 
straighter outer edge of the dark portion of the wing, 
which is not produced along the costal margin. I have 
seen no Huropean specimen possessing this character. 


Tinga, L. 
Tinea pellionella, L. 
Tinea pellionella, Stn., Ann. and Mag. N. H. (3 s.), 
III., 212 (1859) ;” Rbl., Ann..K.. K. Hofmus. VIL, 
283 (1892): IX., 17 (1894). 
Madeira ;' nine specimens (Baker). 
Tinea nigripunctella, Hw. 
Tinea nigripunctella, Wlism., Ent. Mo. Mag., XXX., 
50 (1894)'. 
Madeira ;! four specimens (Baker). 


542 Lord Walsingham’s catalogue of the Pterophoride, 


Tinea trrorella, W\stn. 

Tinea irrorella, Wistn., Ann. and Mag. N. H. (3:s.), L., 
120 (1858);' Wkr.{-Cat. lip. Tnes-B: Si eax x, 
1002 (1864). 

Madeira. * 

Tinea, sp. 

Madeira ; four specimens. 


These are closely allied to irrorella, Wlstn., but differ 
in having smaller and more scattered markings and no 
apical spot ; they are not in good enough condition for 
description. 


Trnrota, H.S8. 
Tineola biselliella, Humm. 
Madeira ; four specimens. 
Tineola allutella, Rbl. 


Tineola allutella, Rbl., Ann. K. K. Hofmus. VI., 270-71, 
Pl. Vil 3.(1892)2 


Canaries! Madeira; one specimen (Baker), 


CHinopHiza, Stph. 
Ginophila V-flavum, Hw. 


Oinophila flava, Stn., Ann.and Mag. N. H. (3 s.), III., 
214 (1859).' 


Madeira ;’ four specimens (Baker). 
Bepe.iia, Stn. 
Bedellia somnulentella, Z. 


Bedellia somnulentella, Stn., Ann. and Mag. N. H. 
(3 8.), IIT., 214 (1859). 
Madeira.’ 


HYPONOMEUTINA. 
Hyronomevta, Z. 


Hyponomeuta bakeri, Wl1sm., sp. n. 
Antenne slightly serrate in ¢; brownish cinereous. Palpi 
porrect, somewhat recurved, cinereous, Head greyish fuscous, 
posteriorly tinged with ochreous, face whitish. Thorax olivaceous. 


Tortricide, and Tineide of the Madeira Islands, 543 


Forewings rather shining, olive-brown, with a distinct white streak 
running from the base to above the anal angle, and containing a 
blackish patch beyond its middle ; this streak is narrow at the 
base, and slightly widened outwards, ending abruptly, but con- 
nected in one specimen by a few whitish scales to the pale base of 
the cilia; before the apex the costal margin is narrowly whitish, 
and the basal half of the cilia around the apex, to a point opposite 
to the end of the white streak coming from the base, is also whitish, 
the outer half of the cilia being brownish-grey. Hindwings 
brownish grey, cilia scarcely paler, with a slight shade running 
through them near their base. Abdomen brownish-grey. Legs 
greyish-cinereous. Hap. al., 20 mm. 

Popes s ¢.. Mus. Wism. 

Hab. Madeira; San Antonio da Serra, five specimens. 

This species is nearly allied to egregiellus, Dp., but’ the 
points of difference appear to be constant, the black 
spot in the white “length-streak ”’ is smaller in eyre- 
giellus, and does not fill up the whole width of the 
streak, the colour of egregiellus is much greyer, less 
brown, there is no conspicuous white shade along the 


base of the cilia, and, moreover, the palpi are shorter 
and more dependent. 


PLUTELLIN. 
Piure.ia, Schrk. 


Plutella cruciferarum, Z. 


Plutella cruciferarum, Stn., Ann. and Mag. N. H. (8s.), 
ITI.; 212 (1859) ;!} Rbl., Ann., K. K. Hofmus. VII., 
283 (1892): IX., 167 (1894). 


Madeira;’ Funchal, and San Antonio da Serra, four 
specimens (Baker). 
GELECHIAN. 
Lana, fr: 
lita pulchra, WIstn. 


Gelechia pulchra, Wistn., Ann. and Mag. N. H. (3 s.), 
i, 22d, (1858)3" “Wks.;, Gat.’ Lip. edng.w By M., 
XXIX., 627 (1864). 


Deserta Grande.’ 


544 Lord Walsingham’s catalogue of the Pterophoride, 


Lita submissella, Stn. 


Gelechia submissella, Stn., Ann. and Mag. N. H. (3:8.), 
IIT., 212-13 (1859) ;' Wkr., Cat. Lp. Ins. B. M., 
XXIX., 628 (1864). 


Porto Santo.’ 
Lita. ocellatella, Stn. 
Gelechia ocellatella, Stn., Ent. Ann. 1859. 151-2 
(14. XIT. 1858):" Ann. and Mag. N. H. (8 s.), 


III., 212 (1859);? Wkr., Cat. Lp. Ins. B. M., 
XXIX., 628 (1864). 


Porto Santo."” 
Inta portosanctana, Stn. 


Gelechia portosanctana, Stn., Ann. and Mag. N. H. 
(3 s.), JIL., 212 (1859) 7 Wkr., Cat. Lp. Ins. B. M., 
XXIX., 628 (1864). 


Porto Santo.’ 
BryorropHa, Hein. 


Bryotropha domestica, Hw. 
Madeira ; one specimen. 


Sirrotroaa, Hein. 
Sitotroga cerealella, Oliv. 


Madeira; Funchal, four specimens. 


Anacampsis, Crt. 
Anacampsis albipalpella, H.S. 


= Gelechia anthyllidella, Stn., Ann. and Mag. N. H. 
(3 s.), ILI., 213 (1859)2 


Madeira ;! one specimen (Baker). 


Anacampsis (?) elachistella, Stn. 


Gelechia elachistella, Stn., Ann. and Mag. N. H. (8 s.), 
TID., 213 (1859)) Wir. (Cat. lip. Ios. 3. Mi, 
XXIX., 628 (1864). 


Northern Deserta. 


The type is unset, but it is apparently an Anacampsis. 


Tortricide, and Tineide of the Madeira Islands. 545 


Anacampsis wollastoni, Wism., sp. n. 


Antenne whitish. Palpi white, the second joint shaded with 
brown externally to near its apex, which is slightly tufted. Head 
smooth, shining white. Thorax whitish, tegule brown, with a 
creamy white streak along their inner edge. Forewings narrow, 
elongate, acuminate ; brown, with a narrow creamy-white streak 
along the dorsal margin extending to the apex and including the 
cilia, but interrupted beyond the anal angle by two oblique brown 
streaks, the first along the apical margin, the second, shorter, in the 
cilia; beneath the apex a slight darkish line in the base of the 
cilia around the apex is preceded by a faint indication of a pale, 
very oblique line, commencing at the anterior extremity of the 
costal cilia and terminating below the apex of the wing; but this 
is not observable in worn specimens. Héindwings shining, pale grey, 
cilia pale ochreous. Abdomen shining, greyish-ochreous. Legs 
greyish-ochreous. xp, al., 14-15 mm. 


Type. &a¢. Mus. Wlsm. 
Hab. Madeira; seven specimens. 


GLYPHIPTERYGIN4. 


Cuoreutis, Hb. 


Choreutis bjerkandrella, Thnb. 
Madeira; Funchal, three specimens. 


SimaEtuis, Leach. 
Simaethis nemorana, Hb. 
Madeira; The Mount, seven specimens. 


Simaethis oxyacanthella, L. 


= Simaéthis fabriciana, Stn., Ann. and Mag. N. H. 
(3 s.), IIL., 210 (1859).* 


Madeira.! 


GuypHipreryx, Hb. 


Glyphipteryx, sp. 
Madeira ; Funchal, one specimen. 


546 Lord Walsingham’s catalogue of the Pterophoride, 


DEPRESSARIAN. 


Depressaria, Hw. 


Depressaria conciliatella, Rbl. 


Depressaria conciliatella, Rbl., Ann. K. K. Hofmus. 
VIL., 272-74, 288, Pl. XVII., 14 (1892).} 


Sicily,! Canaries ;! Madeira, Funchal, and The Mount, 
fourteen specimens (Baker). 

If Iam right in the identification, this appears to be 
an exceedingly variable species, a long series showing 
great variation in colour from pale stone-grey to fawn- 
brown, with intermediate mottled varieties, having much 
the appearance of assimilella, Tr. 

Depressaria applana, F. 
Madeira; one specimen. 
I am unable to distinguish this specimen from the 


common English species, but the antennz are somewhat 
broken towards their ends. 
Sreanorosis, Wlgrn. 
Siganorosis heracliana, D.G. 


Madeira ; one specimen. 


G@COPHORIN 22. 
CicorHorA, Stn. 


(Hcophora pseudospretella, Stn. 


Madeira; one specimen. 


BLASTOBASIN. 


Enprosts, Hb. 


Eindrosis lacteella, Schiff. 


= Endrosis fenestrella, Stn., Ann. and Mag. N. H. 
(3 s.), ILL., 213 (1859).? 
Madeira;! San Antonio da Serra, seven specimens 
(Baker). 


Tortricide, and Tineidex of the Madetra Islands. 547 


Brastopasis, Z. 


Ayeins 3 and 4 of the hindwings coincident, 5 from the same 
point as 3-+-4. 


Blastobasis lavernella, Wism., sp. n. 


Antenne notched in the g¢ above the basal joint ; greyish, basal 
joint ochreous. Palpi ochreous, the outer side of the second joint 
shaded with greyish-brown along its lower half nearly to the apex, 
on the upper side paler than the apical joint. Mead and Thorax 
ochreous, tegule shaded with greyish-brown. Forewings ochreous, 
mixed with reddish-ochreous mottlings, and shaded with patches of 
greyish-brown, forming two irregular oblique fasciaform bands on 
the basal half of the wing and two waved streaks on the apical 
half ; the first oblique band passes outwards to the costal margin 
before the middle from about the inner third of the fold ; the 
second, forming a patch on the dorsal margin at one-third, runs 
parallel with the first to the costa about the middle, or extending 
slightly beyond it, the two are sometimes connected by a slight 
cross-streak on the dise ; at the commencement of the costal cilia 
a greyish-brown quadrangular spot, narrowly produced outwards 
beyond the cell, reverts to the anal angle joining a spot of the 
same colour, which is produced upwards and inwards to the end of 
the cell ; between this and the outer of the two oblique fasciz lies 
a small spot of the same colour immediately above the fold at two- 
thirds from the base; cilia greyish-ochreous with a greyish-brown 
line running around the apex and apical margin near their base, 
but dying out before reaching the anal angle ; this is preceded by 
a faintly-indicated marginal line on the wing within it, Hindwings 
shining, pale stramineous ; cilia pale ochreous. Abdomen shining, 
pale stramineous, the segments indicated by greyish lines. Legs 
shining, pale ochreous. Hap. al., 15-16 mm. 


Type. 8. Mus. Wlsm. 
Hab. Madeira; three specimens. 


This species is apparently allied to Blastobasis de- 
colorella, Wlstn., but has somewhat deeper colouring, 
with a different arrangement of its more pronounced 
markings, and has much the general appearance of 
Laverna ochraceella. 


548 Lord Walsingham’s catalogue of the Pterophoride, 


Blastobasis decolorella, Wl|stn. 


Laverna (?) decolorella, Wistn., Ann. and Mag. N. H. 
(3 s.), I., 122 (1858) ;} Wkr., Cat. Lp. Ins. B. M., 
XXX., 885 (1864); Rbl., Ann. K. K. Hofmus. 
IX., 92-8 (1894). 

Madeira ;' Funchal, and San Antonio da Serra, eighteen 


specimens (Baker). Mr. Leech also took this species at 
Madeira in May. 


This species is a true Blastobasis ; its neuration is as 
follows :— 


Forewings, 12 veins; 7 and 8 stalked, both to above apex. 
Hindwings, 7 veins ; 3 and 4 coincident, 5 from the same point as 
3-+-4, 6 and 7 separate and parallel. 


It exhibits considerable variation in size, in colour, 
and in the distinctness or suffusion of the spots and 
markings. The drawing made by Westwood was 
evidently taken from one of the more reddish-ochreous 
specimens, whereas the more prevalent form appears to 
be whitish-ochreous or whitish-fawn-colour with brown- 
ish and umber spots and scales. The slender dark lines 
across the abdominal segments are persistent throughout 
the series, but these occur also in other species, and must 
not be too much relied on. A description of the well- 
marked paler forms may be useful. 


Antenne distinctly notched and finely ciliate in the g; basal 
joint enlarged, pectinate beneath ; brownish-grey, whitish-fawn 
beneath. Palpi whitish-fawn, second joint externally brown except 
at its apex. Head, face, and thorax whitish-fawn. Forewings 
narrow, elongate, lanceolate towards the depressed apex ; very pale 
fawn, slightly suffused with fawn-brown scales across the middle 
and at one-third of the fold; an umber patch on the middle of 
the dorsal margin, above which is an umber spot on the disc before 
the middle of the wing, followed by a second spot of the same 
colour below and beyond it ; two umber spots at the end of the 
cell, the lower more conspicuous than the upper and almost parallel 
with it; these are followed by a sinuate brownish shade com- 
mencing on the costa in the beginning of the costal cilia and ending 
at the anal angle, much bowed outwards in the middle ; towards the 
apex of the wing a few slight brownish dots and scales are scattered 
about the apex and apical margin ; cilia very pale fawn. Hind- 
wings shining, pale fawn-grey, cilia very pale ochreous. Abdomen 


Tortricidx, and Tineidex of the Madeira Islands. 549 


pale greyish-fawn, each segment narrowly margined with blackish. 
Legs whitish-fawn, the anterior pair much shaded with umber- 
brown. zp. al., 17-22 mm. 

In some specimens the umber dorsal patch is almost 
entirely obliterated, the markings fainter and more suf- 
fused, and the dots around the apex and apical margin 
coalescing and forming a brownish-grey line along the 
base of the cilia, which is reduplicated along their middle 
towards the anal angle. 


Blastobasis fuscomaculella, Rag. 

N. syn. = seevoldiella, Kreithn. = marmorosella, Rbl. 
(nec. Wlstn.). 

(Heophora fuscomaculella, Rag., Bull. Soc. Ent. Fr., 
13795 ex,’ 

(icophora seeboldiella. Kreithn., Sitzb. Z.-B. Ges. 
Wien, XXXI., 20-1 (1881). 

Blastobasis marmorosella, Rbl., Ann. K. K. Hofmus. 
Wil; 276-78, 288), PLO XVI yo 302)" Ex, 
18, 90-1 (1894). 

Portugal,! Spain,” Canaries.3 Madeira (British 
Museum). 

The types of Gcophora marmorosella, Wlstn., in the 
British Museum, show that this species has veins 5 and 
3+4 of the hindwings distinctly stalked, whereas 
(cophora seeboldiella, Kreithn., from Bilbao, which Dr. 
Rebel has identified with this species, has these veins 
arising from a point, and therein agreeing with a larger 
and wider-winged species in the British Museum, and 
with a single specimen in my own collection from the 
Canaries which was determined for me by Dr. Rebel as 
marmorosella. 

The name fuscomaculella must stand for the larger 
form, and marmorosella for the smaller and narrower- 
winged species. I have seen the type of fuscomaculella, 
Rag., and consider it the same as seeboldiella, Kreithn., 
and marmorosella, Rb]. (nec. WIstn.). 


B-veins 5 and 4 of the hindwings coincident; 5 and 3+4 
stalked. 
Blastobasis desertarum, Wlstn. 
N. syn. = Ptervlonche (?) maderensts, Stn. 


Coleophora desertarwm, Wlstn., Ann. and Mag. N. H, 
(3 s.), I., 122 (1858). 


550 Lord Walsingham’s catalogue of the Pterophoride, 


Pterolonche (?) maderensis, Stn., Ann. and Mag. 
N.“H. (3 s.), TI., 203 4859)? W kr, Cat. ip. ins 
B. M.. XXIX., 666 (1864). 

Coleophora desertorum, Wkr., Cat. Lp. Ins. B. M., 
XXX., 879 (1864).4 

Northern Deserta,"? Porto Santo;? Funchal, nine 
specimens (Baker). 

The specimens of Coleophora desertarum in the British 
Museum are small and somewhat more strongly marked 
varieties, but are evidently the same _ species as 
maderensis. 


Blastobasis desertarum, Wlstn. + radiata, Wlsm., var. n. 


Antenne notched at the base in the ¢, basal joint enlarged, 
with a pecten ; fawn-grey. Palpi porrect, the second joint pro- 
jecting beyond the head and clothed with loose scales beneath 
towards the apex, apical joint short, not recurved ; whitish. Head 
and Thorax whitish, the latter shaded with pale fawn-brown. 
Forewings whitish, a pale fawn-brown streak extending from the 
base immediately below the costal margin and gradually widening 
outwards to the end of the cell, whence it is continued to the apex, 
covering half of the outer portion of the wing, the costal margin 
above it being narrowly pale beyond the middle ; a second pale 
fawn-brown streak from the middle of the base extends to the 
anal angle and along the base of the cilia nearly to the apex, but 
not extending beneath the fold, and having above the anal angle 
a small blackish dot opposite to a still smaller one at the lower 
margin of the upper streak; there is also an indication of a 
blackish dot near the outer end of the fold, and the pale inter- 
mediate spaces are slightly speckled with blackish scales ; cilia 
fawn-grey. i/indwings shining, fawn-white; cilia pale whitish- 
ochreous. Abdomen whitish-ochreous, the segments indicated by 
slender dark cross-lines. Legs whitish-ochreous, hind tibiz clothed 
with long loose hair-like scales. Hyxp. al, 14 mm. 

Type. o&. Mus. Wism. 


Hab. Madeira; Funchal, one specimen. 


Blastobasis lignea, W\sm., sp. n. 


Antenne pale greyish-fuscous ; basal joint enlarged, tufted and 
notched in the male, pale cinereous. Pulp? pale cinereous, shaded 
and mottled with greyish-fuscous externally. Head and Thorax 
pale cinereous, the latter shaded with greyish-fuscous. Forewings 
pale cinereous, much shaded and speckled with greyish-fuscous, 


wad 


Tortricide, and Tineide of the Madeira Islands. 551 


with a slight purplish gloss ; five small blackish spots (the first on 
the disc before the middle is sometimes connected with one on the 
middle of the fold below, and somewhat before it ; a small one on 
the lower edge of the cell lies halfway between these and two 
parallel spots, the one above the other, at the outer end of the 
cell) ; around the apex and apical margin is a series of six or eight 
ill-defined greyish-fuscous spots along the base of the cilia which 
are greyish-cinereous, somewhat darker towards the apex. Hind- 
winys very pale cinereous with pale greyish-cinereous cilia. 
Abdomen missing. Legs pale cinereous, somewhat speckled and 
shaded with greyish-fuscous externally. Hxp.al., ¢ 14, 2 19 mm. 


Type. &82.Mus. Wlsm. 
Hab. Madeira; sixteen specimens. 


Blastobasis lignea, Wism. + adustella, Wlsm., n. var. 


In a single specimen the darker shading is more 
conspicuous, and the two inner spots are merged in an 
angulated fascia, leaving the dorsal margin at one-third 
and tending obliquely outwards to the disc before the 
middle, where it is angulated back towards the costa 
from what should be the position of the upper spot; 
but, before reaching the costa, it is again bent upwards 
and slightly outwards to the margin. 


Abdomen very pale cinereous, the segments marked by narrow 
greyish fuscous transverse lines. Exp. al., 19 mm. 

Type. <2 ..Mus., Wism. 

Hab. Madeira; one specimen. 

Intermediate varieties in which the fascia is slightly 
indicated appear to occur, but I have no specimens 
before me in condition for comparative description. 


Blastobasis ochreopalpella, W\stn. 


(Keophora ochreapalpella, Wistn., Ann. and Mag. N. H. 
(3.8:)5 Le, 12101858) 5h Wir. Cat Ep. Ins. BM; 
XXX., 1029 (1864). 

Madeira.' 


Blastobasis marmoreusella, W\stn. 


(Ecophora marmorosella, Wlistn., Ann. and Mag. N. H. 
(3.s.), I., 128 (1858) ;! Wkr., Cat. Lp. Ins. B. M. 
XXX., 1028 (1864). 


502 Lord Walsingham’s catalogue of the Pterophoride, 


Porto Santo,! Northern Deserta.! 


Distinct from seeboldiella, Kreithn. See remarks under 
fuscomaculella, Rag. (ante p. 549). 


Blastobasis nigromaculata, W|stn. 


Gelechia nigromaculata, Wlstu., Ann. and Mag. N. H. 
(3.s.), I., 121 (1858; 3} Wkr., (Cat. pedins.abi Ms 
XXIX., 627 (1864). 

Madeira,! Southern Deserta ;! Madeira, four specimens 
(Baker). 


Blastobasis (?) awrantiaca, W|stn. 


Gracilaria (?) aurantiaca, Wlstn., Ann. and Mag. 
N. -H. (3's.), 1, 122.(1858);) Wkr Cat wip. ime: 
B. M., XXX., 854 (1864). 


Madeira, summer of 1855.1 


Epristerus, Wlsm., gen. nov. 
(emvatnTos, capable of being scientifically known.) 


Tyres, Hpistetus divisus, Wlsm. 


Antenne, g@, strongly bifasciculate (35); basal joint enlarged, 
with a pecten, second and third joints also enlarged, the base of the 
fourth joint attenuate. Mawillary palpi overlapping the base of 
the haustellum. Labial palpi with the second joint somewhat 
longer than the apical, scarcely tufted at the end beneath. Ocelli 
absent. Head somewhat crested above, face smooth. Thorax 
smooth. Forewings narrow, elongate, pointed, the apex scarcely 
depressed, anal angle obsolete. Neuration, 12 veins, 7 and 8 from 
a common stem, both to above apex. Hindwings as broad as the 
forewings, lanceolate, subovate, abdominal angle somewhat pro- 
nounced, cilia (1). Newration 7 veins, 3 and 4 coincident, 5 from the 
same stem as 3+4, 6 and 7 separate. Abdomen, ¢, somewhat 
flattened and laterally dilated. Legs with hind tibie slightly 
clothed, 

Epistetus divisus, Wlsm., sp. n. 


Antenne fuscous, basal joint brownish. Palpi brownish. Head 
and Thorax brouzy-brown ; face shining, pale cinereous. Forewings 
shining, bronzy-brown, with a slightly-waved narrow whitish line 
or fascia across their middle, scarcely farther from the base on the 
dorsal than on the costal margin ; beyond this line the outer half 
of the wing is slightly paler than the base, and shows a faint 
indication of a transverse darker spot at the end of the cell, 


Vortricidx, and Tineidx of the Madeira Islands. 553 


sometimes connected with a few scales of the same colour on the 
costal and dorsal margins, and giving the appearance of a dentate 
shade or fascia ; a few darkened spots around the apex are followed 
by paler scales at the base of the cilia which are bronzy-grey. 
Hindwings and cilia bronzy-grey. Abdomen bronzy-grey. Legs 
bronzy-brown, the spurs pale cinereous, tarsal joints spotted with 
pale cinereous. xp. al., 12-17 mm. 


Type. &2%. Mus. Wlsm. 
Hab. Madeira; five specimens. 


Hpistetus (?) pica, Wlsm., sp. n. 


Antenne, 9, with a pecten on the basal joint ; brownish-grey, 
basal joint white. Palpi white, shaded externally on the second 
joint with brown. Head and face white. Thorax white, a 
brownish-fuscous band across its anterior margin and a spot of 
the same colour posteriorly, tegule white. Forewings narrow, 
elongate ; shining white, with numerous brownish-fuscous patches 
and dots ; an elongate brownish-fuscous patch at the base of the 
costal margin reaches to the fold on its inner half ; this is followed 
before the middle of the wing by a similar costal patch tending 
obliquely outwards and narrowly confluent with a somewhat 
triangular dorsal patch of the same colour which reaches to the 
middle of the dorsal margin, thus forming an oblique fascia 
attenuated in the middle ; beyond the middle of the wing on the 
costal margin is a narrow elongate patch, beneath which are a 
few brownish-fuscous dots tending in the direction of a triangular 
dorsal patch beyond it a little before the anal angle; a con- 
spicuous costal patch lies before the apex and is followed by a 
series of small spots around the apex and apical margin to the 
anal angle ; an outwardly oblique narrow patch of the same colour 
lies near the base of the dorsal margin, reaching to the fold, and 
is continued narrowly alorg the margin to the base ; a single spot 
or dot lies on the lower edge of the discal cell above the outer end 
of the fold ; all these markings are more or less surrounded by pale 
ferruginous scales, but in some specimens these are scarcely 
apparent ; cilia grey, whitish towards the apex. Hindwings grey, 
with grey cilia. Addomen grey. Legs brownish, the joints of the 
tibize and tarsi indicated by whitish spots. zp. al.,15-17 mm. 

Type. ¢%. Mus. Wlsm. 

Hab. Madeira; two specimens. 


In the absence of the ¢ this species cannot be located 
with certainty. 


TRANS. ENT. SOC. LOND. 1894,—PART Iv. (DEC.) 2N 


554 Lord Walsingham’s catalogue of the Pterophoride, 


Hypatima, Hb. 
Hypatima fasciata, Stn. 


Gelechia fasciata, Stn., Ann. and Mag. N. H. (3 s.), 
III., 213. (1859) ;} Wkr., Cat. Lp. Ins. 6. M., 
XXIX., 628 (1864). ; 


Deserta Grande;! Madeira, Funchal, and The Mount, 
five specimens (Baker). 


LAVERNINA. 
Laverna, Crt. 


Laverna vittata, W1stn. 


Laverna vittata, Wistn., Ann. and Mag. N. H. (8 s.), 
L, 122 (1858); Wkr., Cat. Lip. Ins! B. M., XXX, 
885 (1864). 


Madeira.! 


Laverna rebeli, W1sm., sp. n. 


Antenne as long as the forewings; pale fawn. Palpi with 
second joint much longer than the apical, closely clothed and tufted 
beneath ; fawn-whitish on their inner sides, externally clouded 
with umber-brown. Head and face stone-whitish. Thorax fawn- 
whitish. Forewings fawn-whitish, slightly tinted by shining steel- 
grey scales on the paler portions and much clouded on the outer 
two-thirds with umber-brown ; a patch of raised dark umber- 
brown scales lies on the middle of the fold and extends obliquely 
outwards above it, forming the inner margin of a wide umber- 
brown dorsal patch; this patch and the other umber-brown 
shadings beyond it are sprinkled with scattered scales of a darker 
shade which are especially noticeable around the apex of the wing 
at the base of the cilia ; there is also a small umber-brown shade 
at the extreme base of the costal margin ; cilia pale fawn. Hind- 
wings fawn-grey, with paler cilia. Legs fawn-whitish, hind tarsal 
joints shaded with brownish and spotted with whitish externally. 
Abdomen missing. xp. al., 14-16 mm. 


Type. $2. Mus. Wlsm. 
Hab. Madeira; three specimens. 


Caautiopvs, Tr. 


Chauliodus daucellus, Peyr. 
Madeira; five specimens. 


Tortricide, and Tineide of the Madeira Islands. 555 


Asycuna, Stn. 


Asychna insularis, W|stn. 


Asychna insularis, Wlstn., Ann. and Mage NN. H. 
(3 s.), I., 122 (1858) ;! Wkr., Cat. Lp. Ins. B. M., 
XXX., 887 (1864). 


Madeira.! 
COSMOPTERYGIN. 
Cosmopreryx, Hb. 


Cosmopteryx pulcherrimella, Chamb. 
Madeira; nine specimens. 
Hitherto only recorded from the United States. 


GRACILARIANZA. 
Gracitaria, Hw. 


Gracilaria staintoni, Wlstn. 


Gracilaria staintoni, Wlstn., Ann. and Mag. N. H. 
(3 s.), I., 121, (1858) ;! Wkr., Cat. Lip; Ens: 
B. M., XXX., 854 (1864). 


Madeira ;' The Mount, two specimens (Baker). 
Gracilaria roscipennella, Hb. 
Madeira ; San Antonio da Serra, seventeen specimens, 
Gracilaria scalariella, Z. 
Madeira; twenty-one specimens. 
Litxocotietis, Hb. 


Inthocolletis messaniella, Z. 
Madeira; nine specimens. 
LYONETIANZ. 
PHYLLOBROSTIS, Stgr. 


Phyllobrostis daphneella, Stgr. 
Madeira ; one specimen. 


pa Hay oe 
igs Bry eaage 

a) ty 4i8 ree ey nae: 
sil) et aren sf uithdotgn VSR sre “ 


Tek he AN biel ng forty 
Joie teat a nate ark 8 if 
Ret y hie ‘vg Fi Page wae” 
Wire ifs thigh r wile whe: he dior . 
reyarhee. Meena ty 4a mr Pernice tay’ A: ae 


my (eaciearet) oad ant 
ft shone jamal kad mf deo wate OY 


a ental it Aaa 7 Ae ae 
Te Neate 265 eh Hogan’ Ljentvay ce 
RiGar a fom i ie ee fointh ahi OF 
SWS ecto ry ls dine dacietplyy, bo uieee ‘Y i 

Wea. 1 ha abl une (fly Manet tystaley lk 


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tip sgt TMM 5 Qh oleae 

« T rept Ot ive ic ee. 

(ee alka shes.) ‘qbilreagor ce SS 8 


voor cnet (Ceipmieene: ie Ad ebhote fe 
if) bbs a ed |! : ae Y at dh vida ap ae 
TYE a sikh ‘ie ng stay (one oder’? ret 


eal ed 


at fin vy RNY iy tot, Cts a(tanteiy 
f : i vl AY va De see Si ‘tosh eta) 


aiid 


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Ay, ad 
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dehy Aan Ma) 
d | i 
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(oan ay 


XXIII. Palearctic Nemoure. By Kenneta J. Mor- 
Ton, F.E.S. 


[Read Oct. 3rd, 1894.] 
Puates XIII. and XIV. 


At the present time the Perlidz, and especially the 
smaller members of the family, occupy a prominent place 
amongst the less-known of European insects. Pictet’s 
treatise (“Histoire Naturelle des Insectes Névroptéres, 
Famille des Perlides, 1841”), now over fifty years old, 
may still be considered the standard of our knowledge 
of the family, and that work, in many respects of 
exceptional merit, naturally falls short of modern re- 
quirements as far as the separation of critical and closely 
allied species is concerned. 

I have now collected these insects for nearly three 
years, and have, at the same time, made them the sub- 
ject of more or less constant study during my limited 
leisure. The district in which I happen to reside has an 
exceptionally good representation of that aquatic insect 
hfe which belongs to running waters, and materials in 
Perlidx, including most of the British species, were thus 
easily obtained. ‘lhe results of an examination of these, 
in a fresh as well as a dry condition, have led me to the 
conclusion that the only reliable criterion for the separa- 
tion of the species is in most. cases to be found in the 
genitalia and accessory appendages of the ¢. ‘These 
parts have been too much neglected by previous 
writers on Perlidx, and in support of the conclusions I 
have come to regarding their value, I now offer the 
following notes on the species of the genus Nemoura, in 
which genus the appendages appear to be most highly 
specialized. These notes must be regarded as in all 
respects preliminary, but relating as they do to a 
neglected subject, it is hoped they may be of use. 
Monographic completeness is not yet possible, and has 
not been attempted. 

At the outset 1t was not my intention to do more than 
try to make known the species inhabiting the British 
Isles. ‘The expansion of the original idea is due to the 

TRANS. ENT. Soc. LOND. 1894,—PpaRT Iv. (DEC.) 


558 Mr. Kenneth J. Morton on 


co-operation of the following entomologists :—Mr. 
McLachlan kindly lent me all his Palearctic Nemoure, 
numbering several hundreds, from all parts of the 
region, and including the results of his own collecting in 
Savoy, Switzerland, Burgundy, Dauphiné, the Pyrénées 
Orientales, Belgium, etc., Haton’s in Portugal, Zeller’s 
in Carinthia and elsewhere, and Fedchenko’s in Turke- 
stan; Dr. Fr. Ris, now of Rheinan, had the goodness 
to forward his whole collection of Swiss Nemoure, 
an extensive series in perfect condition; Mr. King, of 
Glasgow, supported me with much additional British 
material; Professor Klapalek, of Prague, sent me many 
useful insects; and Dr. John Sahlberg forwarded a 
choice collection of boreal and arctic Perlide, containing 
between three and four hundred examples belonging to 
the Helsingfors Musenm. I have thus had abundant 
material to work from; the apparently inadequate results 
are due to the small proportion of males and the fact 
that isolated females are in many cases practically 
indeterminable. 

At an early stage of my studies the nomenclature was 
seen to be in a hopeless tangle. No clear conception 
existed with regard to the limits of the species; the 
same species was known by several names according to 
the interpretation put upon existing descriptions by 
different entomologists; examples of the same species 
might be seen doing duty as exponents of two or more 
species in the same collection, or the reverse condition 
might obtain, two good species standing under one name. 
J am not sure that I have done any satisfactory work in 
clearing up the synonymy. Through the courtesy of 
M. Frey-Gessner, I have had the opportunity of ex- 
amining the Nemourx of the Geneva Museum, which 
contains what remains of the “ types” of the two Pictets. 
It appears, however, that the collection now contains 
very few examples which can properly be called “‘ types,” 
as the old collection of the elder Pictet was greatly 
destroyed by Anthrenus. Mons. A. Edouard Pictet con- 
tinued the collection, and added to it, and the whole, 
after his death, came into possession of the Geneva 
Museum. A few years ago, Mr. Albarda determined the 
Perlide ; and the Nemoure, as I understand, now repre- 
sent little more than the views of that author. As is 
usual in collections of these insects, females predominate, 


Palzarctic Nemoure. 559 


and it is not easy to speak of right and wrong with 
regard to them; but as there is certainly error in con- 
nection with the placing of some of the males, I have 
been driven, in a great degree, to adopt my own views 
about Pictet’s species. There may be differences of 
opinion regarding my conclusions, but I trust the notes 
and figures given will make easy the identification of the 
species to which I have applied Pictet’s names, and will 
also, in some degree at least, prevent a recurrence of the 
confusion alluded to at the beginning of this paragraph. 

Stephens’ types are much more satisfactory than those 
of Pictet. They are, for the most part, in good condi- 
tion and serviceable ; the drawback in connection with 
them is the one common to collections of Nemowre 
already alluded to, namely, the undue proportion of 
females. My conclusions regarding these types will be 
referred to in the synonymic references preceding, or the 
notes following, the descriptions of the species. I have 
to express my indebtedness to Mr. Chas. O. Waterhouse, 
of the British Museum, for enabling me to examine 
Stephens’ Perlide. 

Before giving the descriptive and other notes on the 
species, it may be well to say a few words regarding the 
characters of Nemoura. The geuus was established by 
Latreille in 1796. Pictet adopted the term to cover his 
three sub-genera, Txniopteryx, Leuctra, and Nemoura. 
These three divisions are now generally regarded as 
separate genera, and the species to be dealt with here 
are those falling under Nemowra proper. 

The following are Pictet’s diagnoses of the three 
genera (very slightly modified) :— 


TENIOPTERYX. 


(Nervures of pterostigma not forming a x.) 


Labial palpi moderate and separated. Abdomen terminated by 
short sets. Three tarsal joints long and nearly equal, Wings in 
rest half cylindrical. 


LructTra. 


(Nervures of pterostigma not forming a y.) 


No caudal setee. Labial palpi very short. Tarsi with 2nd joint 
short, 1st and 3rd long. Wings in rest half cylindrical. 


560 Mr. Kenneth J. Morton on 


NEMOURA. 


(Nervures of pterostigma forming a y.) 


No caudal sete. Labial palpi short and approximate. Tarsi 
with 2nd joint short, Ist and 3rd long. Wings in rest ordinarily 
flat. 


Albarda has shown that the above-mentioned character 
in the neuration of Tniopteryx is not absolutely charac- 
teristic, as his 7’. neglecta* has the wings in the normal 
condition of a Nemoura. ‘The condition of the tarsi will 
at once prevent any confusion between these two genera, 
and the wing-neuration character seems to hold guod as 
a means of separating Nemoura and Lewuctra, which 
agree with each other as regards the tarsi. 

With regard to the position of the wings in rest, 
there appear to be in Nemowra slight deviations in the 
direction of the position they occupy in Leuctra ; this 
has been noticed in N. Meyert when alive. Other 
characters made use of in the above diagnoses will also 
require further investigation ; for example, the presence 
or absence of caudal sete. Seta probably exist in at 
least one of the species here described as a Nemoura. 
Then the appendages of various species here retained 
under Nemoura show so much diversity of structure that 
it is almost certain that, sooner or later, further generic 
subdivision will take place based on these parts. 

The figures are from drawings made under the micro- 
scope with the camera lucida. In most cases, two views 
are given—one from the side, the other from beneath— 
the object beimg held slightly in front of the observer. 
These figures and the descriptions were all made from dry 
examples for the sake of uniformity. These insects are 
subject to great variations in size and colour, according 
to locality, and they have almost all a tendency to 
become blackish or unicolorous when dry, and colour 
characters are, therefore, not of great importance, except 
in a broad way. 

Another feature in these insects (and in all Perlidz) 
is the instability of neuration, and few details under this 
head are sufficiently constant to be of help in the 
separation of species. It has been the practice to lay 


* “ Annales de la Soc. Ent. de Belgique,” Tome xxxiil. 


Palearctic Nemowre. 561 


stress on the form of the pronotum, and this appears to 
be justified to some extent ; but care must also be taken 
here, as shrinkage in drying is unequal, varying with the 
degrees of maturity of the individuals. Amongst living 
examples of the same species of one of the larger Perlidzx, L 
believe that I have noticed a difference between the form 
of the pronotal plate in examples just disclosed and 
those in which the chitin had set hard. While the rugose 
condition of the surface of this segment and the arrange- 
ment of the tubercles may be useful for specific deter- 
mination, I have not yet been able to come to satisfactory 
conclusions on the point. 

In the following descriptions, when reference is made 
to the wings, the anterior pair is alluded to, unless 
otherwise stated. 


Nemowra variegata. 


N. variegata, Olivier, Encye. Méth., vili., p. 186, 
No. 3 (1811) ? 

N. variegata, Pictet, Perlides, p. 386, pl. 1. (L@41). 

N. nebulosa, fuliginosa, pallida, cruciata, affinis, annu- 
lata, luteicornis, pusilla, Stephens, Lllustrations, 
p-. 140 and 141 (1836). 

N. lunata, Rambur, Névropteéres, p. 461 (1842). 

N. uwmbrosa, Kd. Pictet, Névropt. d’Hspagne, p. 20 
(1€65), in part according to types. 


Head and pronotum yellowish-brown, the latter with darker 
clouds. Antenne usually brownish, with paler basal joint. Pro- 
notum very slightly broader than the head without the eyes, with 
a broad, well-defined border, whose margins are gently rounded 
both in front and laterally ; this segment is minutely granulose 
with two vague transverse depressions (or three, counting the one 
behind the anterior border). Meso- and meta-nota dark brown or 
nearly black, the scutella of these segments and the parts adjoining 
sometimes paler. Abdomen brown or blackish, Legs varying from 
brown to yellowish, femora rather darker about the apex, tibiz 
with a dark apical annulation, and tarsi, especially the distal joints, 
fuscous. Wings ranging from nearly hyaline to brownish-grey, 
with blackish or fuscous neuration, often with fuscous markings in 
the region of the pterostigma and x nervures, the pigment in 
strongly-marked examples spreading out on either side of the 
nervure forming the base of the 3rd apical cellule into a fuscous 
triangle, . In ibis species the end of the radius is almost invariably 


562 Mr. Kenneth J. Morton on 


considerably turned up just where it joins the outer margin ; the 
4th apical cellule is biangulate at the base ; and in the hindwings 
there are several transverse nervures in the intercubital areas, a 
state of things not normal in other species. In the ¢ the median 
prolongation of the ventral plate and the lobe on this plate are 
both moderate ; the lateral appendages are slender, slightly curved, 
and narrowing towards the apex, which terminates in two hooks, 
the lower of which is most evident ; seen from above or beneath, 
these appendages are straight, the apex slightly out-turned with 
the tip of the hook turned inwards. In the 9 the apex of the 
abdomen has two large ovate lobes and a pair of longer simple 
appendages. Exp., g, 12} to 20mm.; 92, 15 to 24 mm. 


This species is common in Britain, and probably over 
the whole Palearctic region. I have seen it from Arctic 
Norway, Finland, the Vosges, Schwarzwald, Switzerland, 
Bohemia, Portugal, and Sarepta; it is also mentioned by 
McLachlan in ‘‘ Nedchenko’s Travels,”? as seen by him 
from Turkestan. ‘To what altitude it goes is uncertain; ~ 
very small examples are in McLachlan’s collection from 
the Col du Lauteret, Alps of Dauphiné. It seems 
attached to standing waters, and even occurs in places 
where there is little or no surface-water in the dry 
seascn. ‘Ihe time of appearance varies with locality 
from April to August, and probably later. 

At once one of the largest (but varying greatly in this 
respect, some minute examples occasionally appearing) 
and best-defined species of the whole genus. The granu- 
lose prothorax, the condition of the neuration, and the 
appendages of the ¢ are all characters which can be 
relied upon. For this reason it is the one Nemoura 
regarding whose name there is something like unanimity 
of opinion at the present time. Rambur’s description 
of lunata is good; he refers to the appendages of the ¢. 
The eight species of Stephens’ above narrated are cer- 
tainly all to be referred bere according to the types in 
British Museum ; pusilla is a small ¢. 


Nemoura avicularis, n. sp. 


Head and pronotum shining black, the latter sometimes very 
narrowly outlined in yellow or orange. Meso- and meta-nota also 
shining black with a brown posterior marking. Pronotum very 
slightly broader than the head without the eyes, foremargin and 
sides hardly rounded, borders not distinct in the dry insect, disc 


Palearctic Nemoure. 563 


faintly rugose. Abdomen blackish. Legs: femora above, ends: of 
tibiz, and the tarsi all fuscous. Wings greyish in some lights, but 
almost hyaline, highly iridescent, only very faintly clouded in the 
pterostigmatic region and about the x nervures; neuration 
generally blackish and distinct; the 4th apical cellule rarely 
biangulate at the base; in the posterior wings the superior 
cubitus usually leaves the radius at a considerable distance from 
the basal cell. In the g¢ the median prolongation of the ventral 
plate is comparatively short; the lateral appendages are broad at 
the base, contracted in the middle, with a large head, upturned, 
and acute at the apex externally, and possessing an eye-like 
tubercle ; the distal part of the appendage being very suggestive 
of a bird’s head. Exp., ¢, 174-18 mm.; 9, 22 mm. 


A species easily recognized by the appendages of the 
8, the very black coloration of the body and the highly 
iridescent nearly hyaline wings, which have hardly any 
darker cloudings. With regard to the lateral appen- 
dages, while there appears to be a slight amount of 
variation due to changes in drying, they ordinarily retain 
their character. The chance of confusion most exists 
with regard to the next species, but the notes and figures 
will, I hope, prevent error so far as the ¢ is concerned. 

According to our present British information, this 
species seems to be exclusively lacustrine in habit, and 
has been found in April and May at many of the lochs 
in the highlands of Scotland (King and Morton). It is 
a!lsoin Mr. McLachlan’s collection from the Hnglish Lake 
District. Outside Britain it has been seen from Finland ; 
the collection of the Helsingfors Museum contains 
examples from several localities. 

This species has apparently already been recognized 
in this country, but associated hitherto with one of 
Pictet’s names, a view I cannot in the meantime accept. 
The insect is as yet only kuown as an inhabitant of 
Northern Europe. 


Nemoura cambrica. 


N. cambrica, Stephens, Illustrations, p. 143 (?). 
N. pallicornis and nitida, ibid., p. 148 (7). 


Head and thoracic segments shining blackish. Antenne 
brownish or black. Pronotum nearly as broad as head, with pale 
lateral margins which recede slightly posteriorly. Legs usually 
yellowish, darker on the femora above and on the tarsi, Wings 


564 Mr. Kenneth J. Morton on 


brownish-grey, with darker pterostigma and fuscous or blackish 
neuration. The lateral appendages of the ¢, viewed from the 
side, are rather broad and rounded at the base, the shaft slender 
and subcylindrical, bent beyond the middle and continued out- 
wards into a subacute point ; viewed from beneath, the apical part 
of the appendage is turned outwards in the form of a thick hook ; 
inwardly, the chitinized part is prolonged into a spur, under which 
is a membranous space. Exp., 12 to 18 mm., the ? the larger. 


The smaller size and less hyaline wings will ordinarily 
suffice to distinguish this species from the last; the form 
of the appendages is constant in all the material I have 
examined, and I have never noticed any tendency in 
them to assume the upturned position they occupy in the 
last; in the present species they are also much more 
slender. 

This is one of the commonest Nemoure of all the 
streams of North Britain wherever I have collected. It 
kas occurred by all sizes of running waters, from the 
larger rivers to the tiniest rills, the size of the insect 
varying according to the extent of the waters it inhabits. 
It has not yet come before me from the Continent. 

The application of the name cambrica is open to 
question. The only type seen by me is a ?, and, there- 
fore doubtful. N. pallicornis and N. nitida are both 
represented by females without abdomen ; they are more 
probably large examples of the species which I have 
named N. inconspicua. 


Nemoura lateralis. 


N. lateralis, Pictet, Mém. de la Soc. de Phys. et 
d’Hist. Nat., vu., p. 180 (1836); and Perlides, 
p. 895 (1841); ¢ (2). 

N. nitida, Pictet, Mém. Soc. Phys., p. 179; and 
Perlides, p. 392; 9 (7). 

Head and thoracic segments brownish, or black mixed with 
brown, margins of pronotum paler. Antennz concolorous with 
head, basal joint often paler. Pronotum about same breadth as 
head without the eyes, with margins moderately distinct, receding 
very little posteriorly. Legs yellowish, femora above marked with 
fuscous, as are also the tips of the tibie, and the tarsi. Wings in 
the ¢ usually nearly hyaline, in the ¢? rather more greyish, 
neuration fuscous, membrane faintly clouded about the x nervures 
and pterostigma, The appendages of the ¢, from whatever aspect 


Palearctic Nemoure. 565 


viewed, do not show the distinct hook-like form of the two pre- 
ceding species ; seen from above, their chitinous part is somewhat 
bifid, the furcation filled out with membrane ; viewed from the side, 
their base is broad and rounded, from which they taper in a nearly 
straight direction to the apex, where they again have a broader 
aspect, the upper and more chitinous part of the apex being 
slightly turned outwards, and the lower membranous part produced 
downwards ; from beneath, the appendages are straighter than in 
the two foregoing species, the apex being scarcely curved, but 
turned out at nearly right angles, and terminating in a sub-acute 
point. Exp., ¢,13-17mm; ¢?, 183-223 mm. 


Apparently a very common Swiss insect ; received in 
great quantity from Dr. Ris, taken on the Ziirichberg in 
April and May, and varying much in size and colour. 
What I consider the same insect is present in great 
numbers and variety in Mr. McLachlan’s collection from 
various Swiss localities, including the Hngadine, Hash- 
thal, etc. (McL.); also from Savoy, Chamounix (MclL.), 
Schwarzwald, the Vosges (Mch.), and Carinthia (Zeller). 
1 have also received it from Bohemia (Klapalek). 

An insect easy to distinguish in the $ sex; the ? 
might easily be confused with some of its alles, especially 
N. avicularis. The prothorax in the latter is, perhaps, 
relatively broader, the borders less definite, and the 
margins more rounded; but in the dry insects these 
characters could hardly be relied on with confidence. 
Some of the large ?s from the Alps of Switzerland and 
Savoy look very different from the normal ? ; they have 
an almost square pronotum, are greyer than usual, and 
have the veins more clouded. ‘These deserve attention. 

This species seems to agree pretty well with Pictet’s 
description of N. lateralis. I accept the opinion now 
generally adopted, that Pictet’s N. nitida is the ? of the 
same species. 


Nemoura dubitans, n. sp. 


Dark brown or black, shining, antennze dark brown. Pronotum 
broad, with distinct, usually pale, somewhat rounded margins. 
Legs brownish, femora darker above, as are also the apices of the 
tibiee and the two distal joints of the tarsi. Wings transparent, 
of uniformly brown tint with darker pterostigma ; neuration not 
very strong. The ¢ is easily distinguished from all others nearly 
allied by the genital parts. The ventral plate has the lobe thereon 


566 Mr. Kenneth J. Morton on 


long and narrow; the lateral appendages of about the ordinary 
length, tapering slightly to the apex, which is obtuse and appa- 
rently simple. Exp., ¢,15 mm.; ¢, 20 mm. 


The brownish aspect of this fly is a good prima facie 
character for its determination; the weak neuration and 
the form of the lateral appendages are also of impor- 
tance. All the examples are from Dr. Ris, and are 
labelled “ Oerlikon, April.’ A species requiring further 
elucidation, but I think it is a good one. Confusion 
especially in the @ sex is likely to take place with the 
species 1 here name N. marginata, but the characters of 
the ¢ appear to be decisive. 


Nemoura prxcow, n. sp. 


Phryganea_ regelationis, Scop., Ent. Carniolica, 
ps 209 (ce 


Head and thorax black, antennze concolorous; pronotum 
broader than the head without the eyes, short, borders not well 
defined in the dry insect, lateral margins usually with a slight 
angulation about their middle part, behind which they recede 
somewhat. Legs: femora above, ends of tibiz and the tarsi 
blackish fuscous. Wings greyish, sometimes fuliginous, neuration 
blackish fuscous ; all the veins margined more or less with darker 
cloudings, which are most distinct about the pterostigma and 
x veins. The anal structure of the ¢ is complicated: the lobe of 
the ventral plate is of great size. What appears to correspond 
with the long spinous sheath of N. A/eyeri consists here of a broad 
basal part with a very short spine. Exp, ¢, 18 mm.; 
©, 20-23 mm. 


The complicated anal structure of the ¢ and the some- 
what fuligimous aspect of the wings should make this 
insect recognisable. The form of the prothorax above 
alluded to is constant in the British examples which I 
have examined. 

Apparently not rare in Great Britain. It occurs in 
early March (if not earlier in mild seasons) by the River 
Clyde and one of its affluents, along with Teniopterye 
nebulosa and 1’. trifasciata. I have also seen both sexes 
from Perthshire rivers in April (King), and two ?s are 
in Mr. McLachlan’s collection from the Manchester 
district in March (Cooke). 


Palearctic Nemours. 567 


That this is really Scopoli’s insect is very doubtful. 
Still the name is singularly appropriate, and I would 
have adopted it in preference to creating a new one, if 
such a course had been at all prudent. 


Nemoura maracandica. 


Teniopteryx maracandica, McLachlan, Fedchenko’s 
Travels in Turkestan, Neuroptera, p. 54. 


Black, with greyish pubescence, somewhat shining. Head con- 
cave in middle of disc. Pronotum almost quadrate, sides straight, 
nearly parallel, anterior angles sub-acute, disc faintly rugose. Feet 
sub-testaceous, with greyish pubescence, femora fuscous externally, 
tibize fuscous at base, tarsi black. Abdomen black ; ventral lamina 
of $ triangular, truncate, beneath somewhat concave ; appendages 
testaceous. Ventral lamina of ¢ large, adpressed, sub-testaceous, 
with rounded margin, Anterior wings elongate, greyish, faintly 
irrorated with darker grey dots, neuration blackish fuscous. Exp., 
al., 18-24 mm. 


Found near Samarcand and Taschkent. I have seen 
the types, but the above description is mainly from 
Mr. McLachlan’s diagnosis. he insect is a large one, 
with the facies of a Tseniopteryx, and is very distinct 
on account of its dotted wings. 


Nemoura ornata. 


N. ornata, Mclachlan, Fedchenko’s Travels in Turke- 
stan, Neuroptera, p. 55. 


Blackish with greyish pubescence, shining ; head concave in 
middle of disc. Pronotum transversely sub-quadrate, sides straight, 
parallel, disc almost smooth. Feet black, with greyish pubescence, 
femora with apex testaceous. Ventral lamina of 9 large, ad- 
pressed, entire, transversely striate. Anterior wings short, broad, 
vitreous, with blackish neuration ; two irregular transverse fascie, 
margins of the nervures and some small clouds all smoky. Inferior 
branch of sector much curved. Exp., al., 17-175 mm. (¢). 


This species is from the mountains of Khokan. I am 
indebted to Mr. McLachlan for a sight of the types, and 
the above is mainly a translation of his diagnosis. A 
pretty little species marked somewhat after the manner 
of N. Meyeri, but the markings are much more definite ; 
the femora are also nearly altogether dark externally. 


568 Mr. Kenneth J. Morton on 


Nemoura Meyert. 
N. Meyeri, Pictet, Perlides, p. 390, pl. h. 


Head and thoracic segments black, antenne blackish, hind-head 
and margins of pronotum usually yellowish, the latter segment 
about same breadth as head with very slightly rounded margins 
(the material examined shows great variation in the form of the 
pronotum). Legs yellowish, middle and apex of femora, apex of 
tibiz and tarsi blackish. Wings greyish, often whitish, most of 
the nervures broadly bordered with brownish grey, or the colour 
might be given as brownish grey with white spots ; pterostigma 
pale in the middle and dark on either side; neuration blackish, 
extreme end of radius upturned, lower branches of sector and 
superior cubitus usually much curved. The most characteristic 
feature of the ¢ genitalia is a pair of long upturned spine-like 
sheaths. The large upturned pieces behind these sheaths appear 
to be flanked by blackish pectinated pieces which are often con- 
cealed. Exp. ¢, 153-183 mm.; 9, 183-24 mm. 


In what may be termed typically marked examples, 
this species may be easily recognised by the two-coloured 
pterostigma, the curved neuration above alluded to, and 
the marking on the middle of the femora is also of im- 
portance. With regard to the genitalia of the 3, confusion 
is possible with N. marginata; in the latter species, 
however, the ‘‘ upturned spine-like sheaths” are not 
usually so straight and they end in two points, and the 
pectinated pieces are much more conspicuous and have 
much longer teeth than in Meyert. Well-marked ex- 
amples of the species are usually correctly named. 

I can say little regarding the geographical distribution 
of this species. It is rather common at many streams in 
North Britain in May and June, and even later in 
localities of some altitude. It is also represented 
by large examples from Carinthia in the collection 
of Mr. McLachlan and from Switzerland in that of 
Dr. Ris. 

Under the name of WN. nitida, Pictet, Professor 
Klapalek has communicated a pair of Bohemian insects, 
the ¢ immature and the ? closely resembling N. Meyerv. 
The paired spines are not unlike those of the species 
hereafter described as N. marginata; they appear to be 
shorter and broader before the apex than in N. Meyeri, 
and terminate in two more or less divergent spines. ‘This 


Palzxarctic Nemoure. 569 


may bea good species but more material is required. On 
account of the immaturity of the ¢ the figures are not 
satisfactory. 


Nemoura marginata. 


N. marginata, Pictet, Mém. Soc. Phys., tom. vii., 
p. 181, No. 7; and Perlides, p. 397 (?). 

N. humeralis, Pictet, Mém. Soc. Phys., tom. vii., 
p. 184, No. 11; and Perlides, p. 399 (°?). 

Head and pronotum shining blackish, mixed with brown especi- 
ally on the hind-head. Pronotum slightly narrower than head ; 
borders not well defined, usually paler, the lateral margins recede 
very slightly posteriorly. Legs pale brown. Wings usually 
brownish, sometimes suffused, in other examples nearly hyaline, 
neuration darker especially in the region of the pterostigma and 
x nervures. The structure of the genitalia is complicated ; the 
paired sheaths are in this species much as in N. J/eyeri, but instead 
of ending in a simple acute point the apex is crowned by two short 
spines, and there appears to be another spine at some distance from 
the apex. The strong upright pieces are flanked by a chitinous 
sheath which is deeply pectinate, although the pectinations may 
sometimes be so closely appressed that they are difficult to see. 
Exp., 18-22 mm. 

In applying to this species the name of marginata, 
Pictet, I have been guided chiefly by the fig. of that 
author which agrees tolerably well with a common 
brownish Swiss insect. The limits of the species are, 
however, difficult to define, and there is a certain con- 
fusion regarding it and N. humeralis, a species not yet 
elucidated to my satisfaction. Professor Klapalek sent 
me a ¢ agreeing very well with Pictet’s description of 
N. humeralis, and also forwarded drawings of what he 
considered the ¢, but in the great number of examples 
examined by me I have not yet seen a ¢ which agreed 
with Klapalek’s ¢, and I am inclined to consider the 
latter as a condition of the species under review. In the 
rich materials forwarded by Dr, Ris, I found three forms 
which I have in the meantime referred here; the first 
series is from Glarus, in October, headed by a ¢, from 
which figs. 1 and 2 are taken; the females of this series 
have the wings comparatively transparent with rather 
strong neuration. ‘The second series mainly from the 
Ziirichberg in June have the wings more suffused and 


TRANS. ENT, SOC. LOND. 1894.—paRtiv. (DEC.) 20 


570 Mr. Kenneth J. Morton on 


therefore more typical; unfortunately, it contains no 
males. The third contains larger ¢ examples also from 
Ziirichberg in May ; the wings in these are brownish and 
transparent ; these are most doubtful. 

With regard to distribution, in addition to Switzerland 
and Bohemia, insects which are temporarily referred to 
this species are in Mr. MclLachlan’s collection from 
Carinthia and the Schwarzwald. 


Nemoura lacustris. 


N. lacustris, Hd. Pictet, Névropt. d’Hspagne, 
p. 2h ir). 

Head and pronotum blackish or piceous, antennz concolorous, 
occiput and margins of pronotum sometimes paler. Pronotum 
nearly as broad as head without the eyes, sub-quadrate, lateral 
margins scarcely receding. Legs dingy testaceous, darker at apices 
of joints. Wings greyish, sometimes with a faintly brownish tint, 
subhyaline, iridescent, x nervures faintly clouded, clouding of 
pterostigma often leaving a clear costal space almost as in Meyer ; 
radius slightly upturned at apex. In the ¢ the genitalia are very 
similar to those of marginata, but in none of the examples ex- 
amined are the pectinations of the sheaths flanking the upright 
pieces visible, and the upturned spines are apparently shorter and 
stouter. Exp. al., 19 mm. 


Closely allied to N. marginata, but apparently distinct 
on account of the much shorter and stouter upturned 
spines of the ¢. The examples from which the de- 
scriptive notes are taken are in Mr. Mclhachlan’s 
collection; they came from Portugal and_ the 
Pyrenees. 

There is doubt about the application of Ed. Pictet’s 
name. I have seen the types, and they bear resemblance 
to the insects from which the above description is drawn, 
but they now lack abdomens. 

The group which contains the three species described 
as Meyert, marginata, and lacustris is the least satis- 
factory of all here dealt with; the appendages are 
complicated in the dry insects, and difficult to under- 
stand ; confusion between degrees of exsertion and real 
differences of structure seems to be a danger to be 
specially guarded against. 


Palearctic Nemoure. 571 


Nemoura cinerea. 


N. cinerea, Olivier, Enc. Méth., viii., p. 186, No. 2 (?). 

N. cinerea, Pictet, Perlides, p. 401 (?). 

N. sulcicollis and N. fwmosa, Stephens, Illustra- 
tions, p. 143. 


Shining, black with concolorous antenne. Pronotum usually 
distinctly transverse, about as broad as head, lateral margins 
straight without definite borders. Legs brownish. Wings grey, 
nearly hyaline, but sometimes smoky, with fuscous neuration, 
which is nearly black and usually very strong about the x nervures 
and pterostigma. In the ¢ the median prolongation of the 
ventral plate is of unusual length, and the ventral lobe is long and 
narrow ; there is present a pair of very strong upturned usually 
blackish sheaths which appear to be obtuse at the apex. Exp., 
11-155 mm. 


This species I have found very common in June and 
July at all the Scotch streams where I have collected. 
I have received from Klapdlek both sexes, these 
Bohemian examples having the neuration less distinct 
than in most British specimens. Only one rather doubt- 
ful ¢ example was in Dr. Ris’ collection, but the species 
probably occurs commonly in some parts of Switzerland. 
Examples in Mr. McLachlan’s collection from several 
countries are referred by me to this species: France 
(Autun and the Vosges), Switzerland (Hngadine), and 
Schwarzwald ; probably, also, Portugal  (Haton), and 
Carinthia (Zeller). 

Professor Klapalek considers this species to be Pictet’s 
N. cinerea, and | am inclined to agree with him, although 
the view has been considered by some as doubtful. 
Whether it is Olivier’s cinerea is a point which may be 
questioned with more reason. The species is very 
variable, and I think Stephens’ two species above- 
mentioned are certainly forms of it. 


Nemoura borealis, n. sp. 


Head dark shining brown, antennz usually paler. Pronotum 
brownish, sub-quadrate, narrower than the head. Legs testaceous, 
almost without darker markings. Wings pale, subhyaline, with 
very pale neuration. In the ¢ the ventral lobe or lamina is rather 
narrow and elongate ; appendages rather broad at the base, tapering, 
twisted in the dry insect, expanding towards the somewhat flat- 


Dire Mr. Kenneth J. Morton on 


tened apex, the margins of which are deeply serrate. Exp., about 
15 mm.in g ; the 9 larger. 


So far as the material examined shows, this is a small, 
pale, rather delicate-looking insect, but it is absolutely 
devoid of any striking outward character, The appen- 
dages, however, are very unlike those of any other 
species. 

I have seen it from several Finnish localities, and I 
believe Mr. McLachlan has it from Lapland. 


Nemoura inconspicua. 


N. inconspicua, Pictet, Mém. Soc. Phys., tom. vii., 
p. 185, No. 12 (?). 

N. inconspicua, Pictet, Perlides, p. 404, pl. lin. (?). 

N. pallipes, Steph., Illustr., p. 142 (?). 

N. pallicornis, Steph., Illustr., p. 145 (?). 

N. xitida, Steph., Hlustr., p. 143 (?). 

Shining blackish or brown, prothorax as broad as head with pale 
broad borders and receding lateral margins. Legs yellowish, femora 
hardly marked, tibiae with dark tips, and apical joints of tarsi also 
darker. Wings pale, nearly hyaline, with faint light brown 
neuration. Inthe ¢ the genitala are very prominent, and consist 
principally of two large subcylindrical upper yellowish lobes, and 
a pair of very long slender upturned yellow sheaths. Lobe of 
ventral plate long. Exp., g 13-15} mm., 9 17-18 mm. 


One of the most easily determined of the Perlide, 
Its generally pale colours and the prominent’ genitalia 
in the d separate it from all its congeners. In Scot- 
land I have taken it sparingly in July and August; it is 
attached to smaller streams and to springs. It is well 
represented in the Swiss collection of Dr. Ris. Mr. 
McLachlan has taken it at Chamounix in July. 

This species agrees with Pictet’s description as far as 
that goes. I think, however, it is more than likely that 
Pictet’s species was really a combination of minute forms 
belonging to more than one species, Stephens’ species, 
above alluded to, are probably large females of this 
insect. 

The insects noticed in the preceding pages almost 
certainly represent but a small proportion of the species 
which exist in the Palzarctic region. In addition to 
many females which must remain doubtful, including two 
in Mr. Mclhachlan’s collection from Algeria (Haton), 


Palzxarctic Nemoure. oto 


interesting as being the only Nemowre seen by me from 
the African side of the Mediterranean, there is at least 
one other good species from Turkestan in the same 
collection, but hardly in sufficient material to warrant 
description. 

In commending these insects to the notice of col- 
lectors, I would urge upon them not to despise insigni- 
ficant-looking examples, as these are usually the males, 
and most useful for purposes of determination. If 
possible, all Perlidx should be expanded at once, as few 
insects are so ill-suited for the relaxing box. 


EXPLANATION OF Puates XIII and XIV. 


PLATE XIII. 


Nemoura variegata. 
. Apex of abdomen of ¢ from beneath. 
. Lateral appendage from side, much enlarged. 


no 


Nemoura avicularis. 


. Apex of abdomen of ¢ from beneath. 
35 x side. 


bo 


Nemoura cambrica. 


1. Apex of abdomen of ¢ from beneath. 
ahs 45 side. 


Nemoura lateralis. 
. Apex of abdomen of ¢ from beneath. 
. Lateral appendage from side, more enlarged. 
” ” above ” 


wOnNnre 


Nemoura dubitans. 


. Apex of abdomen of ¢ from beneath. 
si ‘4 side. 


Ne 


Nemoura precox. 


1, Apex of abdomen of ¢ from beneath. 
2. Pronotal plate. 


Nemoura maracandica. 
Apex of abdomen of ¢ from beneath. 


Nemoura meyeri. 


. Apex of abdomen of ¢ from beneath. 
*s as side. 


ror 


574 


Mr. K. J. Morton on Palzarctic Nemoure. 


1. Apex of abdomen of ¢ from side. 
2 
3. 


Nemoura marginata, 


S beneath, 
», pectinated piece, more enlarged. 


Nemoura cinerea. 


1. Apex of abdomen of ¢ from beneath. 
2. 


+5 , side, 


Nemoura inconspicua. 


1. Apex of abdomen of ¢ from beneath, 


2; an i" side, 
Nemoura -————— ? sp. from Bohemia (see page 568). 
1. Apex of abdomen of ¢ from beneath. 
2s ‘i es side, 
PLATE XIV. 


Nemoura variegata. 


1. Anterior wing. 


oo bo 


Q 


al Si tel fact 


Heo OW > 


. Upper branch 
. Lower branch 


Costa. 


. Sub-costa. 

. Radius. 

. Sector radii. 

. Superior cubitus (otherwise median nervure). 

. Inferior cubitus, upper branch (otherwise superior 


cubitus). 


. Inferior cubitus, lower branch (otherwise inferior 


cubitus). 


. Upper ein es fork of superior cubitus or 


Lower branch median nervure. 


of fork of sector radii. 


1, 2, 3, 4, 5, 6. Apical cellules. 


. Posterior wing. 
. Tarsus. 


All much enlarged. 


Nemoura borealis. 


Apex of abdomen of ¢ from beneath. 


Nemoura lacustris. 


. Apex of abdomen of ¢ from beneath. 
. “ Sheath ” from side. 


rs from beneath, more enlarged. 


(ah7er 9 


XXIV. Supplementary Notes on the Scolytidee of Japan, 
with a list of species. By Watrer F. H. Branp- 
FORD, M.A., F.Z.S. 


[Read October 3rd, 1894.] 


THE purport of these notes is to complete my previous 
papers on the Scolyto-platypini and on the Scolytide of 
Japan by the rectification of ene or two errors which 
have crept in, by the addition of three new species which 
have since come to hand, and by the publication, for 
convenient reference, of a list of Japanese Scolytide. 

In my paper on the Scolyto-platypint (Trans. Ent. 
Soc. 1893, p. 425) on p. 430, fourth line above the 
foot-note, for “ anterior”’ read “ posterior femora”; on 
p. 431, thirteenth line, a semicolon should follow the 
word “ remote.” 

In my paper on the Scolytide of Japan (Trans. Ent. 
Soc. 1594, p. 53) on p. 103, last line, for “ separate ”’ 
read ‘‘ asperate.” 

The following species should come after Xyleborus 
brevis, on p. 104 :— 


Xyleborus amputatus, sp. n. 


Fem. Cylindrica, subnitida, breviter pilosa, testacea ; pro- 
thorace semi-elliptico, margine antico convexo subcrenato, dorso 
medio transverse elevato, postice sat dense subtiliter punctato et 
in basi media hirto; elytris prothorace longioribus, parallelis, 
subtiliter lineato-punctatis, interstitiis confuse punctatis, apice 
oblique truncato, truncatura subcirculari, per totum acute mar- 
ginata, opaca, lineato-punctata, utrinque subconcava. Long. 
2°5 mm. 

Hab. Japan, Higo; one specimen (coll. Lewis). 

Fem. Cylindric, moderately shining, testaceous, with short 
pubescence. Head concealed in the type, antenne light testaceous, 
of normal structure. Prothorax semi-elliptic, widest near the 
middle, the sides rounded from the base, slightly behind, more 
strongly towards the convex apex, the margin of which is finely 

TRANS. ENT. SOC. LOND. 1894,.—ParRT IV. (DEC.) 


576 Mr. W. F. H. Blandford’s supplementary notes 


crenate, hind angles obtuse, base subtruncate ; surface with an 
indistinct median transverse elevation, thinly pubescent, its 
anterior half with fine somewhat scattered granular asperities, its 
posterior half rather closely and finely punctate, with a slightly 
depressed patch at base set with close upstanding pubescence and 
intersected by a subelevated median shining line. Scutellum tri- 
angular, rather large. Elytra nearly one-half longer than the 
prothorax, with oblique but scarcely rounded basal margins, the 
shoulders rectangular, the sides straight and subparallel with a very 
slight posterior divergence ; surface cylindric, obliquely truncate 
behind from the posterior third, with lines of very fine punctures, 
interstices with finer irregular piliferous punctures, truncate 
area subcircular, sharply margined all round, subconcave on 
each side and elevated along the suture, dull, glabrous, with three 
rows of punctures on either side, the interstices flat, irregularly 
punctured, its inferior apical border forming a very obtuse angle 
when seen from above. Underside and legs hght testaceous. 

Of the Xylebori known to me, this species is most like 
X. (Tomicus) truncatus, Hr., with which I identify a 
Tasmanian example before me. That species is larger, 
darker and without the patch of hair on the base of the 
prothorax, its elytra are longer, the apical truncate 
surface is nearly vertical, convex, shortly pilose, shining, 
with three impressed striz on either side, the interstices 
convex and muitipunctate. 


The following species should succeed X. validus on 
p: 108 :— 


Xyleborus interjectus, sp. n. 

Fem. Oblonga, sat nitida, nigro-picea; X. valido, Hichh. 
simillima, distinguenda elytris pro portione brevioribus, prothorace 
vix sesqui longioribus, a basi usque ad apicem convexioribus, 
lineato-punctatis, striis non impressis, interstitiis planis, pilis 
erectis longioribus e punctis subasperatis egredientibus per totum 
confertius seriatis, apice minus deplanato, punctis striarum et 
tuberculis minoribus, interstitiis vix convexis. Long. 3:4 mm. 

Hab. Japan; one specimen (coll. Lewis): Cuina, 
Chusan Is. (Walker). 

This species so closely resembles the common Japanese 
X. validus that it is not necessary to give further 
characters than those contained in the diagnosis. It can 
be distinguished without comparison by the non-im- 
pressed lines of punctures on the elytra, by the long and 
close seriate bristles of the interstices, which arise even 


on the Scolytide of Japan. 577 


up to the base from punctures which have at least the 
anterior margin elevated, so as to be subtuberculate. I 
have seen over a hundred examples of X. validws without 
finding any intermediate forms between the two species. 

The next species should follow Trypodendron quercus, 

on p. 124 :— 
Trypodendron sordidum, sp. u. 

Fem. Cylindrica, subelongata, parum nitida, sordide dilute 
testacea, capite, prothoracis antica dimidia parte medio, elytrorum 
lateribus et apice infuscatis, sat dense pilis subtilibus erectis 
adspersa ; antennarum clava breviter ovali, obtusa; prothorace 
longitudine breviori, anterius fortiter rotundato, apice bituber- 
culato, supra mox post medium obscure transverse elevato, basi 
reticulata, punctata ; elytris dense confuse punctatis, punctis hinc 
illinec in lineas ordinatis, apice convexo, substriato, ruguloso, 
densius piloso. Long. 3°8 mm. 

Hab. Japan; one specimen (coll. Lewis). 

Fem. Somewhat elongate, cylindrical, obscurely shining, 
dirty yellow testaceous, the head, the middle part of the anterior 
half of the thorax, the sides and apex of the elytra infuscate, the 
latter less deeply. Head rugosely punctured in front with a 
median smooth line, with thin erect pubescence, denser over the 
mouth ; antennal club short oval, obtuse in front, with rather thin 
pubescence, chiefly at the sides and apex. Prothorax broader than 
long, widest at the base, the sides strongly rounded from behind 
the middle to the apex, its anterior margin nearly circular, with 
two prominent median tubercles ; surface with an obscure trans- 
verse elevation just behind the middle, asperate anteriorly, the 
asperities forming fine transverse lines, posteriorly finely reticulate 
and rather dull, with fine punctures, subasperate in the middle line 
to the base, pubescence fine erect and short, thinner over the 
median area. Scutellum small, rounded, infuscate. Elytra 
two-thirds longer than the prothorax, and narrower than its base, 
their basal margins separately convex, the shoulders rounded, the 
sides parallel, the apex subcireularly rounded; surface cylindric, 
strongly declivous behind, with rather dense fine semi-erect pubes- 
cence, and close irregular punctuation very indistinctly lineate near 
the suture, apical declivity subconvex, more densely pilose, with 
closer subrugulose punctuation and traces of impressed strie. 
Underside and legs pale testaceous. 


This species is very like 7’. pubipenne, Blandf., but is 
more elongate and cylindrical. The club of the antenna 


578 Mr. W. F. H. Blandford’s supplementary notes 


is in form a short, almost orbicular, oval, whereas in 
T’. pubtpenne it is by comparison slightly but perceptibly 
acuminate. 

In my description of the latter insect the club is 
incorrectly described as non-acuminate. It appears so 
when compared with that of 7. quercus, but not with that 
of the present species. The rows of punctures on the 
elytra of 7’. pubipenne, indistinct though they are, are 
much more evident than in 7’. sordidwm, and the sutural 
row is perceptibly, though weakly, impressed; on the 
other hand, the apex of the elytra in the former species 
shows no trace of striz. 

T. sordidum is even more like the description of 
T. politum, Say, than is 7’. pubipenne, but as it appears 
to be a larger insect, and, as far as I can discover, is 
without any suture on the antennal club, I hesitate to 
identify it with that American species. 


Crossotarsus concinnus, n. n. 


C. chapuisi, Blandf., Trans. Ent. Soc. 1894, p. 129. 

I unfortunately overlooked the fact that Duvivier had 
described an African Crossotarsus under the name 
chapuisi, and therefore substitute the above name for 
the one I previously employed. 


SCOLYTIDAi NIPONIC Ai. 


SpHAEROTRYPES, Blandf. 
pila, Blandf. 
Hyuesinus, Fabr. 


SCOLYTINI. 


Hytastes, Er. 


parallelus, Chap. 
attenuatus, Hr. 
plumbeus, Blandf. 
obscurus, Chap. 
ambiguus, Blandf. 


interstitialis, Chap. 


costatus, Blandf. 
nobilis, Blandf. 
laticollis, Blandf. 
tristis, Blandf. 
cingulatus, Blandf. 
scutulatus, Blandf. 


glabratus, Zett. 


Puxoeosinus, Chap. 
decumanus, Hr. i P 


pulchellus, Blandf. 
dubius, Blandf. 
minutus, Blandf. 
perlatus, Chap. 
seriatus, Blandf. 
lewisi, Chap. 
rudis, Blandf. 


Myenopuinus, Hichh. 
piniperda, Fabr. 
minor, Hart. 


Hyorryyncuus, Blandf. 
lewisi, Blandf. 


on the Scolytide of Japan. 579 


Potya@rapuus, Hr. 
oblongus, Blanidf. 
proximus, Blandf. 
miser, Blandf. 


Scotytus, Geoff. 
esuriens, Blandf. 
agnatus, Blandf. 
frontalis, Blandf. 
aratus, Blandf. 
japonicus, Chap. 
claviger, Blandf. 


TOMICINI. 


Cryprurcus, Hr. 
pusillus, Gy/l. 


CrypHatus, Hr. 
exiguus, Blandf. 


Hypotuenemus, Westw. 
tristis, Hichh. 
peritus, Blandf. 
expers, Blandf. 


Cosmoperes, Hichh. 
consobrinus, Blandf. 


PiryorutHoruvs, Hichh. 
jucundus, Blandf. 


ErporxHe.vs, Kichh. 
imitans, Hichh. 
minutus, Blandf. 


Tomicus, Latr. (1807.) 
cembrae, Heer. 
angulatus, Hichh. 


AcantnHortomicus, Blandf. 
spinosus, Blandf. 


Dryocoetss, Hichh. 
autographus, Ratz. 
pilosus, Blandf. 
affinis, Blandf. 
luteus, Blandf. 


nubilus, Blandf. 
moestus, Blandf. 
dinoderoides, Blandf. 
apatoides, Hichh. 


Coccorryres, Hichh. 


graniceps, Hichh. 
perditor, Blandf. 
advena, Blandf. 


Xyiesorvs, Hichh. 


mutilatus, Blandf. 
brevis, Hichh. 
cucullatus, Blandf. 
amputatus, Blandf. 
lewisi, Blandf. 
rubricollis, Hichh. 
apicalis, Blandf. 
atratus, Hichh. 
germanus, Blandf. 
compactus, Michh. 
semi-opacus, Hichh. 
orbatus, Blandf. 
concisus, Blandf. 
validus, Hichh. 
galeatus, Blandf. 
interjectus, Blandf. 
obliquecauda, Motsch. 
aquilus, Blandf. 
praevius, Blandf. 
seriatus, Blandf. 
pelliculosus, Hichh. 
muticus, Blandf. 
festivus, Hichh. 
glabratus, Hichh. 
bicolor, Blandf. 
attenuatus, Blandf. 
sobrinus, Hichh. 
adumbratus, Blandf. 
badius, Hichh. 
vicarius, Hichh. 
minutus, Blandf. 
schaufussi, Blandf. 
defensus, Blandf. 
exesus, Blandf. 


580 Mr. W. F. H. Blandford’s supplementary notes. 


TRYPODENDRON, Steph. niponicus, Blandf. 
quercus, Hichh. contaminatus, Blandf. 
var.niponicum, Blandf. 
sordidum, Blandf. Prarypus, Herbst. _ 
pubipenne, Blandf. modestus, Blandf. 
SCOLYTO-PLATYPINI. Lease ae 
Sconyto-pLatTypus, Schauf. prikicit: eae : 
tycoon, Blah a amus, Blandf. 
shogun, Blandf. amatus, Blandf. 


daimio, Blandf. 


siomio, Blandf. PI OLA 


snikida oR ieae aculeatus, Blandf. 
PLATYPINI. 
Crossorarsus, Chap. Gernyocrerus, Motsch. 
concinnus, Blandf. adustipennis, Motsch. 


chapuist, Blandf. 107 spp. 


( “peri 


XXV. Descriptions of the Pyralidze, Crambide, and Phy- 
cide collected by the late T. Vernon Wollaston 


in Madeira. By Guorcr T. Bernunz-Baker, 
F.L.S. 


[Read Oct. 17th, 1894.] 


My former notes on Mr. Wollaston’s lepidopterous collec- 
tions in Madeira dealt only with Macro-lepidoptera, the 
present paper will therefore treat of the Pyralide and 
Phycide (Lord Walsingham having most kindly cata- 
logued the Pterophoridz, Tortricide, and Tineide). if 
shall, as before, follow the arrangement of Staudinger’s 
catalogue. 


Aglossa cuprealis, Hb. 


Common, very variable in size, the smallest specimen 
being barely 16 mm., or less than half the size of two or 


three of the largest. All are dark, strongly-marked 
examples. : 


Asopia farinalis, L. 


Not uncommon around Funchal. 


Scoparia decorella, Sttn. 


Described by Stainton inthe Ann. and Mag. Nat. Hist., 
8rd ser., 1859, p. 210, thus :— 


“Alis anticis albidis, striga anteriore recta latiuscula, striga pos- 
teriore tenui arcuata et indentata, externe late saturate fusco 
marginata, signo % ochraceo expleto spatio ante marginali angusto 
albo. Exp.al.81]. In the sharp contrast of colour, this resembles 
E. resinea ; but from that species it is abundantly distinct by the 
form of the second striga, the distinct white submarginal space and 
the ochreous filling up of the mercurial marking. Inhabits Madeira 
proper.” 


This species, though like resinea, is also very close to 


lineola; its proper position will therefore be between 
these two species. 


582 Mr. G. T. Bethune-Baker on the Pyralide, 


Scoparia frequentella, Sttu., and var. Concinnella, Curtis. 


I have before me both typical specimens and Curtis’s 
variety, in which the whole of the ceutral area is dark. 


Scoparia scoriella, Woll. (Zell. in litt.). 


Described by Wollaston (Ann. and Mag. Nat. Hist. 
1858, 3rd ser., I., p. 119) as follows :— 


“Alis anticis subangustis, pulverato fuscis, strigis duobus tenuibus 
arcuatis externe latius fusco-marginatis, signo & obscurius expleto; 
spatio ante-marginali latiusculo, superius nigro-lineato, posteriori- 
bus fusco-cinereis. ¢ ? exp. alar. vix 11 lin. Inhabits Madeira 
proper, occurring in similar spots; to the last-mentioned species ”* 
(2.e., tothe next species following this, viz., stenota). 


This is a somewhat darker insect than the following, 
but may be differentiated by the first line being toothed 
interiorly, not exteriorly, whilst the whole of the space 
from the second line to the hind margin is entirely dark. 
The secondaries are brownish grey. Hxp. alar. 20 mm. 

A rarer insect than stenota. 


Scoparia stenota, Woll. (Zell. in litt.). 


Described by Wollaston in the same work and on the 
same page as the preceding, thus :— 


“Alis anticisangustis apice acuto. Strigis duabus albidis postice 
late nigricanti-marginatis, priore acute fracta posteriore tenui 
biarcuata punctis duobus nigris cum striga, prioris umbra confluen- 
tibus signo % obliquo posterioribus cano-albidis, Exp. alar. vix 
11 lin.  Inhabits Madeira proper; abounding in grassy spots, 
chiefly of a rather lofty elevation.” 


This species is fairly constant in markings. The colour 
of the wings is grey, the primaries in fresh specimens 
being almost entirely covered with very dark scales, the 
two transverse lines are white, the first acutely-toothed 
posteriorly, the second doubly arcuate, and dentated 
interiorly, both are broadly margined on the outside with 
pec A small black dot is in the centre of the wane 


* Ann. and d Mag. Nat. Hist. 1858, 3rd ser., I., p. 119. 


Crambide, and Phycide collected in Madeira. 583 


followed below, but nearer to the base, by a short black- 
ish dash, extending into the tooth of the first transverse 
line. Apex subacute, ¢ with very much narrower wings 
than ¢. Hind wings silver grey. xp. alar. ¢ 18- 
20 mm.; ? 15 mm. The ? is generally decidedly 
darker than ¢. This species will follow angustea. 


Scoparia wollastont, n. sp. 

Primaries white, the two transverse lines irregular, each. inclined 
towards the centre, a very limited portion of base black, followed 
up to the first line by greyish greenish yellow; central area black, 
sprinkled plenteously with scales of the aforesaid greenish ochreous 
colour, with which the upper part of the fairly distinct mercurial 
mark is also filled. Beyond the posterior line and right up to 
margin, the wing is entirely of this same colour, which in this 
larger area assumes a lustrous appearance. At the apex near the 
costa, and also at the anal angle, there is a plentiful sprinkling of 
super-imposed dusky scales; near the apex and also at the anal 
angle are two crescentic white marks close to the posterior margin 
Fringes grey, paler at extremities. The wings are rather broad 
and somewhat rounded at the apex. Secondaries translucent grey. 


One specimen from Madeira; abdomen broken, but 
probably from the antenne a g. Mr. Meyrick says of 
this species, ‘‘ Quite new to me, and a fine species.” 


Hellula undalis, ¥. 


Fairly common ; one specimen labelled Funchal, others 
without definite locality. 


Botys sanguinalis, L., var. Hematalis. 
One beautifully deep pink and yellowexample; parent 
form not rare. 
Botys poligonalis, Hb. 
Abundant at low and intermediate elevations. 
Botys asinalis, Hb. 
One unusually large example. 


584, Mr. G. T. Bethune-Baker on the Pyralide, 


Botys maderensis, nu. sp. 


6 Primaries bright umber-brown, the dark basal line just before 
the orbicular very interrupted ; orbicular stigma fairly distinct, 
finely encircled by a dark line; reniform stigma very distinct, 
finely encircled with blackish, and filled in with dark grey. The 
subterminal transverse curved line is composed of a series of dark 
grey scallops, interrupted at each nervure, and preceded by a dark 
dot, the scallops and dots being most distinct in the middle ; a fine 
shading of the ground colour follows these markings, from whence, 
up to the posterior margin, the whole of the space is filled in with 
very dark grey, getting paler near the margin, which (margin) is 
dotted with blackish at the termination of each vein. Antennz 
sub-pubescent. Fringes dark grey, with paler extremities and a 
pale dividing line. Secondaries uniform dark greyish umber, 
darker near posterior margin, which is darkly dotted as in pri- 
maries ; fringes as primaries. Exp. alar. about 32 mm. 


The ¢ differs in no respect from the ¢ beyond 
being duller in hue; antenne simple. ‘his species is 
nearest Walker’s Scopula delineatalis, from St. Helena. 
It can, however, at once be recognized by its uniformly 
dark secondaries, which in delineatalis are whitish, very 
broadly and darkly bordered; the latter is also of a 
smaller size. 


Seven specimens in both sexes. 


Botys atlanticum, n. sp. 


Primaries uniform pale ochreous grey, with a small dark grey 
dot about the centre of the upper part of the discoidal cell, and 
another at the upper extremity, whilst directly beyond the lower 
extremity is a large conspicuous dark grey spot. The sub-terminal 
line is represented by a curved row of dark grey small dots, one on 
each nervure, from the sub costal to the 3rd median ; the termina- 
tion of each of the veins in both wings is darkly dotted. Fringes 
same colour as wings. Antenne simple. - Secondaries rather darker 
grey, with a dark grey border, broad at the apex, but disappearing 
gradually at the anal angle. In the centre of discal cell is an 
irregular dark grey spot. Fringes whitish. One @ specimen. 
Exp. alar. 33 mm. 


I know of no species at all like this insect, but it 


Crambide, and Phycidx collected in Madeira, 585 


evidently belongs to the same sub-division as the pre- 
ceding species. 


Botys ruficostalis, Ld. 


One specimen from Madeira, and I have recently 
received others from this island. 


Botys numeralis, var. Illutalis, Guen. 


There are five specimens of a Botys very nearly allied 
to numeralis. Guenée says his species may be a local 
variety of it, and it is possible my specimens may be the 
same ; at any rate, I shall temporarily place them under 
this name. 


Nomophila noctuella, Schiff. 
Common up to 2000 feet. 


Margarodes unionalis, Hb. 


Not uncommon. 


Diasema ramburialis, Dup. 


Fairly common. 


Crambus atlanticus, W oll.* 


“Alis anticis apice acutis, saturate griseo-ochreis, costa anguste 
albida, dorso basim versus albido, vitta centrali albida a basi 
perducta, pone medium in ramos, fracta plagaque fusea interrupta, 
plaga:heec postice nigro et albido marginata est ; palpis longius- 
culis ; antennis filiformibus, nec pectinatis ; capite vittaque cen- 
trali thoracica albis. Exp. alar. 13 lin. Inhabits Madeira proper ; 
abounding in grassy spots during the summer months at inter- 
mediate and lofty elevations.” 

Primaries greyish ochre, with a moderately broad white longi- 
tudinal stripe from base to four-fifths length of wing, which 
broadens outwards, and, as the short diagnosis says, is broken into 
veins towards the costa, between which are dark dustings (visible 
only in fresh specimens) ; the ground colour from this stripe to the 


* Ann, and Mag. Nat. Hist., 1858, drd ser., I., p. 119. 
TRANS. ENT. SOC. LOND. 1894.—parT Iv. (DEC.) 2p? 


586 Mr. G. 'T. Bethune-Baker on the Pyralide, ete. 


inner margin is very much darker, often, in fact, ochreous brown. 
“Costa anguste albida.” This is not atall constant. Hind margin 
darkly dotted. Fringes grey, with a double dividing line. Second- 
aries brownish grey, dark at the apex, becoming quite pale near the 
anal angle. Fringes whitish, with a dividing line. Palpiaslong as 
head and thorax, white above, head white, thorax white, with 
chestnut patagiae ; abdomen same colour as secondaries. The ? 
is similar to the ¢, but somewhat paler. 


This species belongs to the ¢ristellus and selasellus 
eroup, but is a finer and handsomer insect. 


EHnomene ocellea. 


I have in Wollaston’s collection two fine specimens, 
and have recently received others from Madeira. 


Myelois cinerella, Sttn. 


There are two fine specimens of this insect in the 
National Museum. Mr. Stainton says (Ann. and Mag. 
Nat. Hist. 1859, 3rd ser., III., p. 211) it is not closely 
allied to any known species, and inhabits the Northern 
Dezerta. His diagnosis is as follows :— 


“Alis anticis angustulis dilute griseo-ochreis, fusco valde suffusis 
striga priore obliqua, posteriore sinuata, ciliis dilute griseo-ochreis. 
Exp. alar. 10—11 lin. An obscure-looking insect, not closely allied 
to any known species; the labial palpi are unusually short, and 
the median vein of the posterior wing is trifid, as in /. com- 
positella.” 


Ephestia gnidiella, Mill. 


A few specimens, which present no unusual features. 


Ephestia interpunctella, Hb. 
Abundant at Funchal. 


Galeria melonella, L. 


Common; one specimen is very remarkable, being 
almost black. 


Achreea grisella, F. 


One or two specimens, which differ in no way from the 
ordinary type. 


THE 


PROCEEDINGS 
LNTOMOLOGICAL SOCLETY 


LONDON 
For THE YEAR 1894. 


February 7th, 1894. 


Henry Joun LHtwes, Hsq., F.L.8., President, in the 
chair. 

Donations to the Library were announced and thanks voted 
to the respective donors. 


Nomination of Vice-Presidents. 

The President announced that he had nominated the Rt. 
Hon. Lord Walsingham, LL.D., F.R.S.; Professor Edward 
B. Poulton, M.A., F.R.S.; and Colonel Charles Swinhoe, 
M.A., F'.L.8., Vice-Presidents of the Society for the session 
1894-95, 


Election of Iellows. 


Mr. Walter F. Baker, of 18, Hyde Terrace, Leeds; Mr. 
Percy M. Bright, of Roccabruna, Bournemouth; Professor 
Lewis Compton Miall, F'.R.$., of the Yorkshire College, 


PROC. ENT. SOC. LOND., I., 1894. B 


(ar) 
Leeds; and Mr. Edwin Wilson, of Cherry Hinton Road, 
Cambridge, were elected Fellows of the Society. 


Exhibitions, etc. 

Mr. Jenner Weir exhibited, on behalf of Mr. J. M. Adye, 
a specimen of Plusia moneta, Fabr., which had been captured 
at Christchurch, Hants, and remarked that this species, which 
had been found in this country for the first time so recently 
as June, 1890, was apparently becoming a permanent resident 
here, as it had been since taken in several of the southern 
counties. The food-plant, Aconitum napellus, though rare in 
England as a wild plant, was very common in gardens. Mr. 
Jenner Weir also exhibited a nearly black specimen of Venilia 
macularia, L., the yellow markings being reduced to a few 
small dots. 

Mr. Hamilton Druce exhibited a female specimen of Hypo- 
chrysops scintillans, lately received by him from Mioko, New 
Ireland. He said that only the male of this species had 
been as yet described. 

Mr. EF. Enock exhibited a nest of the British Trap-door 
Spider, Atypus piceus, recently found near Hastings by Mrs. 
Knock, 

Mr. W. F. H. Blandford stated that he had recently obtained 
an additional species of Scolyto-platypus from Japan, which, 
though closely allied to the species he had formerly described, 
showed a very distinct modification of the male prosternum, 

Mr. M. Jacoby exhibited and remarked on a specimen of 
Leptispa pygmea, Baly, which was doing much injury to 
sugar-cane in the Bombay Presidency of India. Mr. G. C. 
Champion stated that he had found an allied species on 
bamboo. 


Papers read, 

Dr. F. A Dixey read a paper—which was illustrated by 
the oxyhydrogen lantern—entitled ‘‘ On the Phylogeny of the 
Pierine as illustrated by their wing-markings and geographical 
distribution.”’? A long discussion ensued, in which the Presi- 
dent, Mr. Osbert Salvin, Mr. Jacoby, Colonel Swinhoe, Mr. 
Jenner Weir, Mr. Hampson, and Mr, Kenrick took part. 


( tit) 

Dr. T. A. Chapman read a paper entitled ‘‘Some notes on 
those species of Micro-Lepidoptera, allied to Micropteryx, 
whose larve are external feeders, and chiefly on the early 
stages of Hriocephala calthella.”’ Mr. Hampson and the 
President made some remarks on the subject of the paper. 

Mr. Hamilton H. Druce read a paper entitled ‘‘ Description 
of the female of Hypochrysops scintillans, Butl.”’ 

The Rev. Dr. Walker communicated a paper by Mr. R. H. 
F. Rippon, entitled, ‘ Description of a variety of Ornithoptera 
(Priamoptera) urvilliana,” 


February 28th, 1894. 


Colonel Cuartes Swinuorz, M.A., F.L.S., Vice-President, 
in the chair, 


Donations to the Library were announced and thanks voted 
to the respective donors. 


Election of an Honorary Fellow. 

Professor August Forel, M.D., of the University of Ziirich, 
was elected an Honorary Fellow of the Society, to fill the 
vacancy caused by the death of the late Professor H. A. 
Hagen, M.D. 


Elections of Fellows, 
Mr. John Pratt, of the Cedars, New Barnet, and Mr, 
Michael Yeatman Woolf, of 1, Marlborough Place, St. John’s 
Wood, N.W., were elected Fellows of the Society. 


Hahibitions, ete. 

Mr. G. C. Champion called attention to a supposed new 
Longicorn Beetle, described and figured by Herr A. F. Non- 
fried, of Rakonitz, Bohemia, under the name of Callipogon 
friedlanderi, in the Berl. Ent. Zeitschr. 1892, p. 22. He said 
that the supposed characters of the insect were due to the 
fact that the head had been gummed on upside down! He 
also exhibited an extensive collection of Coleoptera and 


(4) 


Hemiptera-Heteroptera made by himself in the island of 
Corsica in May and June last. 

The Rey. Theodore Wood exhibited a variety of Saturnia 
carpini, with semi-transparent wings, a large proportion of 
the scales being apparently absent, bred with several examples 
of the type-form at Baldock, Herts; also a pale variety of 
Smerinthus populi, which was said to have been bred, with 
several similar specimens, from larve marked with rows of 
red spots on both sides. 

Mr. R. South exhibited a variety of Argynnis aglaia, 
approaching the form known as var. charlotta, and a variety 
of EHuchelia jacobee, in which the crimson costal streak 
was continued along the outer margin almost to the inner 
margin, taken by Mr. Fowler at Ringwood, Hants, in 1893 ; 
a variety of Argynnis euphrosyne, taken by Mr. Mead in 
Epping Forest, in 1898; and a series of black and other 
forms of Phigalia pedaria, bred during the present year 
from a black female captured last spring by Mr. Rose, of 
Barnsley. 

Mr. South also exhibited, on behalf of Dr. H. G. Knaggs, 
working models of the mechanical decoy and spring net 
described by the inventor in the Entomologist xxvi., pp. 154 
—157, 180—182, 207—210. He remarked that, although 
the apparatus might appear somewhat complicated in 
character, it would be found on examination to be really 
simple in construction. It could be quickly fixed up for use, 
readily taken apart again, and when packed the whole of the 
spring net could be stowed away in a bag measuring only 
twenty-six inches in length, and cight and a half inches in 
eirth, 

So far the contrivance had not been tested in the field, but 
on one occasion Dr. Knagegs had fitted it up on his lawn in 
Camden Road, and caught a specimen of Pieris brassica. It 
would probably be found of the greatest use to collectors of 
Lepidoptera in tropical countries. If specimens of a desired 
species were attracted by the decoy within the area com- 
manded by the net their capture would be certain, as the 
action of the springs was so rapid that the fastest flying 
butterfly would have little chance of escape. 


(tpt) 


Colonel Swinhoe suggested that a dead animal might be 
used as an attraction instead of the decoy. 

Mr. H. Goss exhibited, for Mr. C. B. Taylor, of Jamaica, 
a coloured drawing of the larva of Papilio homerus, Fab. 

Mr. F. W. Frohawk exhibited drawings showing the com- 
plete life-history of Argynis aglaia and A, adippe, every stage 
being figured ; also enlarged drawings of the segments of the 
larve in their first and last stages, showing the remarkable 
difference in structure. Mr. Merrifield commented on the 
beauty of the drawings. 


Papers read, etc. 

. Mr. G. C. Champion read a paper entitled ‘‘On the Tene- 
brionide collected in Australia and Tasmania by Mr. J. J. 
Walker, R.N., during the voyage of H.M. ship ‘ Penguin,’ 
with descriptions of new genera and species,’’ and he 
exhibited the specimens comprised in the collection. Mr. J. 
J. Walker and Colonel Swinhoe made some remarks on the 
paper. 

Mr. Champion also read a paper entitled “An Ento- 
mological Excursion to Corsica,’ in which he described an 
expedition to the mountains of that island in May and June, 
1898, in company with Mr. R. §. Standen, Mr. A. H. Jones, 
Colonel Yerbury, R.A., Mr. Lemann, Mr. Raine, and others. 
The author stated that Mr. Standen had already given an 
account of the Lepidoptera (Rhopalocera) in the ‘‘ Entomo- 
logist,’’? 1898, pp. 2836—288, and pp. 259—263. 

Mr. Osbert Salvin, Colonel Yerbury, and Colonel Swinhoe 
took part in the discussion which ensued. 

Mr. Edward Saunders communicated a paper entitled ‘A 
List of Hemiptera-Heteroptera collected by Mr. Champion in 
Corsica, with a description of one new species.” 

Mr. W. F. Kirby read a paper entitled “‘ Notes on Dorydium 
westwoodi, Buchanan White, with observations on the use of 
the name Dorydium.,” 

Mr. Charles B. Taylor communicated a paper entitled 
‘¢ Description of the larva and pupa of Papilio homerus, Fab,” 


( (vey 


March /4th, 1894, 


Colonel Cuartes Swinnor, M.A., F.L.8., Vice-President, in 
the chair. 


Donations to the Library were announced and thanks voted 
to the respective donors. 


Election of Fellows. 


Mr. William Bateson, M.A., Fellow of St. John’s College, 
Cambridge ; Mr. H. Caracciolo, of the Port of Spain, Trinidad, 
West Indies; Mr. G. C. Dudgeon, of 53, Montague 
Square, W.; and the Rev. Frank E. Lowe, M.A., of St. 
Stephen’s Vicarage, Guernsey, were elected Fellows of the 
Society. 


The Westwood Bequest. 


The Secretary read the following letter on the subject of 
the Westwood Bequest, received by the Treasurer from Miss 
Kmma Swann, Professor Westwood’s niece, 


‘© 141, Woonsrock Roap, Oxrorp. 
“* February 17th, 1894. 


‘‘Drar Srr,—I beg to enclose to you, as Treasurer of the 
Entomological Society, a cheque for £250, payable to your- 
self, from Miss Lucy Swann and myself. 

‘‘ This sum, in accordance with the wish we-expressed in 
our first communication to your Society, to be invested as a 
separate sum, and not to enter into the general funds of the 
Society ; and the income arising from it to be applied, either 
annually or every two years, as the Council may deem best, 
in paying for Entomological engravings or illustrations to 
be published in your Transactions, every such illustration to 
bear the inscription, ‘‘ Westwood Bequest,’’ and a copy 
of each Transaction containing a ‘‘ Westwood Bequest ”’ 
illustration, to be given to us.—I remain, Sir, yours faithfully, 

‘“ HMMA SWANN, 


‘ Ropert McLacuran, Hsq., F.R.5,” 


Ur sae 


Exhibitions, ete. 


Dr. D. Sharp exhibited a collection of White Ants (Ter- 
mites) formed by Mr. G. D. Haviland in Singapore, which 
comprised about ten or twelve species, of most of which the 
various forms were obtained. He alluded to the bearing of 
the specimens found by Mr. Haviland on the question recently 
discussed by Mr. Herbert Spencer and Professor Weismann, 
as to the mode of development of the various forms of 
individuals of social insects. Mr. Spencer, he said, considers 
that these are due to nutrition at a later stage of the 
development than that at which the sex of the individual 
is determined. Dr. Sharp added that Professor Grassi has 
quite recently informed us that he has found, as the result 
of seven years’ observations on the two European Ter- 
mitids, Calotermes flavicollis and Termes lucifugus, that these 
white ants obtain, when necessary, workers and soldiers from 
individuals destined in the ordinary course to become perfect 
insects, and that they do this by varying the quantity and 
quality of the food supplied to the creatures that are to be 
thus deviated from the normal course of their development. 
In some cases, the Termites thus produce what Grassi calls 
neoteinic queens, that is fertile females, that in some portions 
of the development of the body still retain the immature 
condition ; this transformation the white ants can effect at 
more than one stage of the life-history, but, nevertheless, it 
would appear that they prefer to operate at a particular stage. 
Mr. Haviland’s collection contains examples of these neoteinic 
queens, he having found in one nest, instead of a single 
normal queen, eleven of these neoteinic individuals, thus 
confirming Grassi’s statement, that when the ants produce 
these ‘‘ substitution royalties,’ they operate on numerous 
couples, which are ultimately reduced to a single pair. Mr. 
Haviland’s neoteinic queens were accompanied by neoteinic 
kings, and these, according to Mr. Haviland, behaved in a 
similar manner towards their consorts as perfect kings do to 
perfect queens. The collection made by Mr. Haviland con- 
sists of ten or twelve species, of most of which a great 
proportion of the forms was obtained ; the kings and queens 


(| vill .) 


having been procured as well as workers, soldiers, winged 
individuals, and the immature forms. 

Mr. Haviland gave an account of the manner in which he 
examined the nests so as to find the royal couple, which he 
stated frequently required some hours of search. There is 
great variety in the habits of the Termitide, and Mr. 
Haviland observed that at least one species wanders at large 
after the manner of the Hymenopterous ants; this it does at 
night, and individuals of this species may be found returning 
in the morning to their nest, the workers carrying balls of 
food, and being escorted by soldiers, who, however, carry no 
food. Mr. Haviland further stated that one or two of the 
Singapore species made use of mushrooms, which were 
growing in chambers near their nests, after the manner 
described by Smeathman in the Philosophical Transactions 
of the Royal Society. He had examined the alimentary 
canal and found spores of these fungi therein. Mr. Haviland 
mentioned that in the case of a species which is at present 
committing great havoc in the Museum at Singapore, he had 
not been able to find the reproductive individuals. 

Colonel Swinhoe said that it was generally considered in 
India, that the fertile females were never found in human 
habitations, even when these were much infested by the Anis. 

Mr. H. Goss remarked that the fact that the different forms 
of social insects were produced by nutrition was apparently 
known to Virgil, who, he believed, referred to it, and to the 
subject of Parthenogenesis in Bees, in the ‘‘ Georgies,”’ 
Book iv. Mr. McLachlan, Colonel Swinhoe, Mr. Champion, 
Mr. Jenner Weir, and Dr. Sharp continued the discussion. 

Mr. O. E. Janson exhibited specimens of Dicranocephalus 
adamsi, Pascoe, from Sze-chuen, Western China, and D. 
dabryi, Auz., recently received from the neighbourhood of 
Moupin, in the same district; he observed that, although the 
latter had been quoted by Lucas, Bates, and others, as a 
synonym of adamsi, the two species were perfectly distinct ; 
the females of both were unknown to the authors when 
describing them, and presented a remarkable difference, for 
whilst in dabryi this sex is similar to the male in colour and 
sculpture, in adamst it is entirely dull black, with the upper 
surface minutely and densely punctate, 


© m0) 


Mr. C. O. Waterhouse exhibited, for Mr. KE. A. Waterhouse, 
a specimen of Colias edusa, closely resembling Colias erate, 
a Continental species, which was taken on Wimbledon Com- 
mon; a varied series of Chrysophanus phleas, from Barnes 
Common; and a series of Lycena arion from Cornwall. 


Papers read, etc. 


The Rey. Canon Fowler read a paper entitled ‘‘Some new 
species of Membracide.”’ 

Mr. F. Merrifield read a paper entitled ‘‘ Temperature 
experiments in 1893, on several species of Vanessa and other 
Lepidoptera.”” He said that the winter-pupating brood of 
Pieris napi, like the summer-pupating brood, depends for its 
characteristic seasonal colouring partly, but by no means 
entirely, on the temperature to which the individual is sub- 
jected. Pararge egeria was rendered more vivid in its 
markings and colouring by low temperatures, but there was 
no approach to the bright South Kuropean form, either at 
high or at low temperatures. In Cidaria silaceata of the summer- 
pupating brood, low temperatures caused a resemblance to the 
winter-pupating brood, high temperatures caused a duller and 
more uniform appearance, and reduced the size. In Araschnia 
levana, the brood reared from eggs laid in the spring, high 
temperatures in every case produced the blackish normal 
summer or prorsa form, while severe cold in every instance 
transformed the insect completely into the bright fritillary- 
coloured spring or levana type. In Vanessa polychloros, to use 
the language of Dr. Dixey, who had examined the insects, 
forcing invariably tended to produce yellow, whether the 
pupa was previously warmed or cooled, and refrigeration 
produced increased breadth of the dark border, whether 
followed or not by forcing. By a succession of high and low 
temperatures, presumably ancestral markings were brought 
out in the shape of a faint sub-marginal chain on all the 
wings of yellowish cloudy spots with minute black centres. 
Some of the individuals subjected to severe cold were very 
dark and closely resembled V. xanthomelas. In Vanessa 
atalanta very high temperatures developed the golden brown 
parts, both in extent and brightness, new small scarlet spots 

PROC. ENT. SOC. LOND., L, 1894. c 


C29 


beneath and, above, a scarlet cloudy patch on the forewings, 
between the large white costal spot and the third of the 
outer row of white spots, about which it tended to form 
a scattered ring. Low temperatures caused a great exten- 
sion of the blue-green, lavender and white markings, and 
much suffusion. Some specimens obtained or bred under 
natural circumstances, but in very cold weather, and forwarded 
to him by Mr. J. J. Walker and Dr. Chapman respectively, 
showed effects similar to these, but considerably less in degree. 
In Vanessa c-album both the first or July emergence and the 
later or September emergence are affected by moderately low 
temperatures, but the former responds in a much greater 
degree than the latter; this seemed to be quite in accordance 
with Mr. W. H. Edwards’ observations and experiments on 
the allied North American Grapta interrogationis and Grapta 
comma. In Vanessa io low temperatures disintegrated the 
ocellus on the forewing, reducing it to a chain of white spots, 
and brought out other presumably ancestral features. In 
Vanessa antiopa no results had been obtained, but this was 
perhaps owing to the pup being too old. The experiments 
generally confirmed the previous conclusions of the writer ; 
but there was much more to be learned by further experiments 
in the same direction, and probably in the direction of other 
circumstances affecting the insects—moisture, foodplant, and 
possibly light, electricity, and magnetism, though as to the 
three last, such experiments as had yet been tried by the 
writer had produced no results. 

Dr. F. A. Dixey read a paper entitled, ‘‘ Mr. Merrifield’s 
experiments in Temperature-variation as bearing on theories 
of Heredity,” which was supplemental to the previous 
paper. The author said that the interest of Mr. Merri- 
field’s experiments was much enhanced by the fact that 
many of the changes produced were of a reversionary 
character, and that the restored ancestral features differed 
with the nature of the disturbance. The present series of 
experiments not only confirmed former conclusions to this effect, 
but added new evidence of the same kind, referene2 to which 
was made by Dr. Dixey in detail, with the help of diagrams. 


( xi) 

The question might still be raised whether these were, after 
all, genuine cases of reversion, or whether they were not 
simply to be explained as the like effects of like causes, pro- 
duced de novo in both ancestor and descendant. The latter 
supposition, no doubt, was sufficient to account for some of 
the phenomena observed; but there was a residuum, com- 
prising the more special reversionary features, which could 
hardly be so explained. What was the bearing of these 
latter on the general doctrine of reversion? Current expla- 
nations of atavism as a result of disturbance were inadequate, 
inasmuch as they gave no real reason why the more recently- 
established features should be less stable than those with a 
longer ancestral history behind them. As to the two more 
definite explanations afforded by the theories of Darwin and 
Weismann, there was no doubt that if Darwin’s hypothesis 
of centripetal gemmules were granted, the most usual cases of 
atavism (those following hybridisation) could be explained 
under the theory of pangenesis. The present cases, however, 
stood on a quite different footing, as the new conditions 
determining atavism were only applied at an advanced 
period in the life of the individual, and had no reference to 
the ovum from which that individual originated. On the 
other hand, it seemed that if Weismann’s theory of centri- 
fugal carriers of heredity were assumed, the present instances 
could be explained as being due to the critical influence of 
abnormal temperature-conditions on what Weismann called 
‘the struggle of the ids in ontogeny’’; the new external 
conditions favouring some of the ancestral determinants 
(which ex hypothesi exist in the germ plasm) at the expense 
of those more proper to the species. 

Certain observations seemed to show that some, at least, of 
these features might be hereditary ; and it would be most 
desirable to ascertain whether this were so with all or most 
of them. Their transmission, though not their first appear- 
ance, could be accounted for by pangenesis; but under the 
rival hypothesis it would be necessary—in these cases of 
heredity—to postulate, as Weismann now does, a direct effect 
upon some of the determinants wherever they occur, even in 


( ee?) 

the germ-plasm itself. If all could be shown to be hereditary, 
it would seem to follow that the supposed influence upon the 
struggle of the ids was really inoperative, but the issue would 
still remain open between pangenesis and a direct modifica- 
tion of the determinants in the germ-plasm. If some, as 
seemed probable, turned out to be non-transmissible, the 
effect might be tried of varying the period in the ontogeny 
during which the disturbance was applied, with the view 
of ascertaining whether the intervention of new conditions 
at different stages of the struggle of the ids would not 
produce different results. 

In conclusion, Dr. Dixey pointed out the importance of a 
well-established phylogeny as a basis of investigation, since 
ib was only in groups of which the phylogeny was known that 
the ancestral character of these variations could be pro- 
nounced upon with certainty. For many reasons the 
Lepidoptera formed a peculiarly suitable group for such 
experiments, which might fairly be expected to throw much 
additional light on the complex subject of heredity. 

Colonel Swinhoe referred to Dr. Dixey’s remarks as to the 
phylogeny of Argynnis and Vanessa, and asked if he considered 
the male or the female of Argynnis niphe the older form. 

Mr. Hampson pointed out that the peculiar aspect of 
A. niphe $, was generally attributed to mimicry of Danais 
chrysippus, though he was not himself prepared to endorse 
that opinion. 

Colonel Swinhoe thought that the facts of the distribution 
of the two insects were not incompatible with the supposed 
mimicry. 

Dr. Dixey, in reply, said that he believed the marginal and 
apical areas of dark ground colour in A. niphe 2, were of 
more ancient origin than the tawny colour of the male. The 
special features of mimics were often retained rather than 
acquired, and whether A. niphe were a mimic or not, he 
should be inclined to consider these areas as relics of an 
ancestral feature. The evidence as to the original dark 
ground colour of Argynnis was cumulative, and he begged to 
refer to his paper in Trans. Ent. Soc. Lond., 1890, pp. 102-105, 
for a fuller discussion of the question. 


( xiii) 


March 28th, 1894. 


Henry Joun Exwes, Esq., F.L.8., President, in the chair. 


Donations to the Library were announced and thanks voted 
to the respective donors. 


Election of a Fellow. 


Mr. Percy H. Grimshaw, of 58, Comiston Road, Edinburgh, 
was elected a Fellow of the Society, 


Death of Mr, J. Jenner Weir. 

Mr. McLachlan announced the sudden death, on the 23rd 
inst., of Mr. J. Jenner Weir, who joined the Society in 1845, 
and had been one of its most regular attendants. He also 
commented on the scientific attainments of the deceased, and 
his social qualities. Mr. Goss and Mr. Merrifield also spoke 
of their long friendship with the deceased, and of the respect 
and esteem which they entertained for his varied knowledge 
and amiability of disposition. 


Exhibitions, ete. 

Mr. W. Borrer, jun., exhibited a wasp’s nest which had 
been built in such a way as to conceal the entrance thereto 
and to protect the whole nest from observation. He believed 
the nest to be that of Vespa vulgaris (cf. Proc. Ent. Soc. 
London, 1892, pp. xx and xxi). Mr. McLachlan and Mr. 
Blandford made some remarks on the subject. 

Mr. G. F. Hampson exhibited a specimen of Gandaritis 
flavata, Moore, from the Khasi Hills, and called attention to 
the existence in the males of this species, in the closely allied 
British species Cidaria dotata, Linn., and also in two 
Japanese species (C. agnes, Butl., and an undescribed species), 
of an organ on the under side of the forewing, which he sug- 
gested might be for stridulation ; this organ consisting of a 
small scar of hyaline membrane situated just below the 
middle of vein 2, which is much curved: this scar is fringed 
with long hair, and has running down its middle a row of 
sharp spines situated on the aborted remains of vein Ie, 
which is curved close up to vein 2; the spines would natu- 

PROC. ENT. SOC. LOND., I1., 1894. D 


( xv) 


rally rub against part of the costa of the hindwing, but no 
spines or unusual roughening seem to exist on that or on 
any of the veins on the upper side of the hindwing against 
which they could strike; below the scar is situated a large 
shallow fovea or pit in the membrane, slightly developed in 
dotata and flavata, but much more prominently in the two 
Japanese species, and, should the organ prove to be for 
stridulation, it would probably act as a sounding-board. Mr. 
Hampson said that in the Japanese species C. fivsent, Brem, 
exceedingly closely allied to flavata, the males have no trace 
of this organ; and he hoped that entomologists who have an 
opportunity of observing dotata in life would make some 
experiments on living specimens during the ensuing summer; 
probably confining males and females together would lead to 
some results. The President, Prof. E. B. Poulton, Lord 
Walsingham, and Mr. Hampson took part in the discussion 
which ensued. 
Papers, etc., read. 

The Rey. T. A. Marshall communicated a paper entitled 
‘‘ A Monograph of the British Braconide, Part V.”’ 

Mons. Louis Péringuey communicated a paper entitled 
‘‘ Descriptions of new Cicindelide from Mashunaland.” 

Prof. Poulton gave an account of his recent tour in the 
United States, and commented on the entomological and 
other collections contained in the American museums. Lord 
Walsingham, Mr. Hampson, and the President also made 
some remarks on the subject. 


April llth, 1894. 


Henry Joun Ewes, Esq., F.L.8., President, in the chair. 
Donations to the Library were announced and thanks voted 
to the respective donors. 
Election of Fellows. 


My. Frederic Whitworth Jones, of 68, Carlton Hill, N.W., 
and Dr. William Steer Riding, B.A., M.D., of Buckerell, 
Honiton, Devon, were elected Fellows of the Society. 


( xv) 
Exhibitions, etc. 

The Hon. Walter Rothschild exhibited male and female 
specimens of Ornithoptera paradisea, Stdgr., from Finisterre 
Mountains, New Guinea; O. trojana, Stdgr., from Palawan ; 
O. andromache, Stdgr., from Kina Balu, Borneo; (/netus 
mirabilis, Rothsch., from Cedar Bay, Queensland ; and a few 
other splendid species from the Upper Amazons. The 
President, Mr. J. J. Walker, Mr. Osbert Salvin, Lord 
Walsingham, Colonel Lang, R.E., Mr. Champion, and Mr. 
Hampson made remarks on the geographical distribution of 
some of the species and the elevation at which they were taken. 

Mr. H. Goss exhibited, for Mr. G. A. J. Rothney, several 
specimens of a species of Hemiptera (Serinetha augur, Fab.), 
and of a species of Lepidoptera (Phauda flammans, Walk.), 
the latter of which closely resembled and mimicked the 
former, He said that Mr. Rothney had found both species 
abundantly on the roots and trunks of trees in Mysore, in 
November last, in company with Ants (several species of 
Camponotus and Cremastogaster). The Hemiptera appeared to: 
be distasteful to the Ants, as they were never molested by 
them, and he thought that the species of Lepidoptera was 
undoubtedly protected from attack by its close imitation of 
the Hemipteron. Mr. Goss said he was indebted to Mr. C. J. 
Gahan for determining the species. A discussion followed on 
the mimicking species, in which the President, Mr. Water- 
house, Mr. J. J. Walker, Colonel Swinhoe, Mr. Hampson, 
and others took part. 

Mr. J. W. Tutt exhibited (1) a type specimen of Lycena 
corydon, captured in July, 1898 ; (2) a hybrid male (L. corydon 
and L, adonis), taken in copula with a typical female L. adonis, 
May 20th, 1893; (8) a typical male L. adonis, May 20th, 
1893; (4) a female L. adonis, the pigment failing in one hind- 
wing ; (5) a pale var. of L. corydon, probably to be referred to 
var. apennina of Zeller, usually taken in Italian mountains, 
or var. albicans, H. §., taken in Andalusia. Mr. Tutt re- 
marked that, of the first, Staudinger (Cat. p. 12) says 
‘‘ pallidior,” of the latter ‘‘albicans.’’ He also remarked 
that the hybrid retains the external features of the species 
corydon, but has taken on to a great extent the coloration of 


( yee) 


L. adonis. It was captured in copuld with a female L. adonis, 
ata time when L. adonis was very abundant, and some weeks 
before L, corydon occurred (vide Ent. Record, iv., p. 230). 

The question having been raised by the President as to the 
number of meetings of the Society which it was desirable to 
hold during the year, and the most convenient dates for such 
meetings, a long discussion on the subject ensued, in which 
Mr. Waterhouse, Mr. Salvin, the Hon. Walter Rothschild, 
the Rev. T. Wood, Mr. 8. Stevens, the Rev. Seymour St. 
John, and others took part. 


May 2nd, 1894. 

Henry Joun Exwes, Esq., F.L.8., President, in the chair. 

Donations to the Library were announced and thanks voted 
to the respective donors. 

Exhibitions, etc., 

Mr. 8. Stevens exhibited a specimen of Argynnis aglaia 
var. charlotta, taken by the late Rev. James Watson in the 
New Forest in 1870. 

Mr. J. A. Clark exhibited a curious variety of Chelonia caja, 
having an extraordinary wedge-shaped marking extending 
from the outer margin to the base of the left hindwing, and 
also, on the same wing, a small spot. It was brown and 
white in colour, and had the appearance of having been taken 
from the forewing and inserted in the hindwing. The 
specimen was said to have been taken at Abbotts Wood, 
Sussex, in July, 1892. 

Prof. EK. B. Poulton exhibited living specimens of the larve 
of Gastropacha quercifolia, surrounded respectively during the 
early stages of growth by black twigs and lichen-coloured 
twigs, the food being the same in both cases. All the larvee 
were shown upon a white paper back-ground, but examples of 
the surrounding twigs which produced the change of colour 
were shown beside each batch. Mr. Merrifield made some 
remarks on the subject. 

Papers read, 

Mr. E. Meyrick communicated a paper entitled ‘On Pyra- 

lidina from the Malay Archipelago.” 


( xvi ) 


Mr. C. J. Gahan read a paper entitled, ‘‘ A Supplemental 
List of the Longicorn Coleoptera obtained by Mr. J. J. 
Walker, R.N., during the voyage of H.M.S. ‘ Penguin.’ ”’ 


Special General Meeting. 

Prof. E. B. Pounron, Vice-President, then took the chair, 
and a Special General Meeting, convened under Chap. XVIII. 
of the Bye-Laws, in compliance with the following request, 
was held :— 


** To the President and Council of the Entomological Society. 
‘We, the undersigned Fellows of the Entomological 
Society, require you to convene a Special General Meeting of 
the Society on Wednesday next, the 2nd May, 1894, at 
8.30 p.m., or so soon thereafter as the business of the 
Ordinary Meeting, to be held on that day, shall have been 
brought to a conclusion, for the purpose of obtaining the 
approval, by the Society, of the action taken by the Council, 
in declining to interfere in private disputes between Fellows 


f th iety. 
aay ‘¢Ospert Savin. 


‘“F, D. Gopman. 

‘*W. F. H. Buanprorp. 

“C. J. Ganan. 

‘©H. Goss. 

‘¢G. C. Campion. 
“25th April, 1894.” 


Professor Poulton having addressed the Meeting, Mr. A. 
B. Farn made some remarks. 

Mr. C. G. Barrett then moved the following resolution :— 
‘‘That this Society approves the action of its Council in 
refusing to consider it a part of its duties to judge of disputes 
between the Fellows.” This was seconded by Mr. Water- 
house. 

Mr. Arthur Robinson and Mr. G. T. Bethune-Baker having 
spoken in support of the resolution, it was put to the vote by 
the Chairman and carried unanimously. The Proceedings 
then terminated. 


PROC. ENT. SOG. LOND., 11., 1894, E 


( xviii) 


June 6th, 1894. 


Henry Jonn Kuwes, Esq., F.L.8., President, in the 
Chair. 

Donations to the Library were announced, and thanks 
voted to the respective donors. 


Election of Fellows. 


Dr. K. Jordan, of ‘‘ The Museum,” Tring, and the Honble. 
Nathaniel C. Rothschild, of Tring Park, Tring, were elected 
Fellows of the Society. 


Exhibitions, etc. 


Mr. W. F. H. Blandford exhibited a series of eleven male 
specimens of Rhina barbirostris from British Honduras, of 
which the largest and smallest examples measure respectively 
60 and 17 mm. The difference in bulk, supposing the pro- 
portions to be identical, is as 43 to 1. He remarked that 
this variation of the size is especially common in the Brenthide, 
Cossonide, and other wood-boring Coleoptera. The President, 
Dr. Sharp, the Rey. Canon Fowler, Mr. Jacoby, the Honble. 
Walter Rothschild, Mr. Merrifield, and Mr. Champion took 
part in the discussion which ensued. 

Mr. A. J. Chitty exhibited specimens of Cardiophorus equiseti 
taken near Braunton, on the north coast of Devon, in May, 
1891. Mr. Champion and Mr. Blandford made some remarks 
on the species. 

Mr. McLachlan exhibited for Mr. J. W. Douglas, male 
specimens of a Coccid (Lecanium prunastri), bred from scales 
attached to shoots of blackthorn (Prunus spinosa) received 
from Herr Karel Sule, of Prague. Mr. Douglas communi- 
cated the following notes on the species. ‘‘On the 10th May 
‘* last, the males appeared out of scales attached to the shoots 
“ of blackthorn (Prunus spinosa), received on the 9th from Herr 
‘* Karel Sule, of Prague. These males have no special salient 
*‘ characters, but their white scales are very easy to differen- 
‘* tiate from those of congeneric species, for instead of being 
‘** smooth and translucent, they are opaque and covered with 
‘‘ small granulations,”’ . 


(/ ms.) 


‘< Tn the ‘Annales de la Société Entomologique de France,’ 
*« iii, 211 (1834), under the name of Coccus prunastri, Boyer 
‘“de Fonscolombe briefly described the ¢ scales only, but 
‘« Signoret, in 1872 (op. cit., Hssai sur les Cochenilles, p. 258), 
“‘ fully redescribed both the 9 scale and the enclosed insect, 
‘and also noticed the g scales, but they, he says, were too 
‘‘ immature for him to study. The perfect insect has not 
‘‘ hitherto been seen, or at least its appearance has not been 
‘recorded. There are also some mature @? scales of a pre- 
‘vious year, which were sent detached from the shoots. 
‘« The species is common on blackthorn in France and Ger- 
‘‘ many, and should surely be found in Britain.” 

Lord Walsingham exhibited a series of Cacoecia podana, 
Scop., reared from larve feeding on Lapageria and palms in 
Messrs. Veitch’s conservatories in King’s Road, Chelsea, 
including some very dark (melanic) varieties. The Honble. 
Walter Rothschild stated that he had taken the species on 
lime. Mr. Hampson and Mr. Tutt also made some remarks 
on the habits of the species. 

Mr. C. Fenn exhibited a long series of Selenia lunaria, 
bred from one batch of eggs, which included both the spring 
and summer forms; and also two unforced specimens, which 
emerged in November. He remarked that the variation be- 
tween the two emergences, viz., spring and summer, is con- 
siderable, and also the range of variation inter se, especially 
in the spring form ; but it is very remarkable that the summer 
form has one or two representatives among the specimens of the 
spring emergence. The division of the sexes in the time of 
appearance was also peculiar. Mr. Fenn stated that in August, 
1898, 19 individuals were bred, 18 9, 1 ¢. Two ¢ examples 
appeared in November, and the spring brood began to come 
out in April. The first 20 bred were all females, but among 
the 40 which subsequently emerged, the sexes were evenly 
distributed. He said that the parent female was taken at 
Bexley in May, 18938. 

Mr. F. Lovell Keays exhibited, on behalf of Mr. Arthur 
Lovell Keays, a variety of L. alewis (female), having the 
marginal ocelli on the hind wings entirely without the usual 
orange-coloured lunules. The specimen was captured at 


( xx ) 


Caterham, on May 22nd, 1894, and was the first example of 
the species observed by the captor this season. Mr. Barrett 
and the Honble. W. Rothschild made some remarks on the 
specimen. 

Mr. J. H. Durrant exhibited a series of Steganoptycha 
pygmeana, Hb., taken at Merton, Norfolk, between the 25th 
March and the middle of April last. Lord Walsingham made 
some remarks on the species. 

Mr. H. Goss read an extract from a report from Mr. J. R. 
Preece, H.M. Consul at Ispahan, to the Foreign Office, on 
the subject of damage caused to the wheat crop in the district 
of Rafsinjan, by an insect which was called ‘‘Sen’”’ by the 
natives, and which he described as like a flying bug, reddish 
olive in colour, with heavy broad shoulders. Mr. Goss said 
he had been asked by Mr. W. H. Preece, C.B., to ascertain, if 
possible, the name of the species known to the natives as 
“Sen.” Dr. Sharp said that in the absence of a specimen of 
the insect, it was impossible to express an opinion as to the 
identity of the species. 

The Rey. Canon Fowler exhibited, for Miss Ormerod, 
specimens of Diloboderus abderus, Sturm, Hucranium arach- 
noides, Brull., and Megathopa violacea, Blanch., which she 
had received from the La Plata district of the Argentine 
Territories, where they were said to be damaging the grass 
crops. He also read the following notes from Miss Ormerod 
on the subject :— 


‘‘Torrincton Houss, 
“Sr, ALBANS, 
“ June 5th, 1894. 
‘Dear Canon FowiLer,— 

‘“‘T have recently received from Mr. Hy. Watts, Secretary 
of the South American Land Company, specimens of some 
beetles which are attacking the grass on their enclosed land 
in the La Plata district of the Argentine Territories, and of 
which, as far as I am aware, the habits appear to have been 
very little recorded. 

“The information sent was that in a dry spring a white 
grub did serious damage to the roots of the grasses. The 


( Sexy} 


larve (of which specimens were sent, showing them to be 
Lamellicorn), were stated to be 14 inches long, gths inch 
diameter ; body round, with reddish head; and it was said 
that later on thousands of a kind of horned beetle (pre- 
sumably developed from these larve) appeared out of the 
infested ground. 

‘This is all very plain; but the thing that was not clear 
to the local manager, who appears to be a very good observer, 
is—why do the injuries to grass, and also the beetle presence 
(of whatever species), not occur on the unenclosed Pampas, 
but only, or almost entirely, within the enclosures? and why 
should the destruction of grass be followed by such a growth 
of better kinds, that the infestation was decidedly ultimately 
a benefit ? 

‘‘T found from the notes sent me that the great enclosures 
were especially used for grazing stock, as cattle, horses, and 
sheep, and it was mentioned, ‘Hyery year we have a good 
crop of beetles, which, at certain seasons, you find travelling 
along the cattle tracks in hundreds. It appears to be from 
their eggs that the white grub comes. 

‘On turning out the contents of the bottle, I found that 
most of the contents were ¢ and ? of one of the Dynastide, 
but with them were single specimens of what I took to be 
Scarabeide, but had no means here of determining; there- 
fore asked Mr. Janson’s assistance, who was good enough to 
identify the first (the Dynasfes)as g¢ and ? of the Diloboderus 
abderus, Sturm ; and the two other species, of which, un- 
fortunately, only one specimen of each was sent me, respec- 
tively the Hucranium arachnoides, Brullé, and the Megathopa - 
violacea, Blanch. In the case of this latter, Mr. Janson drew 
my attention to the colour being usually of a more marked 
violet than that of my specimen ; but still as the preservative 
fluid has dried out, the violet colour is not wholly absent. 

“The E. arachnoides will be seen to be a very beautiful 
specimen, the peculiar fork, or pair of long processes, in 
front of the clypeus, being very observable. On seeing these 
two beetles, it occurred to me that they might account, to 
some extent, for the improved state of the ground. If they 
are of the manure-burying kinds, this (so to say) ‘ dibbling’ 


C xx} 


in enrichment all about the pasture land could not fail to be 
serviceable. The point, however, of the respective agricul- 
tural habits of the beetles is what I should greatly 
like to be allowed to bring under the notice of the 
meeting. 

‘«‘ With the manure-burying beetles we might havea straight- 
forward process of the lary doing mischief at the roots of 
the grass, but, at the same time, of the manure helping to 
throw up a good new growth; but the case does not seem 
clear as to the Dynastes, the Diloboderus abderus. The note 
sent of the ravage is not clearly applicable especially to any 
one of the species sent. It says, ‘ They literally dug up the 
earth, leaving it as loose as if a spade had been used;’ 
and ‘they work within a couple of inches of the surface, 
eating the roots of all the grass they find.’ But though we 
have a note of the horned beetles coming up by thousands 
from the destroyed pasture, it seems to me we want observa- 
tion as to whether the two other kinds may not also be present. 
I know that in Caffraria, the Keever beetle, Heteronychus arator, 
Fab. (? Burm.) (one of the Dynastide) does a great deal of 
mischief, especially to wheat; and from notes sent me by Mr. 
Fred. R. Schauble, of Stortge, bites off the crops about an 
inch underground ; but any information as to where a pub- 
lished record of the habits of our 8. American pasture beetles 
are to be found, would be a great help to unravelling the 
observations now on hand, and would be of very serviceable 
interest agriculturally as well as entomologically.—Yours 


very truly, 
‘¢Hieanor A. Ormerop, F.E.S.” 


Mr. Hampson raised an important point as to what was the 
legal “‘date of publication’ of Part I. of the Transactions of 
the Society, 1894. He pointed out that the question of the 
priority of the names of certain new species described there- 
in would depend upon the date of publication. Dr. Sharp 
asked Mr. Hampson to explain what he meant by “ publica- 
tion.” Mr. Hampson replied. 

The Honble. W. Rothschild said that he had been informed 
that when a copy, or five or six copies of the Transactions 


( xxiii) 


of a Society, have passed into the museum of a country, that is 
publication. Mr. McLachlan said he believed that when any 
book was ‘‘ on sale”’ that book was published. 

Mr. Goss explained the reason for the delay in the issue of 
Part I. of the Transactions, 1894. 

The Honble. W. Rothschild suggested that immediately a 
Part was printed one or two copies should be sent to the pub- 
lisher for sale. Dr. Sharp pointed out the practical difficulty 
of fixing the date of publication. Lord Walsingham and 
Prof. Poulton continued the discussion. 

Mr. Hampson moved— 

‘‘That, having regard to the Secretary’s statement, the 
date of publication of Part I. of the Transactions 
of 1894, should be fixed at 2nd May, 1894.” 


This motion was seconded by the Honble. W. Rothschild. 


Prof. Poulton moved as an amendment— 

‘That the Secretary be instructed to ascertain the date 
on which the Part was sent to the Publishers, and 
that that should be fixed as the date governing 
priority.” 

Mr. Verrall then moved, as a further amendment— 

“That the matter be referred to a committee, to consist 
of the Treasurer (Mr. McLachlan), the Honble. 
W. Rothschild, and Mr. Hampson.” 

This motion was seconded by Mr. R. W. Lloyd, and carried 
by a large majority. 


Paper read, 


Professor Franz Klapalek, of Prague, communicated a paper 
entitled, ‘‘ Descriptions of a new species of Raphidia, L., and 
of three new species of Trichoptera from the Balkan Penin- 
sula, with criticalremarks on Panorpa gibberosa, McLach,”’ 


Special General Meeting. 

Lorp Wausinenam, Vice-President, then took the chair, 
and a Special General Meeting, convened under Chap. 
XVIII. of the Bye-Laws, in compliance with the following 
request, was held :— 


( xy 3} 


“23rd April, 1894. 
‘¢ GENTLEMEN,— ; 

‘* We, the undersigned Fellows, request, in accordance 
with Chap. XVIII. of the Society’s Bye-Laws, the President 
and Council of the Entomological Society of London to con- 
vene a Special General Meeting to consider :— 

“© (1) The matter of Mr. Farn’s Circular of April 14th. 

‘“‘(2) Whether any amendment of the Bye-Laws is 

necessary in consequence. 


‘«« We are, Gentlemen, yours faithfully, 
‘“« (Rev.) J. Greene, M.A. 


‘‘FRepk. C. Apams. 

“© R. A. Datuas-BEEcuine. 
‘OW. H. Buaser. 

‘¢ Joun E. Rosson. 

‘°C, Fenn. 


**To the SECRETARIES, Entomological Society of London, 
‘11, Chandos Street, Cavendish Square, W.’’ 


The Secretary, having read the above letter of request for 
a Special General Meeting, Lord Walsingham called upon 
one of the Fellows who had signed it to explain the objects 
for which the meeting had been convened. 

Mr. Charles Fenn then offered some explanation of the 
objects of the request for the meeting. 


Mr. Hampson made the following motion :— 


‘‘That this meeting wishes to express its strong disap- 
proval of the printed circular sent round to the 
Fellows by Mr. Farn, and the imputations cast on 
the Council and President, and that having done 
this it declines to further consider the matter.” 

Mr. Bethune-Baker seconded this motion. 

Mr. Farn, Mr. Elwes, Dr. P. B. Mason, Prof. Poulton, 
Mr. Tutt, Mr. Blandford, and Mr. Merrifield having spoken 
on the motion, the Chairman put the motion to the vote, 
and it was carried by a large majority. The Proceedings 
then terminated. 


( eve) 


October 8rd, 1894. 


The Rt. Honble. Lorp Waustnenam, M.A., LL.D., F.R.S., 
Vice-President, in the chair. 


Donations to the Library were announced, and thanks 
voted to the respective donors. 


Election of a Fellow, 


Mr. Alick Marshall, of Bexley, Kent, was elected a Fellow 
of the Society. 


Hahibitions, ete. 


Mr. W.F. H. Blandford exhibited specimens of a sand-flea, 
chigoe or nigua, received from Mr. Szigetvary, of the 
Imperial Maritime Customs, China, who had found them in 
the ears of sewer-rats trapped at Ningpo. Mr. Blandford 
stated that the species was allied to, but perhaps not identical 
with, the American species, Sarcopsylla penetrans, L., one of 
the most troublesome pests in Tropical America and the West 
Indies to man and various domestic and wild animals, the 
female burrowing into the skin, usually of the feet, but also 
of any other accessible region. He said that the distribution of 
the chigoe was recorded over Tropical America and the Antilles 
from 380° N. to 30° §., and of late years it had established 
itself in Angola, Loango, and the Congo. Mr. Blandford also 
exhibited a series of Coleoptera from old collections, in a very 
dirty condition, of which half of the specimens had been 
cleaned by the rapid and effective process of immersing them 
in a solution of potassium hydrate. Colonel Swinhoe, Mr. 
McLachlan, Lord Walsingham, Mr. H. Goss, Mr. Champion, 
Mr. J. J. Walker, Mr. Barrett, and others, took part in the 
discussion which ensued. 

Mr. F. C. Adams exhibited a specimen of Mallota eristaloides, 
a species of Diptera new to Britain, taken by himself in 
the New Forest on the 20th July last. He said that the 
species had been identified by Mr. Austen, of the British 
Museum, and that he had presented the specimen to the 
National Collection. Mr. Verrall made some remarks on the 

PROC. ENT. SOC, LOND., Iv., 1894. F 


( . xxvit ) 


species and on the distribution of several allied species in the 
United Kingdom. Lord Walsingham, as a Trustee of the 
British Museum, expressed his satisfaction at the presentation 
of the specimen to that Institution. 

Mr. Tutt exhibited specimens of Zygena exulans, and read 
the following notes on the subject:—‘The type of Zygena 
“evulans is stated in Staudinger’s Catalog to inhabit the 
‘‘¢ Highest Alps and Pyrenees.’ Its var. vanadis, Dalm., is 
‘stated to be found in ‘Lapland and the Scandinavian 
‘«« Mountains,’ and is differentiated from the type as being 
“¢ narcissime squamata, albo non mixta.’ The assumption 
‘‘ from this then is that the type is more thickly scaled and 
‘‘ mixed with white. In the last week of July, Dr. Chapman 
‘found in the Alps (in the La Grave district), at a consider- 
“able height, a form of Zygena exulans well scaled, as the 
“« species runs, and with the nervures and fore-legs of a decidedly 
“orange colour. The three specimens which he brought 
“away were large. A fortnight later, on a hill about 1000 
“feet above Cogne, specimens were taken, dark in colour, fairly 
‘¢ well scaled, but without any pale nervures (apparently males). 
‘“« A day or two later, more specimens were taken, similar to the 
‘“‘ La Grave district specimens, but with the nervures less orange 
‘« (apparently females), high up the Grauson Valley. Then, on 
‘the 17th August, Dr. Chapman captured a large number in 
“the Grauson Valley, some dark and well scaled, without 
‘the pale nervures, others generally less densely scaled with 
‘strong white nervures and marginal line to anterior wings: 
“to a large extent these characters seemed sexual, but not 
‘‘entirely so. Some years ago specimens of this species were 
‘taken at Braemar, by Dr. Buchanan White, and owing to the 
“‘ lack of scales which some of them showed, they were described 
‘‘as a subdiaphanous form under the name of var. subochracea. 
‘Tn 1886 Messrs. Tugwell and Lachlan Gibb captured a large 
“number of specimens, some of which were in moderate 
‘condition, and it was very evident that Dr. White’s varietal 
‘‘name would hardly answer except for worn specimens, 
‘‘ During the past two years a very large number of specimens 
“have been captured (and some probably bred), and of these 
‘T have seen a very considerable number. It is very evident 


( ‘xxvit ) 


‘from these that the Scotch specimens in good condition exhibit 
‘“‘no difference from many of the Grauson insects. ‘They pre- 
‘sent the two ordinary forms, the well-scaled form without 
‘‘the pale nervures and pale outer margin, and the form 
‘less well scaled with these parts of the wing pale. I 
“exhibit Scotch specimens of each form. It would appear 
“that Staudinger was unaware of the existence of the 
‘« Ta Grave form, in which the orange mixing is so prevalent, 
‘©and which seems a well defined local race. The form with 
“white markings (feste Staudinger) would appear to belong 
‘‘ principally to the female sex. Var. vanadis would appear 
‘to be a rubbed form of the male, in which the scales have 
‘“been partially removed. The Scotch specimens recently 
‘‘ obtained, prove conclusively that in fine condition they are 
‘as densely scaled as the Swiss specimens, and that most of 
‘the specimens in our cabinets regarded as a subdiaphanous 
‘‘ variety ought to be named var. rubbedaria. It would ap- 
‘‘pear easy to explain the rubbed condition of most of the 
‘‘early-caught specimens. ‘The insect is a regular sun lover, 
‘Cand abounds locally on flowers in hot sunshine. In dull 
‘‘ weather, however, it is not at all easy to find specimens. In 
‘Switzerland, sunshine is very frequent, and the consequence 
“ig a collector gets a number of specimens as soon as they 
‘‘have emerged. In Scotland there is comparatively but 
‘little sun, and our collectors only went for an odd day now 
‘“‘and again, because of the nature of the locality. The 
“‘consequence is that they were very much more likely to 
‘“‘choose an unfavourable than a favourable day (most days 
‘being of the former class), whilst the possibility of hitting 
‘the exact day by an occasional visitor is only remote. Now 
‘that our collectors stay in the locality, we get the specimens 
‘‘as fine as they get them in Switzerland.” 

Mr. P. M. Bright exhibited a series of remarkable varieties 
of Arctia menthastri from N. Scotland, also series of Liparis 
monacha (including dark varieties) and Boarmia roboraria from 
the New Forest; Zygaena exulans from Braemar; Noctua 
glareosa from Montrose and the Shetlands ; Agrotis pyrophila 
from the Isle of Portland, and Pitcaple, N.B.; red varieties 
of Teniocampa gracilis; and a specimen of Sterrha sacraria, 


( sxvii >) 


taken at light, at Mudeford, in October, 1893; also living 
larvee of Hmydia cribrum. 

Mr. J. J. Walker exhibited a living specimen of a large 
species of Pulex, which he believed to be Hystricopsylla talpa, 
Curtis, taken at Hartlip, Kent. Mr, Verrall and the Chair- 
man made some remarks on this and allied species. 


Papers read. 

Mr. Kenneth J. Morton communicated a paper, entitled, 
‘‘ Palearctic Nemoure.”’ 

Lord Walsingham read a paper, entitled, ‘‘ A Catalogue of 
the Pterophoride, Tortricide, and Tineide of the Madeira 
Islands, with Notes and Descriptions of New Species.” In 
this paper sixty-six species of Lepidoptera belonging to these 
families were recorded as occurring in the Madeiras, of which 
thirty were noticed as peculiar to the Islands, twelve as 
common to the Madeiras and Canaries, of which two were 
not known as occurring elsewhere, and one extends its range 
only to North Africa. Over thirty species were added to the 
list, and one new genus, seven new species, and two new 
varieties were described. Herr Jacoby and Mr, Bethune-Baker 
made some remarks on the species and their geographical 
distribution. 

Mr. Blandford read a paper, entitled, ‘“‘ A Supplementary 
Note on the Scolytide of Japan, with a list of Species.’’ 


Oct. 17th, 1894. 


Henry Joun Enwes, Esq., F.L.8., F.Z.S., President, 
in the chair. 


Donations to the Library were announced, and thanks 
voted to the respective donors. 


Election of a Fellow. 


Dr. H. G. Breyer, of Pretoria, Transvaal, South Africa, 
was elected a Fellow of the Society. 


(CRsetx, -) 


Exhibitions, etc. 

Mr. G. C. Champion read a letter, dated 15th August 
last, from Mr. J. Y. Johnson, of Funchal, Madeira, on the 
subject of a recent visitation of Locusts to the Island, and 
exhibited specimens. Mr. Johnson mentioned that Darwin, 
in his ‘ Origin of Species,’”’ recorded that in November, 1844, 
dense swarms of locusts visited Madeira. He said that 
since then, until August last, these insects had not visited 
the Island. Mr. Champion remarked that the species sent 
by Mr. Johnson was Decticus albifrons, Fabr., not a true 
migratory locust. Mr. Champion also exhibited specimens 
of Anthaxia nitidula, Velleius dilatatus and Athous rhombeus, 
taken in the New Forest during the past summer. 

Mr. H. Goss read a letter he had received from Captain 
Montgomery, J.P., of Mid-Ilovo, Natal, reporting vast 
flights of locusts there, extending over three miles in length, 
on the 81st August last, and exhibited’ a specimen of the 
locust, a species of Acridiwn. Captain Montgomery stated 
that, as a rule, his district like most of Natal was free 
from the pest, but that an exceptional invasion had occurred 
in 1850. 

Mr. J. W. Tutt exhibited four typical specimens of 
E'mydia cribrum from the New Forest, and read the following 
notes :—‘‘ These four specimens, as will be seen, are well 
‘spotted and streaked, transversely and longitudinally, with 
‘black. The type is stated by Staudinger in his Catalog to 
‘be distributed over ‘ Central and Northern Kurope (except 
“«¢the Polar Region) ; Ural; and, doubtfully, from the moun- 
‘¢<«tains of Andalusia.’ Above these are four specimens of the 
“var. candida of Cyrilli. The specimens came from Cour- 
‘“mayeur, on the Italian side of Mont Blanc. The species 
‘‘ was taken at a height of some 4,000 feet above sea level, on 
‘‘yough ground, on the borders of a larch wood. It was 
‘rather readily disturbed during the late afternoon, settling 
‘‘rapidly again either on the bushes or on dead sticks, with 
‘‘ which the ground was strewn. It was met with again in 
‘the Cogne Valley, from about 6,000 to 8,000 feet, towards 
‘‘Chevanix. Staudinger describes the variety as ‘alis 
‘‘¢anterioribus albis, puncto medio duplo punctisque mar- 


( yee Y) 


‘«¢ ginalibus nonnullis nigris ;’ and gives as its range ‘ Italy, 
«Southern Germany, Alpine Valleys; Northern Spain, 
«Dalmatia, and, doubtfully, Bithynia.’ The species is 
‘very variable. Besides the above, Staudinger describes 
‘a var. punctigera, Fu., as being ‘alis anterioribus punctis 
‘¢nieris paucis;’ with a range spreading over ‘Gaul; 
““¢Southern Germany; and Alpine Valleys.’ He also 
‘‘ describes var. rippertii, Bdy., as ‘alis anticis nigricantibus’ 
‘‘from the ‘Pyrenees’; then a var. inquinata, Rambur, as 
‘“‘¢capite flavescente, alis anticis flavo-albidis, punctis non- 
‘“ ¢nullis marginalibus mediisque nigris,’ and, lastly, a doubt- 
‘ful var. chrysocephala, Hb., from ‘Andalusia, Mauritius, 
‘“¢and, doubtfully, from Southern Russia.’ His diagnosis 
‘runs ‘capite ochraceo, alis anticis albis punctis marginalibus 
6 sempre Ts) 

Mr. R. Adkin exhibited for Mr. H. Murray a specimen 
of Hrebia cthiops, in which the left forewing was much 
bleached, taken in August last, near Carnforth. Mr, Adkin 
also exhibited a series of Acronycta rumicis from Co. Cork, 
Ireland, including light and black forms, with examples 
from the Scilly Isles ; Isle of Man; and North of Scotland 
for comparison. 

Mr. Elwes exhibited a series of Chionobas alberta (male and 
female), Chionobas uhleri, var. varuna, and Hrebia discoidalis, 
from Calgary, Alberta, N.W. Canada, collected in May last, 
by Mr. Woolley-Dod. He said that the validity of C. alberta, - 
which had been questioned by Mr. W. H. Edwards, was 
fully established by these specimens. 

Professor Poulton gave an account of the changes 
he had recently made at Oxford in the arrangement 
of the Hope Collections in the Department of Zoology, 
and as to the laboratory and rooms now available for 
students working at these collections. 


Paper Read. 


Mr. G. T. Bethune-Baker communicated a paper, 
entitled ‘Descriptions of the Pyralide, Crambide, and 
Phycide, collected by the late T. Vernon Wollaston in 
Madeira.”’ 


( Soar 7) 


November 7th, 1894. 


Colonel Cuartes Swinnoz, M.A., F.L.S., F.Z.S., Vice- 
President, in the Chair. 


Donations to the Library were announced, and thanks 
voted to the respective donors. 


Election of Fellows. 

Mr. W. P. Blackburne-Maze, of Shaw House, Newbury, 
Berkshire; and Mr. Bertram George Rye, of 212, Upper 
Richmond Road, Putney, §.W., were elected Fellows of the 
Society. 

EH xhibitions, etc. 

Colonel Swinhoe exhibited a female of Papilio telearchus, 
Hewitson, which he had received by the last mail from 
Cherra Punji. He said that this was the only known speci- 
men of the female of this species, with the exception of one 
in Mr. L. de Nicéville’s collection, which he had described 
in the Journal of the Bombay Natural History Society in 
1898. He also exhibited a male of the same species for 
comparison. 

Mr. C. G. Barrett exhibited abnormal forms of Pararge 
megera, P. egeria, Melitea athalia, Chrysophanus phleas, 
Chareas graminis, Lophopteryx camelina, Plusia gamma, 
Cucullia chamomille, Boarmia repandata, var. conversaria, 
Cidaria psittacata, and other species, all collected by Major 
J. N. Still on Dartmoor, Devon. He also exhibited for Mr. 
Sydney Webb, of Dover, a long series of most remarkable 
varieties of Arctia caja, namely, one spotless, cream-coloured 
and scarlet; one almost spotless, deep brown and black; 
one almost spotless, deep brown and black on one side 
only ; one with the forewings suffused with pink; one with 
the forewings spotless dark brown, hindwings brownish- 
red, spotted; two yellowish, with markings suffused with 
brown; one with all the usual markings delicately pale yet 
distinct ; and others variously suffused on fore or hindwings 
with dark colouring. He also exhibited for Mr. Sydney Webb 
the following varieties of Arctia villica, viz., one wholly 
cream-coloured except the edges of the front wings, which 


( xx) 


were black; two with the cream-coloured spots united in 
varying degrees; two with the hindwings much suffused 
with black; two with wings differently spotted on the 
opposite sides; and one with the hindwings nearly spotless 
but bordered with black. 

Mr. Gervase IF’. Matthew exhibited seven beautiful and 
striking varieties of Arctia villica, bred from larve obtained 
on the Essex coast, near Dovercourt, in March and April, 
1893 and 1894. 

Herr Jacoby exhibited two specimens of Blaps mucro- 
nata, with elytra which had not hardened although exposed 
to the air for a long time, taken on a wall at Hampstead. 
The Rey. Canon Fowler and Mr. G. C. Champion made some 
remarks on the subject of the elytra of immature beetles. 

Mr. H. Goss exhibited a specimen of Periplaneta austral- 
asie, received from Mr, C. E. Morris, of Preston, near 
Brighton. Mr. McLachlan said the species had been intro- 
duced into this country, but was now considered a British 
insect. 

Mr. B. G. Rye exhibited specimens of the following rare 
or local species of Coleoptera :—Cicindela germanica, from 
Swanage; Humicrus rufus, from Shirley ; Triarthron markelt, 
from Lewisham; Mezium affine, from Shoe Lane, H.C.; 
Homaloplia ruricola and Anomala frischi, var. julii, from 
Swanage; Synaptus filiformis, from Sunbury; Livxus para- 
plecticus, from Wicken Fen; Balaninus cerasorum, from 
Wimbledon; Asemum striatum, from the New Forest; and 
Zeugophora flavicollis, from Wimbledon. 

Mr. McLachlan exhibited for Mr. G. C. Bignell, of 
Plymouth, two new species of Ichneumonide, from Devon- 
shire, viz., Pimpla bridgmani, Bign., a parasite on a spider, 
Drassus lapidicolens, Walck., and Praon absinthii, Bign., a 
parasite on Siphonophora absinthii, Linné; together with 
Pimpla epeire, Bign., also a parasite on a spider. 

Mr. C. O. Waterhouse stated that the Acridium received 
by Mr. Goss from Capt. Montgomery, and exhibited at the 
last Meeting, was Acridium septemfasciatum, and he 
exhibited the species with the wings extended. 

Mr. Ridley exhibited a species of a scale insect (? Lecantwm) 


( ‘XXxXHI}) 


found on a nutmeg tree in Malacca, and made some remarks 
on Formica smaragdina, which makes its nest on the trees, 
joining the leaves together by a thin thread of silk at the ends. 
He stated that the first step in making the nest is for several 
ants to bend the leaves together and hold on with their hind 
legs, and one of their number after some time runs up with a 
larva and irritating it with its antenne makes it produce a 
thread with which the leaves are joined; when one larva is 
exhausted a second is fetched, and the process is repeated. 


Paper read. 

Mr. Waterhouse read the following paper entitled, ‘‘ Some 
‘‘remarks on the Antenne of Insects.” 

““T have lately been preparing a series of antenne for 
‘‘exhibition in our Museum. There were one or two points 
‘upon which I wanted information, but which I failed to 
‘‘ find recorded in a satisfactory manner; and there are one 
‘‘or two points, which, although touched upon by Herr 
‘‘ Kolbe in his excellent work, ‘ Kinfiihrung in die Kenntnis 
‘“« «der Insekten,’ have not had sufficient prominence given to 
‘‘them. I venture to call attention to these this evening. 

‘Tn the first place, | wanted to know the extreme limits in 
‘the number of joints in the antenne. I found these varied 
“from two in Platyrrhopalus and some other species of 
‘** Coleoptera, to 480 in Meroncidius, one of the long-horned 
‘‘ Locusts. Articerus, one of the Pselaphide, is said to have 
‘an antenna consisting of a single joint, but my examination 
‘‘did not satisfy me that this was really the case. Possibly 
‘“some Locusts may have even a greater number than the 
‘species I counted. In the Lepidoptera I found the number 
‘‘ of joints varied from 17 in Oncopera, one of the Hepialide, 
‘to 120 in Thysania, a large Mexican Noctuid. 

‘“‘The next point I would call attention to is the club of 
“the antenna. In certain groups the antenne always 
‘terminate in a club; but it is curious to note certain 
‘species appearing with a clubbed antenna, when all their 
‘allies have simple antenne. In the Longicorn Coleoptera, 
‘‘for example, there are some 10,000 species, all with more 
‘or less linear or acuminate antenne, but Telocera, an 


( &Zxexv ) 


“ Australian insect, has a distinct club. In the Aculeate 
‘“‘ Hymenoptera again the antenne are almost always of a 
‘‘simple character, and more or less acuminate, but Nomia 
“ antennata has a distinct club, as has also Steganomus, and 
‘in another genus, Thaumatosoma, a different form of club is 
‘met with. The bi-pectinate antenna of Psammotherma, one 
“of the Mutillide, is another case of a form of antenna 
‘presenting nothing special in itself, but remarkable as 
‘ standing alone among the Aculeata. 

“In the Diptera we see filiform antenne, consisting of a 
‘series of cylindrical or bead-like joints (e.g., Tipula, 
‘* Cecidomyia), and others of the disk and arista type, the 
‘intermediate forms being seen in Xylophagus, Oxycera, 
** Chloromyia, and Sargus. 

‘“‘The antenne of this latter type are generally very small 
‘“and inconspicuous. In certain Dolicopide, however, the 
‘‘arista is sometimes very long, and here, again, we meet 
‘“‘with a club where it might be least expected; I would 
‘‘particularly mention a Psilopus from Ceylon, a_ pretty 
“‘oreen fly five millimetres long, having antenne six milli- 
‘metres in length, terminating in a flat club. 

‘“The clubbed antenna appears among the Neuroptera in 
“the Myrmeleonide and Ascalaphide, being most pronounced 
‘‘in the latter. 

‘“‘In the Orthoptera a club is very rare, but is seen in 
“‘ Gomphocerus. In the Hemiptera a few species have a 
‘slight club, as in Verlusia. 

‘¢T will now just mention another point. In the antenna of 
‘‘the little water-beetle, Parnus, the antenne are very short, 
‘‘ and the second joint is much enlarged and produced forwards. 
‘« It is curious to notice a very similar development in Gyrinus, 
‘a water-beetle belonging to a totally different family. And 
‘in the great water-bug, Belostoma, the second joint is also 
‘‘ produced forwards, but in this case the third joint is simi- 
‘larly formed, and the fourth has a small hook also turned for- 
“wards. It is difficult to suggest a satisfactory reason for this.”’ 

Mr. Champion, Herr Jacoby, Mr. McLachlan, and Mr, 
Gahan took part in the discussion which ensued. 


( fuxxy) 


December 5th, 1894. 


Hewry Joun Exwes, Esq., F.L.8., F.Z.5., President, in the 
chair. 


Donations to the Library were announced, and thanks 
voted to the respective donors. 


Election of Fellows, 


Mr. EK. Augustus Bowles, M.A., of Myddelton House, 
Waltham Cross, Herts; Mr. E. C. Cotes, of the Indian 
Museum, Calcutta; Mr. Joseph W. Green, of West Lodge, 
Blackheath, S.E.; Mr. Henry Keeble, of 10, Coleman Street, 
E.C.; Mr. Thomas Turner, of Cullompton, Devon, and 
Mr. F. H. Wolley-Dod, of Calgary, Alberta, Canada, were 
elected Fellows of the Society. 


Hahibitions, ete. 


The Secretary read a letter from Mr. McLachlan, the 
Treasurer, on the subject of the Committee,* which had been 
appointed, consisting of the Treasurer, the Honourable Walter 
Rothschild, and Mr. Hampson, for the purpose of settling the 
date of publication of Part I. of the Transactions of 1894. 
Mr. McLachlan reported that the Committee had ascertained 
that the Part was first obtainable from the publisher’s on 
May the 11th, 1894, and that this was therefore the date of 
publication. 

Mr. F. Merrifield exhibited hybrids belonging to the genus 
Saturnia, obtained by Dr. Standfuss, of Ziirich; viz. a male 
and female hybrid from a male of Satwrnia pavonia and a 
female of Saturnia pyri, to which he had given the name of 
Saturnia emilie; also hybrids from what Dr. Standfuss 
described as ‘‘a male of Callimorpha dominula var. persona ”’ 
(received from Tuscany) and a typical female of Callimorpha 
dominula, to which he had given the name of romanovi. Mr. 
Merrifield remarked that the so-called var. persona differed 
entirely from the type of Callimorpha dominula. 


* See Proc. of Meeting on June 6th, 1894, p. xxiii, ante.—H. G. 
PROC. ENT. SOC. LOND., v., 1894. G 


( =>) 


Mr. J. W. Tutt exhibited specimens of a very small form 
of Huchlcé, and read the following notes: 

‘* As the Fellows of this Society well know, the Rey. F. B. 
‘“‘Newnham, of Church Stretton, Salop, has sought during 
‘‘the year in the pages of ‘The Entomologist’s Record, etc.,’ 
‘‘to establish two species of /Huchloé, distinguishing from 
“ EF, cardamines, » form ‘much smaller than FZ. cardamines, 
‘«¢ measuring, on an average, only about an inch and a quarter 
‘‘<from tip to tip of the forewings; the discoidal spot being 
‘“‘¢ placed, as in LH. turritis and EH. gruneri, at the juncture of 
“« «the orange and white spaces, not, as in ’. cardamines, well 
‘¢¢ within the orange tip. When viewed under the microscope, 
‘“¢ the wing-scales appear very different from those of FH. 
‘*“¢cardamines. He then proposed for this supposed new 
‘species the name of L’. hesperidis. Mr. Newnham sent me 
‘Can example of #. hesperidis and asked me whether I would 
«exhibit it at one of the Societies’ meetings where there 
‘‘¢are some Entomologists acquainted with Continental 
“‘¢insects.’ I have, therefore, brought the specimen for 
‘‘ exhibition to-night. 

‘‘ With regard to this, I may say that Staudinger describes 
‘an Italian form as ‘ab. (et var. ?) turritis, Och., iv., p. 156, 
‘““¢ab, minor, alis anterioribus puncto, nigro part. albam 
‘‘¢attingente.’ Mr. Newnham, owing, it would appear, to 
‘‘an error made (and corrected ‘ Ent. Ree.,’ v., p. 146) by 
‘Mr, Kirby, considered turritis distinct ; I do not know that 
‘there is any evidence tending to prove that it is anything 
“but a small form of cardamines. 

‘“‘There has been, of course, considerable discussion as to 
‘‘these small specimens of Muchloc, but until the supposed 
‘species is bred and some really good characters obtained 
‘from the earlier stages, it appears totally impossible to 
‘‘accept it as a species on such slender characters as the 
‘‘imago affords. KH. cardamines of normal size varies end- 
‘lessly in the position of the discoidal spot with regard to the 
‘‘ orange blotch (vide, ‘ Ent. Ree.,’ etc., vol. v., pp. 173, 174), 
‘‘where I have given a somewhat long table. The difference 
‘in scaling, if only comparative, is to be expected in small 
‘‘specimens, for it is well-known now that most small 


( “xxzvit *) 


‘aberrations owe their diminutive stature to a deficiency of 
‘food, and this would show as strongly in the scales as in 
‘‘other structures. Reference to these small specimens are 
“scattered through many Entomological Magazines, and 
‘« these records extend over many years. We find such in ‘ The 
‘Northumberland and Durham Catalogue,’ by Mr. Wailes ; 
‘‘in Newman’s ‘ British Butterflies;’ in the ‘EK. M. M.,’ 
‘¢ vol. xxv., by Mr. C. G. Barrett, etc. Mr. T. D. A. Cockerell 
*«(¢ Kntom.,’ vol. xxii., p. 176) calls this small form Fuchloé 
‘* cardamines var. minor, a name I presume that will have to 
‘‘sink in favour of turritis, Och. 

‘« Personally, however, I am quite open to conviction and 
“shall be pleased to agree with Mr. Newnham as soon as I 
‘‘am satisfied as to its specific distinctness, even by one 
‘single indubitable character. 

“To exemplify the difference in size which cardamines 
‘“reaches, even in the same locality, I exhibit three specimens 
‘captured in Chattenden. The large female (al. exp. 20625 
‘‘inches) was taken there, May 22nd, 1888. The small 
“‘female (al. exp, 1°25 inches) and the male (al. exp. 1813 
‘‘inches) were both taken on May 28th, 1892. The speci- 
‘men of hesperidis is rather larger than my small female, 
‘‘being 1°313 inches.” 

Mr. Tutt also exhibited specimens of Noctua dahlii, and 
read the following notes : 

‘The study of the variation of our Nocrum.is certainly one 
“of the most interesting pursuits of British Entomologists. 
‘‘The specimens exhibited to-night are those of Noctua dahlii, 
‘a species which shows not only a considerable range of 
‘sexual dimorphism, but also of variation according to 
‘‘geographical distribution. Of those exhibited twenty 
‘“specimens were captured in Delamere Forest by Mr. Geo. 
‘Day of Knutsford. ‘These specimens, as will be observed, 
‘¢ show the usual sexual dimorphism which is so conspicuous 
‘in Great Britain. This dimorphism is well marked in the 
‘‘specimens which I have from Aberdeenshire, Yorkshire, 
‘‘ Hgsex, and other localities, the males being of a bright 
‘‘chestnut colour, the females being of a purplish colour 
‘‘with a wide range extending from red with a purple tinge 


(| XXxvill_ ) 


‘‘to purplish black. The males, it will be observed, have a 
‘strong tendency to develop a dark central transverse band 
‘“‘or a dark subterminal band, but very rarely are both 
“present in the same specimens. ‘The distinct sexual 
‘‘dimorphism here exhibited, however, does not occur 
‘throughout the British Isles. At Morpeth the males con- 
‘sist of a mixture of the ordinary chestnut form, together 
‘‘ with purple forms, the latter being coloured like the females 
‘‘which show the usual purple-red range of variation. If, 
‘« therefore, in this species the purple be considered the older 
‘form, it is clear that at Morpeth the evolution of the sexual 
‘colour difference has not yet reached so complete a stage as 
‘‘in many other localities. I exhibit Morpeth specimens show- 
‘‘ing the two male forms and usual female form found there. 

‘In Aberdeenshire, where the species is usually abundant, 
‘the sexual dimorphism is again very marked, the females 
‘‘ varying, as usual, very much in the intensity and depth of 
‘colour. I have, however, out of some 200 specimens 

examined from Aberdeenshire, only seen one male of a 
‘‘ purplish-red tint. This specimen, which I exhibit, has a 
‘‘remarkably pale subterminal band to the anterior wings. I 
‘‘also exhibit two remarkable male aberrations from Aber- 
‘deen, one with a clear central band with scarcely a trace of 
*‘ discoidal spots between a dark basal and a dark subter- 
“minal band; the other deeply suffused with purplish- 
*‘fuscous. There is also among the Delamere Forest speci- 
‘“mens a female with the basal area chestnut coloured, the 
‘‘outer part being very dark purplish. 

‘Four Aberdeenshire females exhibited are the darkest I 
‘‘ have ever seen, and have been selected from a very large 
‘‘number of specimens. One of the Delamere Forest speci- 
‘‘mens is similar. They are, as will be seen, of a fuscous- 
‘“‘black coloration with the normal purple almost obsolete. 
‘‘These dark specimens have, as a rule, very indistinct stig- 
‘‘mata and transverse markings. 

‘Perhaps the most peculiar fact about this species, how- 
‘ever, is, that in Ireland, so far as I know, the males and 
‘females show none of the sexual dimorphism so distinct 
‘‘and prevalent in Great Britain, but both sexes are of the 


C ge J 


‘‘normal purple-red coloration of the female. The common 
‘‘mottled chestnut male either does not occur or exists as a 
‘‘very rare aberration in Ireland, just as in Great Britain 
‘‘purple-red males only occur as a rare aberration. It is this 
‘‘igolation of the purple form in both sexes in Iveland, 
“together with the fact that this form approaches more 
‘“nearly to its nearest generic congeners—stiymatica, brunnea, 
“and ditrapezium —in colour which makes me inclined to 
‘look upon this as the older form, and the bright chestnut 
‘‘males as more recently evolved. 

‘‘As I have already noted, there is a tendency in this 
‘‘ species to show a considerable range in minor aberrational 
“forms. The redder females are the ab. rufa of ‘The 
«British Noctue,’ etc., which is distributed rather commonly 
‘‘among the males and females of the Irish and the females 
‘‘of the English and Scotch specimens. The ab. fusca has 
‘a large quantity of fuscous mixed with the purple colour, 
‘‘ becoming in many specimens almost black, whilst the ab. 
‘* candelisequa of Stephens is a glaucous tinged form of dahlit, 
‘‘yare enough in this species, although glaucous tinted speci- 
‘“mens are quite a feature in the colour aberrations of this 
‘“senus, becoming quite common in VN, festiva, and almost 
“the rule in N. sobrina. A very pale reddish form of this 
‘‘ species was recently exhibited, by Mr. A. F. Bayne, at the 
‘“City of London Entomological Society, and although those 
“of us who know the extent of variation that N. festiva 
‘undergoes in Scotland, will scarcely agree with Guenée that 
“<¢the species varies no less than festiva,’ our recent know- 
“ledge has convinced British lepidopterists that dahlit is an 
‘exceedingly variable species. 

‘‘The triple colour variation of the female dahlii is very 
‘‘interesting —red, purple, and black being the extent of 
‘‘variation through which stiymatica, brunnea, and c-nigrum 
‘extend, whilst in brunnea ochreous mottling in the males is 
‘far from uncommon. This parallel range of colour varia- 
‘tion in closely allied species is proving very general 
‘‘wherever sufficient material, collected over a sufficiently 
‘‘wide area, has been obtained.” 

Herr Jacoby read a letter received from Mr. Buxton 


(my 


Forman, one of the Assistant Secretaries of the Post Office, 
to the effect that the Postal Union had decided to make a 
rule not to admit natural history specimens by sample post, 
which was intended for the transmission of bona fide trade 
patterns or samples of merchandise, and consequently that 
the forwarding of such specimens at the sample rate would in 
future be irregular. 

Lord Walsingham stated that he had had a long corre- 
spondence with the Post Office authorities on the subject, and 
that the late Mr. Raikes, when Postmaster-General, promised 
him in 1891 that such specimens should, so far as the British 
Post Office was concerned, be transmitted at the sample rates ; 
and a letter to the same effect, from the late Sir Arthur 
Blackwood, when Secretary to the Post Office, was published 
in the Proceedings of the Society for 1891. 

Mr. C. G. Barrett exhibited for Mr. A. J. Hodges a speci- 
men of Hydrilla palustris, from Wicken Fen ; also specimens 
of Caradrina ambigua, from the Isle of Wight. He remarked 
that of the latter one specimen had the hind margin of right 
forewing indented, and the wing broadened as though from 
an injury to the pupa. In this wing the margins of the large 
orbicular and reniform stigmata had become so joined that 
the dividing. lines had disappeared, and the stigmata were 
fused into one irregularly formed blotch. The left wing of the 
specimen is normal. The effect of the injury seems to have 
been as though the two stigmata had been in a semi-fluid 
condition and had run together. 

Mr. McLachlan exhibited, on behalf of Mr. G. F. Wilson, 
I'.R.8., of Weybridge, a ‘‘ grease band ” which had been tied 
round trees to prevent the females of Cheimatcbia brumata 
from ascending the trunks for the purposes of oviposition ; 
the band was thickly covered with the bodies of the females, 
together with a few males. 

Surgeon - Captain Manders exhibited a pair of Chelura 
bifasciata, from the Shan States, and called attention to the 
‘Cagssembling’”’ habits of the male, some hundreds of which 
were attracted by the numerous females which emerged from 
the cocoons at sunset. 

Mr. B. A. Bower exhibited a beautiful variety of Zygana 


( xii) 


loniceré, Ksp., having the spots confluent, taken at Chatten- 
den Wood, North Kent, in June last; also a specimen of 
Incurvaria tenuicornis, Stn., taken at Chislehurst, in May, 
1893. 

Mr. H. Goss exhibited, for Mr. F. W. Urich, of Trinidad, a 
series of males, females, and workers of Sericomyrmez opacus, 
Mayr, a species of Fungus-growing and Fungus-eating Ant. 
He said he was indebted to Mr. G. A. J. Rothney for kindly 
mounting the specimens on card. 


Papers Read, 


Colonel Swinhoe read a paper entitled ‘“‘A List of the 
Lepidoptera of the Khasia Hills, Part III.” 

Mr. C. J. Gahan read a paper entitled ‘‘On the Longicorn 
Coleoptera of the West India Islands.’ 

Mr. F. W. Urich communicated a paper entitled ‘‘ Notes 
on the Fungus Growing and Hating Habit of Sericomyrmex 
opacus, Mayr.”’ 

Prof. E. B. Poulton read the following paper, by Prof. E. B. 
Titchener, of Cornell University, U.S.A., entitled ‘‘ An 
apparent case of Sexual Preference in a Male Insect.” 

‘«T have had under observation during the past summer a 
“number of Diapheromera femorata. The insects were all 
‘taken from a single clump of Rubus odoratus, and have been 
‘‘fed upon the leaves of this shrub. Seven were taken in the 
‘second week in July, five females and two males. All but 
‘‘one backward female were in the same stage of develop- 
‘‘ment, having two skin-castings to come before maturity 
‘‘was reached. One of the females had lost her two front 
‘lees, and there was no renewal of these with either of the 
‘‘new skins. The first male to become sexually mature was 
‘small and weak; though he has proved to be exceptionally 
‘long-lived. He ‘chose’ the maimed female, from the group 
‘‘of four. Two days later the other male, which had left 
‘the other females untouched, was found in connection with 
“this same one, having driven his weaker rival away. The 
‘‘yemaining females, with the exception of the immature one, 
‘“‘ were fertilised subsequently. In the last week of August I 


( xiii) 


‘took two more insects, a mature male and female, separ- 
“ately. Although the cage was quite large, there was a good 
‘deal of scrambling and fighting among the males when the 
‘‘newcomers were introduced. The new male, with five 
‘females to choose from, all already fertilised, settled upon 
‘‘the maimed female. A few days later, I found two insects 
‘‘in connection. They were placed in the cage with the rest 
‘‘in this condition. Next morning I saw that the fresh male 
‘had left its female, and was in connection with the maimed 
‘fone. 

‘* Four males in succession, therefore, had ‘ preferred ’ this 
‘mate, out of a possible four, five, or six. Moreover, she 
‘‘was constantly in connection with one of them, while the 
‘“‘rest were often left unsought for some days together. I 
‘thought at one time that the reason of these phenomena 
“‘might lie in a purely mechanical fact: the female in 
‘‘ question, being maimed, could not move so quickly as the 
‘rest, and so could not escape the advances of the male. 
“But I only once saw a male shaken off by a female—the 
‘‘first male mentioned above: and that female was a virgin. 
‘* When once fertilisation by one male had taken place, the 
‘‘ female remained passive at the approach of others. When 
‘‘the third and fourth males were introduced, all the avail- 
‘able females had had connection. The experiment is not, 
“of course, ‘pure;’ no experiment made in captivity is. 
‘« Still, the insects took to their confinement very well; feeding 
‘‘heartily and breeding readily. I record the facts for what 
‘they were worth, and shall be glad to answer any question 
‘“as to possible sources of error that may occur to the readers 
‘‘of them. Another watcher beside myself has had the 
‘‘insects under pretty constant observation, and can confirm 
‘the above account.”’ 

The President and Prof. Poulton made some remarks on 
the observations recorded in Prof. Titchener’s paper. 

The Rey. H. 8. Gorham communicated a paper entitled 
‘* Notes on Herr A. Kuwert’s * Revision der Cleridengattung 
** Omadius, Lap.” 


* Annales dela Société entomologique de Belgique. Tome Trente-huitiéme, 
lii., pp. 62-97, 1894. 


( xliii_ ) 


“ Herr A. Kuwert in giving analytical tables and descrip- 
‘‘tions of 44 species, presumed to be new, of this most 
“difficult genus, labours under the disadvantage of not 
‘having consulted the records of previous workers in the 
‘family of the Cleride., 

‘‘Had he done so he could hardly have failed to discover 
‘that I published in 1876 (Cistula Entomologica, p. 57), 
““¢ Notes on the Coleopterous family Cleride,’ with critical 
‘remarks, at pages 92 and 101, on this genus and on Stig- 
‘“matium, with descriptions of eleven new species of Omadius, 
‘and many of Stiymatium, and with some synonymy founded 
‘‘on the examination of typical specimens from the collec- 
‘tions of Guérin Meneville, Chevrolat, W. W. Saunders, 
‘‘ Wallace, Semper, and others mentioned at p. 58 of the 
‘‘ work mentioned. 

“These will all be found recapitulated in the ‘ Zoological 
‘¢ Record,’ for 1876, at pp. G0—62. 

“Tf Herr Kuwert had done thus, and had he, also, looked 
‘Cup his references, he would not have fallen into the error 
‘‘of referring the type of the genus O. indicus to Spinola, and 
‘‘of substituting for that name prolicus, Klug, whereas it 
‘“«(O. indicus) is referable to Laporte, and was published in 
‘‘Silberman’s Revue (1833—39). Nor would he refer 
‘««Chevrolat’s species to Guérin,’ they having been expressly 
‘‘ published in M. A. Chevrolat’s name. 

‘One species of mine, O. notatus, has apparently somehow 
‘‘come under Herr Kuwert’s notice. It is briefly placed as a 
‘synonym of kamelianus, White, thus (? notatus, Gerb.) sic. 
‘‘ Beyond the fact that both are described as from the Philip- 
“pine Islands, no reason is given for thus conjoining 
‘«¢ them. 

‘Some names such as Cramensis for an insect for Ceram, 
‘‘ which is corrected in the description, may be overlooked as 
‘‘ misprints, and it is a little remarkable that Herr Kuwert 
‘‘has escaped colliding with any of my names, either in 
‘‘ this genus or in Stigmatium ; although ‘ angustifrons,’ No. 26, 
“ig very suggestive of some relation with ‘angusticeps,’ Gorh., 
‘one being from Sumatra, the other from Borneo. 

‘© O. nigropunctatus, Cheyr., has been, by comparison of 


( *xlivel 4) 


“types and otherwise, identified by me with O. medio- 
“ fasciatus, Westwd., and is so beyond all doubt, being a very 
‘‘ easily identified species. 

‘The redescription of a species from Perak as O.javanus is 
‘very futile, that name having been used by Dejean and 
‘‘ placed as a synonym of O. indicus in Gemm. and Harold 
‘Catalogue, teste Spinola. 

‘‘The finest and most conspicuous species, O. prioceroides, 
‘“Thoms., escapes notice altogether. 

‘“‘ Synthesis, we may remind Herr A. Kuwert, is an impor- 
‘‘tant part, nay, essential to revision. 


‘* Revision des genus STiGMATIUM. 


‘Under this title, in the ‘Annales de la Société Ento- 
‘“‘mologique de Belgique,’ Tome xxxviii., Pt. viii., pp. 398— 
‘“457, Herr A. Kuwert has published an analytical and 
‘‘descriptive paper of such species of the genus Stigmatium, 
‘“‘Gray, as were known to him. These are, apparently, con- 
‘tained: (1) in his own collection; (2) in that of the Honble. 
‘Walter de Rothschild, Tring Park (England); (3) in the 
‘“Museum at Berlin; (4) in the Museum at Vienna. Of 
‘however, the majority of species, either newly described or 
‘identified with described species, no clue is given as to how 
‘they have been identified, or by whom collected. Herr 
‘« Kuwert has made no attempt to identify the many species 
‘‘of this genus described by M. A. Chevrolat and myself 
‘‘ during the past eighteen years, although we have been well 
‘‘known to have paid special attention to the family Cleride. 
‘In 1876, Chevrolat published a very considerable contri- 
‘‘bution to the knowledge of this group, entitled ‘ Mémoire 
‘‘sur la famille des Clerites,’ in this paper ten species of 
“* Stigymatium from the collections of Boucard, Haag de Ruten- 
‘berg, Sahlberg, Sallé, etc., are described. In the same 
‘“‘ year, after communication with M. Chevrolat, I described in 
“«¢ Cistula Ent.,’ no less than 22 species with one new allied 
‘genus, ‘ Hemitrachys.’ Since then I have printed many 
‘papers on Cleride, in which reference to these species and 
‘‘some additions appear. The species thus described formed 
‘part of the world-known collections of Bates and Wallace ; 


calves} 


‘and most of the important collections, both of this country 
‘and of France, Holland, Belgium, and Italy, have passed 
‘through my hands, and many of the types are in my own 
‘collection. Herr Kuwert, however, passes the whole of 
‘this over in silence. I may add that several of his species 
“appear at first sight to be identical with my own. 

‘“The species of this genus are more difficult and obscure 
“than perhaps any other genus of the family, perhaps as 
‘“much so as those of any genus of Coleoptera. 

‘“The student now will find himself in a hopeless con- 
‘fusion, from which nothing but the collation and close 
‘examination of the types of my own, and the scattered 
‘types of Herr Kuwert’s descriptions can extricate him. 

‘‘The types, however, of all my species are in my posses- 
‘“ sion, and are, of course, accessible to any one coming here to 
“see them.”’ 


ANNUAL MEETING. 
January 16th, 1895. 


Henry Joun Ewes, Esq., F.L.8S., President, in the chair. 


Mr, W. F. H. Blandford, one of the Auditors, read the 
Treasurer’s Balance Sheet, showing a balance in the Society’s 
favour of £29 2s. 7d. 

Mr. H. Goss, one of the Secretaries, read the following :— 


Report of the Council. 


During the Session 1894-95, one Honorary Fellow, Pastor 
Wallengren, and four ordinary Fellows have died, viz., 
Major-Gen. George Carden, the Rey. Alfred Forbes Sealy, Mr. 
John Jenner Weir, F.L.8., and Dr. F. Buchanan White, M.D.., 
¥.L.8.; eight Fellows have resigned; three have been 
restored to the list; and twenty-six new Fellows have been 
elected. 


( (sive) 


The number of Fellows elected during the year is slightly 
above the average. At the same time it is desirable to in- 
crease our numbers more rapidly, so as to enable us to pub- 
lish more papers, and allow more plates, and in other ways 
to advance our interests and promote our objects. The 
Council, therefore, again appeal to the Fellows to do their 
utmost to induce their friends to join the Society and thus 
increase its revenue. 

At the present time the Society consists of 9 Honorary, 
49 Life, and 824 Fellows paying the Annual Subscription, 
making the total number of Fellows now on our List 382, 
which, after allowing for the losses by deaths and resigna- 
tions, is an increase of 16 since the Annual Meeting last 
year. 

Our Transactions for the year 1894 form a volume of 586 
pages, containing 25 Memoirs contributed by the following 
authors, viz., Mr. Edward Meyrick, B.A. (2 papers); Mr. Charles 
Owen Waterhouse; Mr. George T. Bethune-Baker (2 papers); 
Mr. Walter F. H. Blandford, M.A. (2 papers); Mr. Hamilton 
H. Druce ; Colonel Charles Swinhoe, M.A.; Mr. George C. 
Champion (2 papers); Mr. Edward Saunders; Dr. Frederick 
A. Dixey, M.A., M.D. (2 papers); Dr. Thomas A. Chapman, 
M.D.; Mr. Charles B. Taylor; Mr. William F. Kirby; the 
Rev. Canon Fowler, M.A.; Mr. Frederic Merrifield; Mons. 
Louis Péringuey; Mr. Charles J. Gahan, M.A.; Professor 
Franz Klapalek ; the Rev. Thomas A. Marshall, M.A.; the 
Right Hon. Lord Walsingham, LL.D., F.R.S.; and Mr. 
Kenneth J. Morton. Of these 25 papers 12 relate to Lepido- 
ptera (or to enquiries in which they were the subjects of 
experiment), 7 to Coleoptera, 2 to Hemiptera, 2 to Neuro- 
ptera, 1 to Hymenoptera, and 1 to an entomological excur- 
sion to Corsica. The Memoirs above referred to are illustrated 
by 14 plates, of which 5 are coloured. The Society is in- 
debted to Mr. G. T. Bethune-Baker for half the cost of Plate L, 
to Colonel Swinhoe for part of the cost of Plate IL., and to 
Mr. F. Merrifield for the entire cost of Plate IX. 

The Proceedings, containing an account of the Exhibitions 
and Discussions at the Meetings, in addition to several papers 
not published in the Transactions, and to several Notes of 


( ‘xdvii.°) 
interest in connection with Exhibitions, extend to 48 
pages. 

During the past year about 250 Books, Pamphlets, 
Journals, and Papers have been added to the Library ; and 
the Meetings have been better attended than in any previous 
year. 

Another matter for congratulation is the increased use made 
of the Library by the Fellows of the Society, which is shown 
by the following figures, furnished me by our Resident Libra- 
rian, Mr. W. R. Hall. In 1889 the number of books borrowed 
or referred to was 95. In 1890 this number was raised to 
108, and in 1891 to 170. During the past year the number 
has been increased to 209! The figures speak for themselves, 
and show that the value and advantages of the Library are 
fully appreciated. 

The Balance Sheet which you have just heard read, and 
which is appended to this Report, includes, under the head of 
Donations, one of £45 from our generous benefactor Mr. 
J. W. Dunning, on the anniversary of the 45th year of his 
election as a Member, with a suggestion that it might be 
applied partly to general purposes and partly to additions 
to the Library, which suggestion has been acted upon. 

The Subscriptions received for the year 1894 amount to 
£309 15s., asum in excess of that for any former year. 

Subscriptions in arrear were paid up to an extent exceed- 
ing expectations ; but, on the other hand, the amount still 
due for 1894 is much in excess of that which is usual at the 
end of a year. 

The sales of our Transactions have been satisfactory, and 
show a notable increase on those of the years immediately 
preceding. 

In February last the Misses Swann, nieces of our lamented 
Honorary Life President (the late Professor Westwood), 
handed over the ‘‘ Westwood Bequest”’’ of £250, and it has 
been invested in the names of Trustees in Birmingham Cor- 
poration 8 per cent. Stock. 

Three Life Compositions haye been received during the year, 
and invested in Consols. The total sum now so invested 
being £455 18s., representing £478 11s, stock. 


( xni ) 


Commencing the year with a balance in hand of £10 6s. 1d., 
this was increased to £29 2s. 7d. on the 31st December, with 
no outstanding liabilities; and after carrying forward £7 7s. 
for subscriptions received in advance to the credit of 1895, a 
result that cannot be considered other than satisfactory. 


11, Chandos Street, Cavendish Square, W. 
January 16th, 1895. 


The Secretaries not having received any notices of objec- 
tion, the following Fellows of the Society were declared duly 
elected members of the Council for the Session 1895-1896 :— 
Mr. George T. Bethune-Baker, F.L.8.; Mr. Walter F. H. 
Blandford, M.A., F.Z.S.; Mr. George C. Champion, F.Z.8. ; 
Dr. Frederick A. Dixey, M.A.; Mr. Henry John Elwes, F.L.S. ; 
Canon Fowler, M.A., F.L.S.; Mr. Charles J. Gahan, M.A.; 
Mr. Herbert Goss, F.L.S.; Mr. Robert McLachlan, F.R.S.; 
Professor Raphael Meldola, F.R.S.; Professor Edward B. 
Poulton, M.A., F.R.S.; Dr. David Sharp, M.A., F.R.S.; and 
the Rt. Hon. Lord Walsingham, M.A., LL.D., F.B.S. 

The following were the Officers elected :—President, Pro- 
fessor Raphael Meldola; Treasurer, Mr. Robert McLachlan ; 
Secretaries, Mr. Herbert Goss and the Rev. Canon Fowler ; 
Librarian, Mr. George C. Champion. 

Mr. Elwes then read his Address, at the conclusion of 
which Professor Meldola moved a vote of thanks to Mr. Elwes 
for his Address, and for his services as President during 
the past year, and he also moved a vote of thanks to the 
Treasurer, Secretaries, and Librarian for their services during 
the past year. This was seconded by Mr. F. Du Cane Godman, 
and carried unanimously. Mr. Elwes, Mr. McLachlan, Mr. 
Goss, and Canon Fowler made some remarks in acknowledg- 
ment. 


¢ xlixe ) 


ENTOMOLOGIAL SOCIETY OF LONDON. 
Balance Sheet for the Year 1894. 


RECEIPTS. PAYMENTS. 
d. a5 & Bl 
Balance in hand, Jan. Ist, Printing Transactions, &c. 216 13 6 
1894 . . . + 10 6 1 | Plates, &. 69 16 0 
Subscriptions for 1894 . 30915 0 | Rent and Office Ex- 
Arrears 1818 0 | penses - ; F Leos 
Admission Fees 48 6 0 | Books and Binding . By th 
Donations 57 4 6 Investments :— 
Westwood Bequest . 7 200) 00 Consols . £47 5 0 
Sale of Transactions, &e. 77 5 6 Westwood Bequest 250 0 0 
Interest on Investments :— A 
Consols £11 8 2 au ee te es 297 5 0 
ubscriptions in advance 
Westwood Bequest 3 9 a a Bee ATEG ASOE: (a) eat Ae 
Life-Compositions . 47 5 © | Balance 292 7 
Subscriptions in advance 7 7 0 
£841 4 9 £841 4 9 
ASSETS. 


Subscriptions in arrear. (considered good) £16 16s. 


Investments :— 


Od, 


Cost of £473 11s. 0d. Consols = £455 18s. Od. 
Cost of £239 12s. 4d. Birmingham Corporation 3 per cent. 
Stock (Westwood Bequest) £250. 


LEAB TI TT Hs: 
(Nil.) 


Ropert McLAcuHian, 


Treasurer. 


Examined and found correct, 


9th January, 1895. 


SAMUEL STEVENS. 
A. Hueu JoNnEs. 


Watter F. H. BLANDFORD. 


CHARLES J. GAHAN. 
OsBERT SALV:N, 


Oe see 


ADDRESS, 


ETC,, ETC, 


GENTLEMEN, 

The time has now arrived when, in accordance with 
the rules of this Society, I have to resign the Presidency into 
other hands, and I do so with less regret, because, as I said 
in my address last year, I find that the numerous duties 
and pleasures of life, and the greatly increased amount of 
business which my connection with agriculture at present 
entails, leave me too little time to attend to the work of 
the Society ; and because I know that I am resigning it into 
the hands of a gentleman who will, as a Fellow of the Royal 
Society, resident in London, be able to do much more to 
advance the interests of the Society than I can do. 

Iam happy to think that, as I leave the chair, the Society 
is more numerous, more active, and richer than when I 
took it, and hope that it will continue to make as satisfactory 
progress in numbers and ability as it has done for some years 
past.* 

No event of great importance has, so far as I know, taken 
place in the entomological world during the past year. The 
number of memoirs and papers registered by the indefatig- 
able labours of our colleague, Dr. Sharp, in the Zoological 
Record, is no less than 1069 against 1026 in the previous 
year, and I fear that the difficulty of keeping pace with, and 
studying this enormous mass of material, to which I made 


* The increase in the number of Fellows is from 254 in 1884 to 383 in 1894. 


(Glas) 
special allusion in my address last year, continues to in- 
crease. 

We have not lost by death during 1894 either so many or 
such distinguished men as those whose names I had to 
mention last year. Of our Fellows, the oldest and best known 
is Jonn JENNER WEIR, F'.L.S., etc., who died at Beckenham 
in August. He had been a Fellow since 1845, and was a 
most regular attendant at our meetings during nearly fifty 
years. He was often a Member of Council, Treasurer from 
1876 to 1879, and vice-President in 1886. Though not a pro- 
lifie writer, he had a large general acquaintance with British 
and Exotic Lepidoptera, as well as a good knowledge of 
ornithology and botany. Of late years he had formed a fair 
general collection of butterflies, which was dispersed at Stevens’ 
rooms soon after his death. He was a most amiable and 
popular man, who will be much missed at our meetings. 

Major-General Grorcr Carpen, F.E.5., died at Bromley on 
Feb. 12, aged 56, from the effects of influenza. He had 
served with the 77th regiment in the Crimean War, and with 
the Northumberland Fusiliers during the Indian Mutiny, 
and commanded this regiment for some years. Though not 
a scientific entomologist, he was a close observer, and an 
active collector of British Lepidoptera. 

Francis Bucuanan Wuite, M.D., died at Perth, on Dec. 8rd, 
aged 52, having been a Fellow of our Society since 1868. 
He devoted a great part of his life to the study of natural 
history, especially in the Highlands of Scotland; and though 
perhaps more a botanist than an entomologist, had con- 
tributed many papers on his discoveries to the ‘ Kntomo- 
logist’s Monthly Magazine’ and other periodicals. Perhaps 
his best known and most important works were his paper ‘ On 
the Male Genital Armature in the European Rhopalocera,’ 
published in the ‘Transactions of the Linnean Society,’ and 
a memoir on the Pelagic Hemiptera of the Challenger Expe- 
dition. He was for some years President of the Perthshire 
Society of Natural Science, and editor of its Proceedings, and, 
though not so well known in England, was a leading man 
among Scottish naturalists. 

Dr. Hearper of Carmarthen, and Winuram Macuin of 

ul 


(li } 


London, were also known as at one time collectors of British 
Lepidoptera, but Iam not aware that either has written any- 
thing of permanent interest. ' 

Among foreign entomologists who have died in 1894, I find 
but very few names of note. The most eminent perhaps were 
Lucien Francois Leruterry, who died at Lille on April 4th, 
aged 64. Together with Dr. A. Puron, he probably held the 
first place among Hemipterists, and had commenced to pub- 
lish a general Catalogue of the Hemiptera of the world, 
of which the first part appeared in 1893. He had been a 
member of the Entomological Society of France since 1857. 

Dr. Jacop Spinepera was a well-known Swedish entomo- 
logist, who had written on Hemiptera, and published a mono- 
graph of the Scandinavian Psocida, and some papers on 
Lepidoptera as well. He was a leading member of the Swedish 
Entomological Nociety. 

Hueco Tueopor Currstopn died at St. Petersburg on 
Oct. 24, aged 63. He was one of the most energetic collec- 
tors of insects in Europe, and had travelled extensively for 
many years in various parts of the Russian Empire, especially 
in Amurland, North Persia, Transcaspia, and the Caucasus. . 
A Saxon by birth, he went to Sarepta as a teacher in 1858, 
and at once began to make his name well known among 
Kuropean lepidopterists. Of late years he had been employed 
by the Grand Duke Nikolas Mikhailovitch as curator of his 
collections, and had resided at St. Petersburg, except during 
the summer months, which he usually spent in the Caucasus. 
He had written numerous papers on the results of his expedi- 
tions, and described many new species of Lepidoptera. He 
will be a great loss to science, as there was probably no one 
better acquainted with the Lepidoptera of the Russian Empire. 


The subject which I have chosen for my address is one 
in which for many years I have taken great interest, 
namely, the Geographical Distribution of Butterflies; for 
though a great deal has been written of late years on 
the distribution of plants, mammals, birds, fishes, and 
reptiles, comparatively little has yet been done, so far as I 
know, by entomologists to show how far the natural divisions 


(a bad 8) 
of the earth’s surface which have been established for other 
classes are applicable to insects. I think that the pro- 
portion of known as compared with unknown insects is still 
too small, and the classification of the known species still too 
uncertain, to allow the same methods to be applied to them 
that have been used for mammals by Wallace, for birds 
by Sclater and Sharpe, and for plants by Hooker, Dyer, and 
Hemsley; but still I think we might do more than we have 
done in this branch of our subject. 

Personally, I believe that it is still too soon to investi- 
gate, with much hope of success, the reasons for a thousand 
curious and apparently inexplicable facts which turn up in 
the study of distribution, and of which, I think, the speci- 
mens I am showing to-night, in illustration of these remarks, 
afford abundant proof; but I hope to draw the attention of 
our Fellows, and entomologists generally, to the backward 
position we occupy in this branch of zoological study. 

No one man has, or could have, sufficient knowledge of all 
orders to work out a scheme of geographical distribution for 
insects generally ; but we might individually do a good deal to 
throw light on the subject, in order to see how far the facts, 
as far as we are able to judge, agree with those derived from 
the study of plants, mammals, and birds. 

T hope eventually to review the subject more completely ; but 
in writing of the regions where I have personally travelled and 
collected, and of whose butterflies I have a fair knowledge, 
namely, Europe, Asia, and North America, I find so many 
doubtful and difficult questions, that I thought it best not to 
attempt more than a slight sketch of the affinities of regions 
like South America, Africa, and Australia, whose butterflies 
I know but very slightly, from the writings and observations 
of others; and shall, therefore, touch but very lightly upon 
them at present. 

In attempting to compare the distribution of insects with 
that of birds, animals, or plants, we are met by two great 
difficulties, which must make any exact definition of the 
regions and sub-regions impossible at present. One is the 
want of knowledge as to the insect fauna of many of the most 
important areas of which the animals, birds, and plants are 


(diy) 

fairly well-known. The second is the uncertainty of the 
elassification which at present exists in many of the most 
important groups. It would seem, as far as one can judge 
from what we do know, that the families and genera of 
butterflies are more cosmopolitan in their distribution than 
those of either animals or birds; but this may be partly owing 
to the fact that the recognised genera are not so much sub- 
divided. 

And on the question of sub-division of genera I must say a 
few words. Since the publication of Westwood and Double- 
day’s work, many years ago, there has been no attempt made, 
so far as I know, to revise the genera of butterflies except 
in Schiitz’s (continued by Rober) ‘ Familien und Gattungen der 
Tagfalter,’ published in 1892, as a volume of the ‘ Exotische 
Schmetterlinge ’ of Staudinger and Schiitz. 

This is a careful and fairly complete work, based on an 
analytical study of the rich collection of Dr. O. Staudinger, 
but the genera adopted are so large in many cases, that some 
extent of sub-division seems necessary for purposes of con- 
venience. The very numerous sub-genera proposed, though 
insufficiently characterised, by Moore, Butler, and Scudder, 
but in many cases not accepted by other lepidopterists, are 
not sufficiently criticised by Schiitz, and genera such as Saty- 
rus, containing many divergent species, are adopted en bloc. 
Some of our best systematists, of whom Messrs. Godman 
and Salvin are the leaders, consider that it is impossible to 
sub-divide large cosmopolitan genera such as Thecla in a 
satisfactory way when dealing with local faunas, and in this 
I fully agree with them. We have a good recent instance 
of the extent to which the contrary system may be carried 
in Scudder’s ‘ Butterflies of the Eastern United States and 
Canada.’ In his preface, p. ix., Mr. Scudder attempts to 
justify this practice by saying, “if the characters I 
‘‘ have pointed out, as pertaining to such groups and their 
‘‘ relations to those placed above and below them, are not 
‘‘in themselves a justification, then I have none, and no- 
‘‘ words of mine could or should alter such a fact ; if, how- 
‘‘ever, these characteristics represent actual categories, 
‘“‘andif, at the same time, such groupings make clearer the 


Clg) 


‘relations which the life histories and the distribution of 
‘‘ the groups bear to their structure, then words are not 
*“ needed.” 

Now let us apply these excellent principles to the way in 
which Mr. Scudder has acted. 

He allows Gineis maccount to remain in the same genus 
with (!. semidea, though there are not perhaps in the whole 
genus two species more divergent in their general appearance, 
habits, and distribution than these ; maccouni representing a 
group characteristic of the Coniferous Forest of the Pacific 
coast, the other being typical of the most Arctic form of the 
genus, found on high mountain summits and ice-swept coasts. 
On the other hand, he makes out of the twelve species of 
Thecla found in his region six genera, of which four are said 
to be restricted to America, though it is evident that the 
author has little acquaintance with the genera of the 
Neotropical region. Amongst them IJncisalia is said to be 
confined to the United States, but closely allied to the 
European Callophrys, Billberg, of which he gives* our Thecla 
rubi as type; but does not mention Thecla dumetorum from 
California, which is so close to rubi that I cannot distin- 
guish them without the labels, and I find much variation in 
the characters by which Jncisalia is defined. Thus I am 
justified in saying that until Scudder shows us how to 
distinguish rubi from dumetorum and Callophrys from In- 
cisalia, the latter genus is not at present to be relied on 
for the study of distribution. 

Moore carries the same practice as far as Scudder, and 
though he attempts by analytical tables to characterise his 
so-called genera, they cannot be accepted on his authority 
alone, though, no doubt, some of them are based on good 
characters. In dealing with the Indian Satyrine, he has 
apparently, without reference to their allies in other parts of 
Asia, Africa, and Europe, sub-divided Ypthima, Satyrus, and 
others into many so-called genera bearing names such as 
Chazara, Crebeta, Hemadara, Kanetisa, Kolasa, Karanasa, 
Thymipa, etc., which being without meaning or sense, are 
very hard to remember. 


* Scudder, Gen. Butt. in Proc. Amer. Acad. Sci. Boston, 1875, p. 182. _ 


Cae”) 


Tf I adopted them and based a study of the distribution of 
Indian Satyrinw on them, I should probably arrive at grossly 
distorted and unreliable results. 

This being the case in regions which have been well worked, 
and of which the butterflies are fairly known, how much 
more difficult it is to decide as to regions which, like many 
parts of Africa, are almost unknown. I cannot dismiss this 
part of the subject without referring to some remarks by 
Canon Tristram, F'.R.§., in the Ibis for January, 1895, p. 180. 
He says, 

‘‘The question is:—is the multiplication of genera, each 
‘¢ containing one or two species and those closely allied, an aid 
‘‘or a hindrance to the study of the subject? To quote the 
‘words of Dr. Bowdler Sharpe on another point in nomen- 
‘‘clature, and which I would apply to many of the new- 
‘‘fangled genera, ‘An arrangement we shall never adopt, 
‘«¢ag we consider it a clumsy and unnecessary method of 
“ «nomenclature, and one that in the hands of unscrupulous 
‘‘¢ writers may be employed ad lib. to gain a little tem- 
‘« «porary notoriety and end in making the study of birds 
‘« ‘impossible. Can any science bear the weight of such 
‘« ¢a system of nomenclature?’ ”’ 

I say decidedly, No. 

The Rhopalocera are perhaps the best adapted for analysis, 
as being better known and less numerous than Coleoptera or 
Heterocera, but as soon as you begin to compare lists of butter- 
flies from different regions and sub-regions with the object of 
‘finding out the proportion of peculiar genera and _ species 
characteristic of them, you are at once involved in a host of 
minor difficulties arising from the two causes above men- 
tioned. And though one is able to give the general features of 
the butterfly fauna of many of the sub-regions, one cannot 
be sure how far their relative degree of specialization may 
not have to be modified by future discoveries and _ better 
systems of classification based on larger materials for study. 

Shortly after the publication of Dr. Sclater’s classical 
paper on the Distribution of Birds, Mr. W. F. Kirby pub- 
lished, in the Journal of the Linnean Society, Zool., 
Vol. XL, p. 481, 1878, a short paper on the Distribution 


( ivity 

of Butterflies, which was, as far as it went, quite in accord- 
ance with the facts then known, but is now somewhat out of 
date; and, moreover, errs in this respect, that many of the 
genera mentioned as characteristic are, from my point of 
view, not truly so. For I cannot regard as characteristic 
forms, genera which are not present throughout the greater 
part of the regions which they belong to, or which are found 
in equal abundance in other regions; but rather I consider 
those as characteristic which are dominant in number of species, 
found almost everywhere in these regions, and not in others, 
except in isolated cases or as stragglers. 

Dr. Adalbert Seitz has written a long and interesting paper 
in the Zoologischer Jahrbucher of Jena, vol. v., pp. 281— 
348, on the ‘Geographical Distribution of Butterflies and 
its Dependence on Climatic Influences,’ in which he at- 
tempts, inter alia, to show that the influence of trade winds 
has largely contributed to the abundance and variety of 
Lepidoptera in various parts of the world, and, so far as 
trade winds have affected the abundance or scarcity of vege- 
tation, there is no doubt some truth in this; but this 
theory is utterly insufficient to account for many of the most 
remarkable features we meet with; and though it is easy 
to find facts to prove almost any theory in such investigations 
as this, yet Dr. Seitz’s knowledge of many of the faunas 
which he mentions in proof of his theory is evidently very 
slight. He attributes to the influence of trade winds a fact 
which is only partially true, viz., that islands lying in the path 
of trade winds eastward of continents, such as Cuba, Mada- 
gascar, the Philippines, Borneo, and Formosa, are all very 
rich in insects, whilst islands lying to the west of conti- 
nents out of the course of a trade wind, are usually poor in 
insects. 

He ignores the difference between oceanic islands sur- 
rounded by deep seas which have probably been long isolated, 
and continental islands surrounded by shallow seas which in 
former ages were probably connected with continents. 

I am myself disposed to consider that the influences which 
contribute principally to the development of a large number 
of species and individuals of Rhopalocera, are largely climatic. 


( apiiy » 

In temperate and cold regions, the prevalence of regular 
seasons having a warm and sunny summer, with a dry and 
cold winter, and in tropical regions abundant rainfall, coupled 
with sufficient sunshine, being the conditions which favour 
abundance of individuals, A great range of elevation in a 
limited area has an overwhelming influence on the number 
and variety of genera, 

One would reasonably suppose that as the greater number 
of Lepidoptera depend on plants alone during their laryal 
stage of existence, their distribution would closely coincide 
with that of plants. So far as I am able to judge, how- 
ever, the general features of their distribution agree far 
better with that of birds than with that of plants. 

The best general reswmé of the distribution of the plants 
I know is that by Mr. W. B. Hemsley, F.R.S., in his intro- 
duction to the botanical part of ‘ Biologia Centrali-Americana,’ 
published in 1888; and I am glad to see that in this 
admirable paper the same principle of working out the facts 
is adopted that I had previously used in 1873 in my paper 
on the ‘ Distribution of Asiatic Birds,’** namely, by per- 
centage. Wallace states that nothing like a_ perfect 
Zoological division of the earth is possible, and in this I 
quite agree with him. But when Wallace, Sclater, and 
Sharpe are so nearly agreed, the main difference between 
them being that Sclater makes the Pacific region separate 
from the Australian, whilst Wallace and Sharpe unite them 
as a primary division, it is clear that there cannot be any 
great error in their conclusions. 

Hemsley suggests two alternative schemes, in one of which 
he admits as primary botanical regions the following :— 


1. Northern, corresponding to the Palearctic and Nearctic 
regions of Wallace, Sclater, and Sharpe. 

2. African, corresponding to their Ethiopian region. 

3. Indian, corresponding to their Indian region, with the 
addition of the Austro-Malay sub-region of Wallace. 


* Proc. Zool. Soc , 1873, pp. 645—682. 


G chi >) 


4. South American, corresponding with the Neotropical 
region of Zoologists. 

5. Australasian, corresponding with the Australian 
region of Wallace and Sharpe, less the Austro-Malay 
sub-region, and with the Pacific and Australian region 
of Sclater, less the same sub-region. 

Hemsley’s alternative primary division, which he says is 

more in accordance with the writings of many botanists, is as 
follows :— 


. Northern Region. 
. Neotropical. 

. Paleotropical. 

. Andean. 

. Cape. 

» Australasian. 


om Ot Hm OO bo Re 


In both his divisions he omits two small but exceptionally 
interesting floras, namely, those of the Sandwich Islands 
and Antarctic regions, which though not large enough to be 
treated as primary regions, cannot be consistently included in 
either of the others. 

Sir Joseph D. Hooker in reviewing Hemsley’s arrangement 
on pages lxy.-lxviii. of the same work, states that though he 
disputes neither his facts or his methods, yet he differs from 
him to some extent in his limitation of the primary floras 
of the globe, and divides the world into two primary regions 
or Botanical Empires, Tropical and Temperate, which he 
again divides into the following regions or Botanic King- 
doms :— 


1. The North Temperate region of the Old World. 
This coincides with the Palearctic region of Sclater, 
except that it includes Greenland. 

. The North Temperate region of the New World. 

. The Tropical region of the Old World. 

. The Tropical region of the New World, 

. The South Temperate region of America, 

. The South Temperate region of Africa. 

. The South Temperate region of Australia. 


co bo 


10 OF ee 


( Ix) 


Prof. Thistleton Dyer, F.R.S.,in an admirable lecture on 
‘Plant Distribution as a Field for Geographical Research,’* 
admits three primary floras :— 


1st. The Northern Flora, practically the same as Hemsley’s 
northern region, which he divides into— 
la. The Arctic Alpine. 
1b. The Temperate. 
1c. The Mediterraneo-Caucasian. 
2nd. The Southern Flora, divided into— 
2a. The Australian. 
2b. The South African. 
2c. The Temperate South American. 
2d. The Antarctic Alpine. 
8rd. The Tropical Flora, divided into — 
8a. The Asiatic. 
3b. The American. 
8c. The African. 


There is really no great difference between the views of these 
three most competent authorities, except in the value that is 
attached by them to primary and secondary divisions. 
Hooker, it is true, whose personal knowledge, gained by travel 
in many of these regions, which alone will enable ‘a man 
to appreciate the physical features of the earth’s divisions in 
the highest degree, seems to consider the north temperate 
regions of the old and new world as divisions of primary 
importance, and in this agrees with most zoologists: whilst 
Dyer and Hemsley agree in uniting them. But, judging 
from the study of butterflies alone, I say without hesitation 
that the Nearctic and Palearctic regions are inseparable, and 
that the Nearctic region has fewer peculiar endemic genera 
and species than the Mediterranean-Asiatic, or, as I call it, 
_ Mediterraneo-Persic sub-region ; and, in consequence, I believe 
that the Nearctic or North Temperate region of the new world, 
is at best a sub-region or province of the great Temperate or 
Palearctic region. It is true that a number of species and 
some genera occur all over the United States, which are not 
Palearctic, and that in the Gulf States, New Mexico, 


* Proc. Royal Geogr. Soc., vol. xxii, 1878. 


( ia’) 


Arizona, and Southern California these may give an abso- 
lutely un-European appearance to the lepidopterous fauna ; 
but when you come to examine these elements you will find 
nothing, or almost nothing, of an endemic or peculiar 
character amongst them. 

What you do find are— 

1st. A number of stragglers or immigrants from the Neo- 
tropical region, belonging to cosmotropical genera, such as 
Papilio, Callidryas, Terias, Danais, Junonia, and Thecla. 

2nd. A few purely Neotropical butterflies, such as Heliconia 
charitonia (in the Gulf States only). Leptalis, one species ; 
Meganostoma, two species ; four or five species of Hrycinide ; 
one species of Eurema; one of Paphia, and a few others 
almost entirely confined to the borderland of Texas and 
New Mexico, or to Florida, where the vegetation is sub- 
tropical. 

8rd. Several species of Phyciodes and Neonympha, which, 
though Neotropical rather than European, have extended 
their range throughout a great part of the United States, but 
of which Phyciodes only is a constantly-present and dominant 
genus. 

4th. Three species belonging to monotypic genera confined 
to very limited areas in the United States, viz., Meniseca 
Tarquinius, which occurs in various spots east of the Rocky 
Mountains, from Maine to Missouri and Florida, but usually 
a scarce or local species; a butterfly of very obscure alliance, 
but having a marked superficial resemblance in form and 
colour to the curious Malayan Liphyra brassolis, with which 
also if agrees in the aphidivorus habit of its larva. 
Neophasia menapia, a monotypic Pierid confined to the 
pine forests of the Northern Pacific States, and Neominois 
Ridingsi, which is found on the western slopes of the Rocky 
Mountains. 

If we adopt Scudder’s genera, this number would be 
increased, but I have shown reasons for not doing so, until 
they are better established by a study of the allied forms in 
other regions. 

Among the Hesperiida, it is true, the apparently Neotropical 
element in places overwhelms the Palearctic forms; but in 


( xii) 


this family the classification is not sufficiently certain to 
enable us to analyse the generic distribution, though Watson’s 
‘Revision of the Hesperiida,’* based on a somewhat hasty 
study of the species in the British Museum, shows that the 
majority of the American genera, and some of the subfamilies, 
are confined to the Neotropical region. 

Among the Heterocera the Neotropical element appears to 
be greater than among the Rhopalocera; but here again we 
have not sufficient knowledge. Packard, in his Monograph of 
the Geometrid Moths of the United States,+ gives a good 
account of their distribution (pp. 567, et seq.), with lists of the 
species inhabiting the east and west of the United States of 
America; but he does not compare these lists with those of 
Europe, and the European genera he mentions as not found 
in the United States, are probably subject to much modifi- 
cation at the present time. 

He states, however, two facts, which I can abundantly con- 
firm from my personal knowledge of the Western butterflies, 
namely, the absence on the Pacific slope of forms character- 
istic of Japan and China, and the presence there of some 
European types which do not occur in the Atlantic States. 

One of the most remarkable facts about the plant distri- 
bution of North America is that which has been abun- 
dantly proved both by the late Dr. Asa Gray and Sir J. D. 
Hooker, namely, the resemblance between the flora of the 
Eastern States and that of North-east Asia, whilst almost 
every characteristic form in the vegetation of the Atlantic 
States is wanting in California. There is, however, as far as 
I know, but little trace of a corresponding Japanese or 
Chinese element amongst the Lepidoptera, except the 
following :— 

Midea, Herr-Schaeff, a section of the genus Anthocharis, 
comprising four known species, of which one occurs in the 
Alleghany States, one in California, and two in China and 
Japan ; and Achalarus, of Scudder, of which two species, as 
recognized by him, are found in the Southern States east 


* P. Z. 8., 1898, pp. 3, et seq. 
+ U.S. Geological Survey, Vol. X., 1876. 


ll 


(Ist) 
of the Rocky Mountains, one extending into Mexico, 
and the other, A. lycidas, to the southern portions of New 
England. Of the remaining species, A. liliana ranges from 
the Naga Hills to Yunnan, A. bifasciatus is found in North 
China, and three or four more nearly allied species in 
Central and Western China. 

This is the more remarkable, because of the extraordinary 
resemblance between the Lepidoptera of the Rocky Mountains 
of Colorado, Montana, and Alberta, and those of Northern 
Europe and the Alps. This resemblance was, *I think, first 
pointed out by Packard in a paper on the Geographical 
Distribution of the Moths of Colorado ;* but it is not there 
shown with anything lke the force that later and _ better 
knowledge of the high-level insects of the Rocky Mountains 
afford. 

I have had personal experience of this in two collecting 
trips to the Pacific and Rocky Mountain States, and was 
astonished at the number of butterflies, identical with or very 
closely allied to those of Hurope, which occur at high levels in 
Colorado and Alberta, and the small number of species which 
belong to non-Huropean genera. I have not yet been able to 
bring together the results of these journeys, as I hope to do 
when I have explored the higher mountains of Montana, 
Wyoming, and Idaho; but I am convinced that many of 
the species which have been separated by W. H. Edwards 
and others are identical with Kuropean forms. 

A few of the most striking examples are shown among the 
specimens I have brought here, and I would call special 
attention to Lrebia magdalena, lL. tyndarus, FE. epipsodea, FE. 
sophia, and Cenonympha typhon among the Satyride; 
Argynnis chariclea and A. freya among the Nymphalide; 
Papilio machaon amongst the Papilionide ; Colias hecla, C. 
nastes, Anthocharis ausonides among the Pieridze, and Pamphila 
palamon and Hesperia centauree among the Hesperiidae. 

There can be no doubt that the subdivisions proposed by 
Dr. Bowdler Sharpe, in his admirable paper on the ‘ Zoo- 
Geographical Areas of the World, illustrating the distribution 


* Ann. Rep. U.S. Geological Survey for 18738, p. 543, et seq. 


( lxiv ) 


of birds,’* are too minute to be adopted for butterflies, and I 
should be disposed at present only to admit the following :— 

Ist. The Boreal, which would prebably include his first four 
subregions, namely the Arctic, Alaskan, Aleutian, and Cold 
Temperate. Though we know almost nothing of the 2nd and 
3rd, we cannot expect many Lepidoptera to exist there ; and 
the other two, including the higher elevations of the Rocky 
Mountains and of California, are inseparable from the Boreal 
region of Hurope and Asia, and coincide with the Polar and 
British N. Anferican floras of Hooker. 

2nd. Sharpe’s Humid Province, which he divides into the 
Appalachian and Austro-riparian subprovinces, and which 
coincides with the Great Eastern Forest Reyion of Hooker, 
of which the Southern, especially a part of the Littoral area, 
is so largely tinged with a West Indian element, both in plants 
and butterflies, but not apparently in birds, that it forms a 
transition area to the Antilles. 

8rd. Sharpe’s Arid Province, which includes his Campestrian 
and Sonoran sub-Provinces, and the southern part of the 
Prairie Region of Hooker, together with the great deserts of 
Utah, New Mexico, Arizona, and Western Texas, which is 
characterized by the comparative scarcity of most of the domi- 
nant northern genera, and shows a Mexican and Neotropical 
element by the presence of many Hesperiida, a few Evycinida, 
and other genera not found to the north, and for the most 
part represented by a few species with Mexican affinities. 

This forms a transition area to the Mexican Province of the 
Neotropical region, and being almost entirely a region of great 
aridity, and without forest, forms, in the same way as the 
African Sahara does, a barrier to the northern extension of all 
the Neotropical species which inhabit forests. 

I have mentioned these divisions in greater detail, because 
it is clear, from the remarks of Dr. Dixey in a recent paper in 
our Transactions (Trans. Ent. Soc., 1894, p. 822), that there is 


* Natural Science, Vol. III., Aug., 1893. 

+ The Distribution of the North American Flora. A lecture delivered on 
April 12, 1878, before the Royal Institution cf Great Britain. ‘ Gardeners’ 
Chronicle,’ Aug., 1878. 


(dee) 


ereat confusion in the minds of some entomologists as to the 
affinities of these regions. 

Mr. Salvin informs me that the boundary, though some- 
what indefinite, must be drawn through Northern Mexico, 
and coincides closely with the region of forest. 

Judging from the little I was able to see of the butterflies 
during the short time I spent in the Highlands of Central 
Mexico, at a season unfavourable for collecting, the northern 
element is there far weaker than among birds, two species of 
Grapta being the principal evidence of it; and is by no means so 
well marked as in the high Andes of Ecuador, Bolivia, and 
Chili, where several species of Argynnis and Colias are found, 
together with other genera resembling, if not very closely 
allied to northern forms, which do not exist in the Highlands 
of Mexico. 

Of the Andean and Chilian butterflies we know too little to 
analyse their elements. Butler’s paper,* although probably 
very incomplete, is the best on Chili; and Staudinger’s 
‘ Hochandine Lepidopteren’ + is of great interest. It seems, 
however, pretty certain that the genera do not show anything 
like the necessary amount of divergence from those of the 
Neotropical region to support the separation agreed in by three 
such eminent botanists as Hooker, Dyer, and Hemsley, of an 
Andean or Temperate South American region. 

What is most remarkable is the existence, at high eleva- 
tions in various parts of the Andes, and at sea level in South 
Chili and Patagonia, of several genera and species elsewhere 
unknown in the Neotropical region, and which are isolated 
from their congeners in North America by an cnormous area 
of country. 

Among these T'rifurcula huanaco is a remarkable species 
which occurs in the Andes of Bolivia, at 16,000 to 17,000 feet, 
and has a marvellous likeness to Baltia shawi, found at a 
similar elevation in Ladak. 

Phulia, a genus of three or four nearly allied species, also 


* List of Butterflies collected in Chili by T. Edmonds, Hsq., Trans. Ent. 
Soc. Lond., 1881, p. 449-486. 
+ Iris. Dresden. Band vii., p. 43, 1894. 


(. Teya" #) 


occurring at great elevations in the Andes and Chili, has a 
striking resemblance to Synchloe Dutleri, a species which 
accompanies Baltia in Ladak. If similar conditions of envi- 
ronment do not produce similar effects, how can these extra- 
ordinary cases of resemblance in remote and disconnected 
areas be accounted for ? 

But the Chilian Satyrid@ are the most numerous and peculiar 
of these mountain forms. Of seventy-two species recorded by 
Butler from Chili, no less than thirty-two are Satyrid, 
including several endemic genera, which resemble if they are 
not congeneric with /pinephele, Frebia, and Hipparchia, of the 
northern region. 

Argyrophenga and Aryyrophorus are two genera of most 
striking coloration, on account of the metallic silvery colour 
of the upper surface, whose only near ally is the still more 
remarkable A. antipodum of New Zealand. 

Next I may notice some species of Aryynnis which, though 
of a type different to anything in the North Temperate region, 
as A. inca and A. cytheris, have not been generically separated, 
and four or five species of Colias, a genus which occurs from 
Kcuador to the Straits of Magellan. 

Of butterflies found on the eastern side of extra tropical 
South America, I know none of Northern type except Colias 
lesbia, and a species described as (nets antarcticus, Mab., 
which I have never seen. Muryades, however, is a genus 
of very remarkable structure, haying an extraordinary 
horny excrescence on the abdomen of the female, analogous 
to that found in Parnassius, in Luehdorfia from N. EK. Asia, 
and in Hurycles from Australia, These four genera are the 
only ones in which this appendage is observed, though there 
is something like it in Pareba. I showed in my paper on 
‘ Parnassius ’ * that this organ was developed during copula- 
tion, but its function still remains unknown. 

With regard to the butterflies of the Neotropical region I 
shall say nothing, as I know too little of them, and they are 
better left to those who haye made this region a special study. 
Mr. Salvin, the highest authority on the subject, does not 
see his way to subdividing it on the butterflies without going 


* Proc. Zool. Soc. 1886, p. 6, et seq. 


€ levy) 


into elaborate statistics, which neither my time nor space 
would allow me to introduce here. But his general im- 
pression is, that when the forms of the higher Andes and 
extreme South are eliminated, no very marked divisions exist. 
He points out that the great features of the region as a 
whole, when contrasted with the rest of the world, are—the 
presence of peculiar subfamilies, such as Heliconine and 
Brassoline ; the enormous development of Ithomiine (a 
section of Danaiine) to the exclusion of Kuplaine; the 
large number of characteristic genera of Nymphaline and 
Morphine, as well as of Satyrine; the vast extent of 
Erycinida as compared with their poverty in other regions ; 
the presence of the highly peculiar Dismorphine (Pieridae), 
and wholly peculiar species of Papilionide ; the comparative 
poverty of true Lycwne, in place of which Thecla is vastly 
developed. And I would add the enormous development of 
Hesperiidae. 

I find in Watson’s paper on this family, above alluded to, 
that out of some two hundred genera recognized by him in 
the whole world, about half, including the whole of the sub- 
family Pyrrhopyyine and almost the whole of his first section 
of Hesperiine, are confined to the Neotropical, while about 
seventy occur in the Indian, and thirty only in the Ethiopian 
region, And at certain seasons and places, as I found at 
Orizaba in Mexico, in March, the species of Hesperiide out- 
number the species of all the other families together. 

Now let us turn to the Old World section of the Northern 
region, of which the butterflies are better known than those 
of any other region. Though some of the genera, such as 
Satyrus, Pararge, and Lycena, require subdivision to bring 
their generic value into accordance with those adopted in 
the Indian region, we have no great difficulty in reviewing 
them. 

I recognize four sub-divisions of this region :— 

1. The North American, which I have already dealt with. 

2. The Eurasian, which extends from Britain to N. E. 

Siberia, and coincides with that adopted by Sharpe. 
3. The Mediterraneo-Persic, or Mediterraneo-Asiatic of 
Sharpe. 


( xvi) 


4, The Eastern Temperate, which includes the Mantchurian 
and Mongolian provinces of Sharpe in part—though a 
great part of these provinces have more affinity with 
the Himalo-Chinese subregion, and are difficult to 
separate from it. Cf. Elwes on the Butterflies of 
Amurland, North China, and Japan.* 

The following genera are characteristic of the North 

Temperate region. 

Parnassius is absolutely confined to it, and has its greatest 
development in the eastern part of the Eurasian sub-region, 
but is poorly represented by two or three species in North 
America. 

Doritis, Thais, and Hypermnestra are small genera confined 
to, but hardly characteristic of, the Mediterraneo-Persiec sub- 
region. 

Luehdorfia and Sericinus are strictly confined to, but hardly: 
characteristic of, the Hast Temperate sub-region. 

Of the Pierina, Leucophasia is confined to, but not specially 
characteristic of, any part of it. 

Mesapia, Davidina, and Baltia are small genera: confined 
to the highest altitudes of Tibet. 

Colias is one of the most widely distributed and dominant 
genera over the whole region at all altitudes, especially well 
developed in Alpine regions, and extending as far north as 
any butterflies. It has developed some outlying species in 
other regions, as C. nilgherrensis in South India, C. imperialis 
in Patagonia (?), and five species in extra-tropical South 
America and the Andes. It also extends to various parts 
of Eastern and South Africa, but nowhere, as far as I know, 
occurs in a strictly tropical climate. 

Anthocharis is found almost throughout the region, but is 
not well developed in any part of it but the Mediteranean and 
North American sub-regions. 

Neophasia and Midea I have already spoken of above. 

Zegris is still more peculiar, being found only in isolated 
localities, namely, Andalusia, South Russia, Turkestan, and 
the South-Western States of North America. 


* Proc. Zool. Soc. 1881, p. 856. 


( tix) 

Of the Nymphalina, Argynnis is the largest, most wide- 
ranging, and characteristic genus of this region, occurring in 
every part of it. It has, like Colias, developed some outlying 
species which are somewhat aberrant, namely, 4. hanningtont 
in East Africa; A. niphe in India, Australia, and Java; and 
three or four species in extra-tropical South America. It 
does not, however, like Colias, extend to South Africa. 

Melitea, though it occurs in every part of the region, is 
more characteristic of Europe and Asia, most of the American 
species being somewhat aberrant. 

Timelaa is confined to North and West China, where there are 
one or two species only. 

Araschnia is found from Western Europe to Eastern Asia 
and J apan, but not in America; it is hardly characteristic of 
any particular sub-region. 

Grapta, which is perhaps only a section of the cosmopolitan 
genus Vanessa, is found throughout the region, and extends to 
the Indo-Malayan region and Mexico in isolated cases. 

Limenitis is characteristic of the whole region in a minor 
degree, but it is also well-represented in the Himalo-Chinese 
sub-region, and on this account should perhaps be excluded. 

Of the Satyrinz, Pararge is not a very homogeneous genus, 
but its more typical species are widely spread, not, however, 
extending to North America. 

Canonympha is very widespread and characteristic of every 
part of the region, except the Eastern States of North 
America, where it is absent, though everywhere present in 
the North Western States and California. 

Erebia is a very dominant and wide-ranging genus, specially 
_ characteristic of Alpine and Arctic regions. There is no 
true Erebia outside the region, but two doubtfully congeneric 
species are found in the mountains of New Zealand. 

' Satyrus is a very well developed and characteristic genus 
of the Eurasian sub-region, but is scarce or absent in the 
North American and East Temperate. 

Epinephele has almost the same distribution as Satyrus, 
but is absolutely wanting in North America. 

Melanargia is a very well marked and peculiar genus, 
characteristic of the Eurasian sub-region ; one or two species 


(six 3 


extend to Eastern Asia, but none are found in North 
America. 

(Hneis is somewhat similar in its distribution to Frebia, 
and is characteristic only of Alpine and Arctic districts, 
though some species in Europe, Asia, and America are con- 
fined to grassy steppes having an extremely cold winter 
climate. 

The Hrycinide are represented only by two small almost 
monotypic genera, Nemeobius in Europe and Polycena in 
Central Asia. 

Among the Lycenide, notwithstanding the general abun- 
dance of the family throughout every part of the region, 
hardly a single genus can be called characteristic of it, and 
only one, which is monotypic, is peculiar to it, namely, 

Lasiopsis roboris, confined to South-Western Europe. 

Cigaritis and Thestor are two small genera, almost confined 
to the Mediterraneo-Persic sub-region, and both seem to be 
of African affinity. ; 

Chrysophanus or Polyommatus is largely developed through- 
out the whole region, of which it is highly characteristic, 
though several species are found also in the Himalo-Chinese 
sub-region, 

Lycena is a very dominant genus, which, until its species 
are grouped into the genera adopted by de Niceville on the 
Butterflies of India, cannot be analysed for my present 
purpose. It is not more characteristic of the region, however, 
than of the Indo-Malayan. 

Hespertida.—In reviewing the Hesperiidae of this region, to 
see how far they may agree with the distribution of other 
families, I have adopted the genera defined by Watson. A 
careful paper by Dr. A. Speyer on the genera of the European 
Hespertide, with special reference to their occurrence in North 
America, was published in the Stettiner Ent. Zeit., 1878, pp. 
167-193, and is translated by A. J. Lintner in the Canadian 
Entomologist, 1878, pp. 121, 144, 163, et seq. 

I have not been able to accept the conclusions of this 
paper either as regards the genera adopted, or the statement 
that no species except H. centauree is common to Europe 
and North America, because the materials at Speyer’s dis- 


( these 3) 


posal were absolutely insufficient for the work he undertook. 
Watson’s arrangement, though not by any means final, is based 
on amuch wider study, and the revision which I am now 
engaged on, with the assistance of Mr. J. Edwards, of his 
genera, so far as they are found in the North Temperate and 
Indian regions, leads me to prefer it to Speyer’s. 

Carcharodus, Hubn. (Spilothyrus apud Staudinger, Pyrgus 
in part apud Speyer) has three or four species occurring in 
Europe and West Asia, but all characteristic of the Mediter- 
raneo-Persic sub-region, and absent from N. America. 

Hesperia, Fabr. (Syricthus apud Staudinger, Scelothrix 
apud Speyer) is the most numerous of all the genera in this 
region, but is cosmopolitan in distribution, though poorly repre- 
sented in N. America. Over twenty species may be distin- 
guished, and probably more exist, as Mr. J. Edwards, in study- 
ing the genital organs of the species in my collection, has 
found constant and striking differences, which he considers of 
specific value, between H, malva and what is here shown as 
H. distincta 

Thanaos (Nisoniades apud Staudinger) is widely spread 
throughout the region and in N. America, where it seems 
most abundant. 

Adopea (Hesperia in part apud Staudinger, Thymelicus pt. 
apud Speyer), of which A. thawmas is typical, according to 
Speyer, occurs sparingly throughout the region, and also in 
N. America, if 7’. garita is congeneric, as Speyer asserts. 

Erynnis, Schranck (Hesperia in part apud Staudinger, 
Pamphila part apud Speyer), is also very widely spread, and 
perhaps most of the supposed species composing it may be 
regarded as forms of H. comma. 

Pamphila, Fabr. (Carterocephalus apud Staudinger and 
Speyer), is a small genus represented in Northern Europe, 
Asia, America, and E. Tibet. 

Heteropterus (Cyclopides apud Staudinger and Speyer), a 
small genus peculiar to N. Europe and Asia. Butleria aurei- 
pennis, from Chili, seems to be a representative form. 

Gegenes, Hubn. (Hesperia in part apud Staudinger, Goniloba 
apud Speyer), is a genus of African affinity, only occurring in 
the Mediterraneo-Persic sub-region. 


() dean? 9 


There are also a few species, mostly of Indian genera—but 
in one case Achalarus only found elsewhere in the U.S.A.— 
which occur in the extreme east of the North Temperate 
region, such as Ismene aquilina, Satarupa nymphalis, Aero- 
machus inachus, and one small genus, Augiades, of which A, 
sylvanus is the type, which has its headquarters in the 
Himalo-Chinese sub-region. 

Eruiorran Recron.—Of Africa, as a whole, we have less 
knowledge than of any other region of so large an extent. 
Many collections and local lists of butterflies from various 
parts of tropical Africa have been made, but none of them 
have any pretension to being at all complete, and, as far as 
I can judge, they give no data sufficient to decide whether 
the sub-regions which are recognized for birds will hold 
good for butterflies. Cf. Kirby.* 

It seems, however, that the insects of tropical Africa are 
widely distributed from west to east, and extend on the south- 
east coast to Natal, whilst those of the more arid parts of 
North-Eastern Africa are comparatively poor in endemic genera, 
and rich only in species of Acrea and Teracolus. When we 
come to South Africa, a country so rich in its flora that it is 
considered by botanists to form a region apart, we find the 
poverty of butterflies even more striking. 

Mr. Trimen’s remarks on the subject are worth quoting.{ 
‘There can be no doubt that, with the exception of the 
‘‘ eastern coastbelt, from about the Kei river to Delagoa Bay 
‘‘ and Inhambane, South Africa is very scantily supplied with 
‘‘ butterflies.”’ ‘‘ So accustomed are we to associate butter- 
“ flies with flowers, that I well remember how much the 
‘“‘ dearth of these insects surprised and disappointed me when 
‘* first I contrasted it with the unrivalled variety and beauty of 
‘* the flora of the Cape district.” 

‘* | believe that when the Cape flora comes under investiga- 
‘« tion as regards fertilization by insect agency, it will be found 
‘‘ that a great proportion of its large and brilliant blossoms are 
ee eee to the visits of Diptera, and a good part of the 


¥* The ‘Buttartlics and Moths of Africa, a paper read before the V Ak 
Institute, June 8, 1889. 
+ South African Butterflies, vol. i., p. 42. 


( Ixxiii_) 


‘‘yemainder for those of Hymenoptera. The great number 
‘of densely hairy flower-frequenting Coleoptera in South 
‘« Africa must also play a large part in plant fertilization.” 

‘© A residence of nearly twenty-five years at Capetown 
‘enables me to state with some certainty that the species 
‘‘ inhabiting the neighbourhood do not number more than 
‘< forty-seven.” 

‘© This remarkable poverty of butterflies is rendered all the 
‘more striking from the circumstance that twenty-nine of 
‘« the species are small Lycenide (twenty-two) and Hesperiide 
‘« (seven), and the bulk of the remainder consists of sombre 
‘‘ Satyrine (ten) of medium size.” 

“It is only when we progress eastward along the belt 
‘«‘ between the first mountain range and the sea-coast that 
‘the Rhopalocerous fauna finds conditions more favourable 
‘for its development. Thus, at Knysna, where extensive 
‘ forests of large trees clothe a large area, I collected, during 
‘¢ nine months’ residence, sixty-two species.”’ 

« Across the Kei river, Colonel Bowker collected one hun- 
‘‘ dred and seventeen species, and it is only when we reach 
‘© D’Urban, on the coast of Natal, that the augmentation in 
‘their ranks is remarkable. At this spot the Rhopalocera 
‘‘ become a constant and beautiful feature of the scenery.”’ 

Mr. Trimen goes on to remark that he took in one day, 
near Natal, no less than fifty-four species, which, however, is 
less than I have taken in one day, with Mr. Salvin, in the 
Italian valleys of the Alps. 

It is not more easy to account for this poverty at the Cape 
than it is in South and West Australia, where, notwith- 
standing a rich, varied, and peculiar vegetation, we have 
extreme poverty in the Rhopalocera. The desert country 
which separates the Cape Colony from tropical Africa on the 
north, is, no doubt, a good natural boundary against the 
extension southwards of tropical forms, for which the climate 
of littoral South Africa would appear to be far better 
adapted than the coast of Korea and South Japan are for 
the tropical Indian forms of butterflies which we find there ; 
or than the coast of New England for some forms of Neo- 
tropical affinity which extend so far north; but though the 


(. dskiv >) 


desert may have prevented the migration from the north, the 
tropical coast-line on the south-east would have formed no 
obstacle to the extension of many East African tropical species 
to the Cape Colony. 

It is perhaps a question on which zoologists are not agreed, 
whether the Ethiopian region should be considered a region 
of primary value or be united with the Indian into a Paleo- 
tropical region, as is done by Hooker and in Hemsley’s alter- 
native scheme. 

Iam not prepared to accept the latter course. There are, 
no doubt, a great number of dominant genera, which are 
common to both continents; but the almost total absence 
of some of the most typical and dominant Malayan genera 
in Africa, such as Euplaa, Ornithoptera, Delias, Neptis, 
Amblypodia, and the presence in Africa of many large 
genera which are almost, or quite, absent in India, such as 
Acraa, Euryphene, Romaleosoma, Azxiocerces, Pentila, etc., 
quite outweighs the fact that some rather numerous groups, 
such as Mycalesis, Ypthima, Charaxes, Hypolimnas, Eurytela, 
are common to both. 

Of Madagascar and the Mascarene islands, which, on 
account of their many peculiar genera of mammals and 
birds, stand quite apart from Continental Africa, I cannot 
say much, as the only comprehensive account of the 
butterflies we have, by Saalmuller,* is too incomplete to 
analyse with profit. A large number of peculiar species 
occur, and some of them show Malayan rather than 
Ethiopian affinities. It has the only two species of Huplea 
which are found out of the Malay region ; also a single species 
of Hypanartia and Crenis, which are Neotropical genera. The 
beautiful moth Urania Ripheus, is also a striking instance of 
a genus found nowhere else but in South America; but, 
according to Schatz, Heteropsis is the only genus of butterflies 
peculiar to Madagascar. 

Inpo-Matayan Reaion, on East Tropica Recion or THE OLD 
Wortp.—No other region, except the Neotropical, is anything 
like so rich in peculiar, dominant, and characteristic forms 


* Lepidopteren von Madagascar, vol. i., Frankfort, 1884. 


Coley) 


as this. It may be subdivided with greater ease and conveni- 
ence than the North Temperate; but, with the exception of 
the Himalo-Chinese sub-region, its divisions do not coincide 
very well with those adopted by Mr. Sharpe or myself for birds. 
What I called the Indian sub-region, and Sharpe calls the Indo- 
Peninsular sub-region, has almost no peculiar forms among but- 
terflies, which are not equally abundant in, and characteristic 
of,a great partof the Ethiopian region. In thecultivated plains 
and jungles of Peninsular India, the dominant genera are all 
either cosmopolitan, like Papilio, Lycana; cosmotropical, like 
Terias, Catopsilia, and Junonia; or abundant in Africa like 
Teracolus, Idmais, Danais, Precis, Messaras, Atella, Ergolis, 
Hypolimnas, Charaxes, Mycalesis, Ypthima, Aphneus; or Indo- 
Malayan, like Delias, Ivias, Eronia, Neptis, Lethe, Elymnias, 
Curetis. 

Excepting the monotypic Parantirhaa, confined to the 
mountains of Travancore, and therefore belonging to the 
Indo-Malayan sub-region, I know of no genus of butterflies 
peculiar to the Peninsula of India. I will therefore strike it 
out, and include it as a province only of the Indo-Malayan 
sub-region, forming a link with the Ethiopian region. 

The north-western arid part of the Peninsula, including a 
great part of Rajputana, Sind, and the Punjab, is in butterflies, 
as in birds and plants, an eastern extension of the Mediter- 
raneo-Persic province. 

An extraordinary case of the recurrence of a species at 
widely remote spots in this sub-region is that of Anthocharis 
charlonia, which is found in various forms in the Punjab, 
Mesopotamia, Transcaspian Desert, Algeria, and in the 
Canary Isles (on Fuerteventura), though I have not seen a 
specimen from this locality. This range corresponds almost 
precisely with that of the Desert Bullfinch, Krythrospiza 
githaginea. 

The mountains of Southern India, Ceylon, and the Malabar 
coast must, however, be included in the Malayan sub-region, 
as they have a larger proportion of Malayan than of African 
forms, and many of the species found there are peculiar forms 
‘of Malayan genera. This part of the Indian region, how- 
ever, is poor in species and genera as compared with the 


( bexvi 4) 


Indo-Malay islands; while the mountains of Ceylon seem 
especially poor in peculiar forms. 

In Mr. Hampson’s list of the Butterflies of the Nilgiri 
district,* only two hundred and seventy-five species are in- 
cluded, of which Hestia and Parthenos are perhaps the only 
Malayan genera not found in Sikkim. There is only a single 
species of Nemeobiine, one of Morphine, and two of Ambly- 
podia, all of which are characteristic of the Indo-Malay region. 
Zipaetis is, perhaps, the only genus not found in the Hima- 
layas which occurs here, and Colias nilgiriensis is a unique 
instance of a Northern genus which extends so far south, 
showing that an insuperable obstacle has been opposed by the 
low and hot plains and the arid hills of the Peninsula to the 
extension southwards of either Northern genera from the 
north-west or Himalayan forms from the north-east. 

The dominant and characteristic genera of the Indo-Malay 
region, as a whole, are not very numerous, but for the most 
part have developed in the Malay archipelago a great number 
of peculiar species. 

The following are most conspicuous among them :— 

Danais, which, however, is equally well represented in 
Africa. 

Euplea, which has an immense number of species, and 
many groups extend to every part of the region. 

Cethosia, Cynthia, and Cirrhochroa, with about thirty species 
amongst them. 

Neptis, with at least ninety species, many of them confined 
to particular islands, but everywhere, except in the Pacific 
islands and extra-tropical Australia, a dominant genus. 

Athyma and Futhalia, with, probably, over one hundred 
species. 

Lethe, Mycalesis, Ypthima, and Elymnias are all dominant 
genera, but the first is better developed in the Himalo-Chinese 
sub-region, and the other three are all represented in Africa. 

The important though not numerous family of Morphine 
is characteristic of the central parts of this region, though 
entirely wanting in continental Africa. Eleven genera 


* Journ. As. Soc. Bengal, 1888. 


(xen 1) 


and about fifty species occur, of which Hyantis alone with 
one species is peculiar to the Papuan islands; as compared 
with this we have one genus only, Morpho, with about fifty 
species, in the Neotropical region. 

The subfamily Nemeobiine are almost confined to this region, 
though two genera with three species occur in the North 
Temperate region, and four or five species of Abisara occur in 
Southern Africa and Madagascar. 

All the rest of the Erycinide, about eighty-seven genera 
and one thousand species, are peculiar to the Neotropical 
region, only a few species spreading to the Southern parts of 
the United States. 

Among the Lycenide there are many genera which, as far 
as at present known, appear to be confined to this region, and 
which form a conspicuous and varied element in Indo- 
Malayana, but probably many of them will be found to 
occur also in tropical Africa when that country is better 
known. 

Of the subfamily, Liptenidw, only one, the very rare 
and curious Liphyra brassolis, occurs in various places from 
Sikkim to the Malay islands. 

Amblypodia is by far the largest and most characteristic 
group among the Lycenidw, and contains with its subgenera 
probably as many as one hundred and fifty species. 

Zephyrus is very characteristic of the Himalo-Chinese 
region to which it is confined. 

Satsuma is a Japanese and Chinese genus, which may have 
nearer allies in North America than is as yet supposed. 

Taraka, Phengaris, and Orthomiella are all monotypic 
genera characteristic of the Himalo-Chinese sub-region. 

The latter sub-region, which I defined many years ago 
when writing on the Birds of Asia (Proc. Zool. Soc. 1878, 
p. 659), has now been generally accepted as a natural and 
homogeneous one, and though the Rhopalocera do not appear 
to have developed in it, as many peculiar endemic genera as 
the birds and plants have done, yet relatively they are fully as 
rich in species. 

There are, however, many well-marked and homogeneous 
genera entirely confined to the Himalo-Chinese sub-region, 


( lxxvim )) 

and nearly all of them have a wide range, from the 
Himalayas through the mountains of Upper Burmah to the 
valley of the Yang-tse-kiang. This is evident from a com- 
parison of the list of Chinese. butterflies in Leech’s recently 
completed and most valuable work,* with the lists of Hima- 
layan and Malayan butterflies given in my paper on the 
Butterflies of Sikkim,+ where we find that, out of 477 species 
in China, excluding the Hesperiide and a few others which 
are quite abnormal in their distribution— 

145 belong to thirty-four genera which are highly charac- 
teristic of, and twenty-four of them absolutely con- 
fined to the sub-region. 

84 belong to thirty-six genera characteristic of the Indo- 
Malay region generally, of which two genera, 
Euthalia and Neptis, contain thirty-three species, 
leaving only fifty-one species to the remaining thirty- 
four genera. 

107 belong to genera of North Temperate type, but most 
of these (perhaps more than half) are confined to the 
highlands of East Tibet, which more properly form 
part of the North Temperate region. 

45 belong to genera such as Danais, Ypthima, Melanitis, 
Charaxes, Junonia, etc., which are common to the 
Ethiopian region, and 

96 belong to cosmopolitan genera. 

Out of the first category eight genera, all monotypic, are 
peculiar to China, namely, Mandarinia, Callarge, Palao- 
nympha, Isodelma, Timelaa, Amblopala, Davidina, and 
Sericinus. Luehdorfia is found elsewhere only in Japan. 

Besides these, the following genera only are not found in 
the Himalayas :— 

Acropthalmia, which occurs in the Philippines. 
Midea inthe United States. 

Polycena in Central Asia. 

Melanargia in Europe and West Asia. 
Phengaris in the Naga Hills. 


* The Butterflies of China, Japan, and Corea. London, 1892-94, by 
J. H. Leech. 


+ Elwes on the Butterflies of Sikkim, Trans. Ent. Soc. Lond., 1888, p. 269. 


Cj lisse) 


The number of genera found in the Himalayas, but not 
as yet in China, is greater ; but it must be remembered that 
a large part of the very centre of this region, including the 
head waters of the Irrawaddy, and the greater part of the 
south-western provinces of China, are as yet quite unex- 
plored. Most of them are Lycenide of Malayan type. 
Among them, perhaps, the following may be mentioned, 
and of these, the first two only are confined to the Himalayas: 
Anadebis, Orinoma, Zipatis, Elymnias, Penthima, Neurosiyma, 
Allotinus, Miletus, Nacadaba, Horaga, Cheritrella, Pontia, 
Prioneris, Hebomoia and Eronia. 

Of typical Malayan genera, not found in the Himalo- 
Chinese sub-region, the following are the principal :— 

Hestia, Idaopsis, Erites, Zeuxidia, Xanthotenia, Prothoé, 
Terinos, Rhinopalpa, and several genera of Lycanida, 

The boundaries of this sub-region are extremely indefinite, 
as they depend greatly upon altitude, the region being essen- 
tially one of considerable elevation. So far as we know at 
present its outlying districts in the high mountains of the 
Malay Peninsula, Borneo, Sumatra, and Java, are not as well 
distinguished by peculiar mountain forms of butterflies as they 
are by birds. Formosa, of which we know almost nothing, is 
an island which would probably now repay the exploration of 
an entomologist better than any other in the Hastern Seas, 
and the recent discovery of a species of Hestia, so far north as 
the Liukiu islands, shows that there is much of interest to be 
expected there. 

The division between the Indo-Malayan and Austro- 
Malayan regions laid down by Wallace, and which has since 
been generally accepted by zoologists as ‘ Wallace’s line,’ is 
much less marked in butterflies than in birds or mammals, and 
the few traces of an Australian element, which are found to 
the west of this line, are comparatively unimportant. 

In Java, Mr. Snellen, who is now engaged in a work on 
the Lepidoptera of this island, informs me that though there 
is a difference between the butterflies of East and West Java 
yet it is not strongly marked. The greater part of the species 
are spread over the whole island. Papilio Van de Polli, Pap, 
Now, Prioneris philomene, Delias Peribaa, Cethosia Lamarckii, 


(“lex 


and CU. Leschenaultii, are instances of butterflies which are 
common in East but not found in West Java. Tenaris is a 
typical Austro-Malay genus found in Java, but not I think 
in Sumatra or Borneo. In the island of Nias we find a 
purely Indo-Malayan fauna, to which perhaps the most 
notable exception is Miletus celisparsus, Butt., a Lycenid of 
Papuan type. 

In the Philippine group, of whose butterflies we have now 
an excellent account by Semper,* I find only a very few 
genera which are not characteristic of the whole Indo-Malay 
sub-region ; Acropthalmia, Ptychandra, and Hypothecla being 
exceptions. Thysonotis and Phrissura are perhaps the only 
forms showing Australian affinities. 

In Sumba and Sambawa, Doherty’s explorations have 
shown an overwhelming proportion of Indian as compared 
to Australian types, as I hope to show more clearly when I 
have time to work out his collections. 

It is only when we come to Celebes and the Moluccas that 
any marked change becomes evident, and even here the 
difference is nothing like so great as among birds and 
mammals. 

Wallace, in his classical paper on ‘‘ The Papilionide of the 
Malayan Region,’’+ states that Celebes is the most peculiar of 
all the islands in the Archipelago, for though it has a smaller 
number than either Borneo or Java, no less than eighteen of 
its Papilionide are peculiar, whilst Java, Sumatra, Borneo, 
and Malacca, with forty-five species, have only twenty-one, or 
less than half peculiar. 

Wallace shows that of the nineteen groups into which he 
divides the Malayan Papilionide, only three, viz., the 
Priamus, Ulysses, and Erecthus. groups, are peculiar to the 
Austro-Malayan sub-region, and when we come to inquire 
what are the elements in this sub-region which connect 
it with the Australian continent we hardly find any. 
Though far too little is yet known of the insects or even of 
the butterflies of Papua and its surrounding islands, to 


* Semper, Die Schmetterlinge der Philippinischen Inseln. 1892. 
+ Trans. Linn. Soc., vol. xxv (1865). 


( <issca) 


enable us to analyze their constituent elements, yet I am 
strongly of opinion that from my present point of view the 
Austro-Malayan sub-region must be attached to India and 
not to Australia, and in this case I am in accordance with 
the views of botanists. My knowledge of this sub-region 
is, however, too slight, and my collection entirely inadequate 
to illustrate it this evening, and I therefore propose to leave 
the question in abeyance for the present. 

We now arrive at a very critical point in the investigation, 
which is to see what is the relative value of the Australian 
fauna, as compared with those of other regions, and 
what is its position as a primary division. I must confess 
that Iam unable, from personal knowledge, to answer this 
question, as, after taking a general survey of the Australian 
Lepidoptera, I was inclined to say that, on their evidence 
alone, the Australian region could hardly be said to exist. 
But on applying to Mr. E. Meyrick, whose personal knowledge 
of the region is extensive, and whose critical accuracy in 
describing and classifying the Lepidoptera of Australia makes 
his opinion most valuable, I received the following remarks, 
which I quote in extenso, He writes as follows :— 

*‘Confining myself to the Lepidoptera, New Zealand 
‘and Australia cannot possibly be classed together. They 
‘“‘have hardly any species in common; the few they 
‘‘ have are undoubtedly either (1) recent immigrants from 
‘one to the other, or (2) insects of world-wide range, as 
‘* Heliothis armigera, or (8) artificially introduced. Their 
‘* genera are equally distinct ; no genus is prominent in both, 
‘‘ except such as are also prominent throughout all regions. 
‘« In fact, on a consideration of general affinity, New Zealand 
‘ig really much more related to England that to Australia. 

‘*(2) Australia is quite as distinct from the other continents 
** in Lepidoptera as it is inmammalsand plants. The follow- 
‘‘ing are some instances. A peculiar family of Pyralidina 
“(the Tineolide), consisting at present of four monotypic 
‘« genera, is confined to Australia (it is more ancient than any 
‘‘ Knglish family, being the ancestor of the Pterophoride) ; 
“this may be compared with the Monotremata, which consist 
‘* of three monotypic genera. 


ce 
ce 


ce 


a3 


( lexan) 


‘*The Ccophorida are enormously developed in Australia 
(I have nearly nine hundred species), whilst those known 
from the rest of the world do not probably reach a hundred 
and fifty. This is a parallel case to the excessive develop- 
ment of the Myracea, and of such genera as Acacia. The 
Geometrid family Monoctenide are known to have about one 
hundred species in Australia, and about thirty in the rest of 
the world. The Xyloryctida, a group of Tineina, are perhaps 
not entitled to rank as a family, but only as a group of the 
Gelechiada. However, in any case they form a natural and 
distinct group, including already over one hundred species 
in Australia, whilst only one or two others have been found 
elsewhere. I could give many other cases of large 
genera, or groups of genera, mainly or entirely confined 
to Australa. 

‘‘A more peculiar case (not, I think, paralleled in other 
forms of life) is that of Crambus, which is largely developed 
throughout the whole world except Australia and the adja- 
cent islands, though Australia is eminently suited for it. 
There are two species in Australia, one an insect which 
occurs throughout most of Europe, Asia, and Africa, the 
other a Malayan and South Pacific species, so that both 
must be supposed to have immigrated; in New Zealand 
there are about thirty endemic species. 

‘**(3) New Zealand is remarkable for its poverty; only. 
about six hundred species are known, and there cannot be a 
great many more, but probably some have already become 
extinct. Most of the groups usually classed as Bombycina 
and Sphingina are altogether absent. The larger genera 
are all such as are common to all regions. Where, how- 
ever, there is a marked affinity in respect of any more 
peculiar genera, the affinity is undoubtedly with South 
America. There is also a strong archaic element; various 
small genera of one or two species, belonging to ancient 
types, probably once generally prevalent, but now every- 
where nearly extinct. Noticeable genera are— 

“« Diptychophora, fourteen species (several others in South 


‘* America, one in Australia); Scoparia, sixty species, more 
‘‘ than are known from the rest of the world, though the 


CIs) 


‘genus is cosmopolitan; Heliostibes, three species, only 
‘‘ known otherwise from South America. The Micropteryyide, 
‘‘ eight species, are only known otherwise from Europe, North 
“and South America. Many large families, such as the 
‘* Phycitida, are wholly, or almost wholly, absent. 

(4) The distinctness of the Australian fauna is to some 
“extent masked by the great influx of Malayan species 
‘into the northern portions, a comparatively recent phe- 
** nomenon. 

(5) So far as I am. acquainted with the fauna of the 
‘‘ South Pacific islands, it seems very fragmentary and miscel- 
‘«¢ Janeous, more Malayan than anything else, but it is very 
** inadequately known as yet.” 

This evidence is very striking, and if confirmed by other 
groups of insects would justify the retention of Australia as one 
of the primary regions ; but it must not be forgotten that our 
knowledge of these obscure groups in most parts of the world, 
and especially in the tropics, is, one may say, infinitesimal, 
and though I am quite as ready to attach weight to evidence 
drawn from obscure and little-known insects, as from the 
more striking and better-known groups, yet the classification 
on which the whole theory rests must be sound and well 
determined, and I fail to see how any system of classification 
can be more than provisional which is not based on a much 
greater amount of material than I think Mr. Meyrick at 
present possesses. 

Lord Walsingham, however, whose special knowledge of 
the Tincina of the world gives his opinion great weight, and 
to whom I submitted Mr. Meyrick’s remarks, agrees with him, 
Provisionally accepting, therefore, Australia as one of the 
primary divisions, can it be subdivided ? 

New Zealand must, according to Meyrick, stand apart, 
though if it is not a province of the Australian region it can 
hardly be included in any other. 

Its very few species of butterflies comprise one or two wide- 
ranging forms which are probably immigrants. The principal 
others are 

Two endemic species of Vanessa. 


( ixxxig,} 


Four species of a genus resembling and nearly allied to 
Chrysophanus, all of which are endemic. 

Two species of a genus allied to Frebia, peculiar to the 
mountain ranges of the South Island. 

And one very striking butterfly, Argyrophenga antipodum, 
whose nearest allies are found in the mountains of Chili. — 

Papua and its islands, with the extreme northern pro- 
montory of Australia, ought, on the evidence of its butterflies, 
to form with the Moluccas and Celebes a province of the 
Indo-Malay region ; and the Pacific Islands do not, as far as 
we know, possess anything like the number of peculiar or 
characteristic forms of insects to justify their separation, 
the majority of the Lepidoptera known from them belonging 
to very wide-ranging and Malayan genera. Thus we find, 
that even if the Australian region is to rank with the other 
great divisions of the earth, it must do so for the present 
rather on negative than on positive evidence. 

I must now conclude what I fear to some of my hearers 
has been a somewhat tedious address, but which I hope may 
give rise to an interesting discussion at our next meeting. 
The specimens illustrating it will be, by the kind permission 
of Sir W. Flower, deposited in the British Museum for a 
time, in order that they may be more closely examined by 
anyone who wishes to do so. If I shall have succeeded in 
leading some of our Fellows to pay more attention to a most 
interesting branch of our subject, I shall feel, however, that 
your time has not been wasted. The subject of distribution 
is one which requires sound systematic work as a basis, but 
cannot be profitably studied by systematists alone without 
some knowledge of the geographical, meteorological, and 
geological features of the earth. All of these are essential 
aids to the proper consideration and appreciation of the 
numerous difficulties we meet with in trying to explain the 
facts which are presented to us, and a knowledge of all of 
them will add immensely to the interest and pleasure of 
travel, which I look on as an essential part of the education 
‘of a modern naturalist. . 


(. Ixuxy., 


INDEX. 


Notr.—Where the name only of the Species or Genus is mentioned, the 
description will be found on the page referred to. 


The Arabic figures refer to the pages of the ‘ Transactions’; the Roman 
numerals to the pages of the ‘ Proceedings.’ 


The New Genera and Species, and those which have been redescribed, will 
be found in detail, but certain of the longer papers are arranged generi- 
cally under their headings. 


Page Page 

GENERAL SUBJECTS .......... _Ixxxv HEMIPTERA ......... oantecttess lxxxvili 

APHANIPTERA....se.ssseeeees. XXXVI FELYMENOPTE RA. cc scocccteciess lxxxix 

ARACHNIDAsee...ciscenessseneee Ixxxvi | LEPIDOPTERA ..........00 0.0000 Ixxxix 

COLEOPTERA ...ceesesccessesee = [XXXVI NEUROPTERA  ...cccsvccencseoes xevi 

DIPTERA scshiccssisacciesssceses LXEXVIIE ORTHOPTERA cccsscsevssnsentine xevi 
oo 


GENERAL SUBJECTS. 
Annual Meeting, xlv. 


Antenne of Insects, some remarks on, xxxiil. 

Diloboderus abderus, Eucranium arachnoides, and Megathopa violacea, 
notes on, xx. 

Emydia cribrum, notes on, xxix. 

Formica smaragdina, notes on nests of, xxxiii. 

Gandaritis flavata, notes on, xiii. 

Lecanium prunastri, notes on, xviii. 

Mechanical decoy and spring net, iv. 

Pierine, a large number of genera and species alluded to, 249-334. 

President’s Address, 1. 

Publication of Transactions, Part I., 1894, discussion concerning date of, xxiii. 

Sarcopsylla, notes on a species of, xxi. 

Selenia lunaria, notes on, xix. 

Special General Meetings, xvii, xxiv. 

Temperature experiments in 1893, on several species of Vanessa and other 
Lepidoptera, ix. 

Temperature-variation as bearing on theories of heredity, Mr. Merrifield’s 
experiments in, x. 

The Westwood bequest, vi. 

White ants, notes on a collection of, formed by Mr. G. D. Haviland in 
Singapore, viii. 

Zygena exulans, notes on, xxvi. 


( < lxxxvi ) 


APHANIPTERA. 
Hystricopsylla talpe (?), from Kent, exhibited, xxviii. 
Sarcopsylla, species of, from Ningpo, exhibited, xxv. 


ARACHNIDA. 
Atypus piceus, nest of, from Hastings, exhibited, ii. 


COLEOPTERA. 


Acanthotomicus, n. g., 89. A. spinosus, n. s., 90. 

Adelium nodulosum, n. s., 404.—tasmanicum, n. s., 408. 

Anthaxia vitidula, from the New Forest, exhibited, xxix. 

Arrhenoptera exilis, n. s., 372.—pygmea, n. s., 371. 

Athous rhombeus, from the New Forest, exhibited, xxix. 

Australia and Tasmania, Tenebrionide collected in, by Mr. J. J. Walker, 
during the voyage of H.M.S. ‘ Penguin’ (G. C. Champion); species 
belonging to the following genera described or alluded to :—Adelium, 
401. — Alphitobius, 379.—Amarygmus, 407. — Arrhenoplita, 371.— 
Brycopia, 405.—Caanthus, n. g., 378.—Cediomorpha, 358.—Cedius, 
357.-—Cestrinus, 354.—Chalcopterus, 406.—Coripera, 401.—Corticeus, 
369.—Cotulades, 352.—Crypticus, 362.—Decialma, 3 5.—Dinoria, 
404.—Diphyrrhynchus, 366.—Dipsaconia, 377.—Docualis, 352.— Ectyche, 
397.—Edylius, n. g., 353.—Hlascus, 352.—EHnnebeopsis, n. g., 375. 
—Ennebeus, 375.—Ganyme, 377.—Heleus, 882 —Heterocheira, 366. 
—Hymea, 397.—Hyocis, 368.—Hypaulax, 389.—Lepispilus, 393.-— 
Lyphia, 370.—Melytra, 397.—Menephilus, 390.—Meneristes, 392.— 
Micrectyche, 399.—Nyctozoilus, 388.—Omolipus, 398.—Opatrum, 358. 
—Palorus, 369.—Paratovicum, n. g., 380.—Phennis, n. g., 399.— 
Phycosecis, 364.—Platydema, 373.—Prionotus, 358.—Promethis, 392. 
—Pseudocedius, 357.—Pteroheleus, 382.—Saragus, 385.—Scymena, 
385.— Synuypetes, 384.—Titena, 396.—Toxicum, 380.—Trachyscelis, 
365.—Tribolium, 379.—Ulodes, 377. 

Brycopia tuberculifera, n. s., 405. 

Caanthus, n. g., 878. C. gibbicollis, n. s., 378. 

Cardiophorus equiseti, from Devonshire, exhibited, xviii. 

Cestrinus brevis, n. s., 856. 

Chreonoma basalis, n. s., 487. 

Cicindela mashuna, n. s., 452. 

Coccotrypes advena, n. s., 100.—perditor, n. s., 99. 

Corticeus australis, n. s., 369. 

Cosmema dolosa, n. s., 452. 

Cosmoderes australis, n. s., 369. 


Crossotarsus chapuwisi, n. s., 129.—concinnus, name substituted for chapuist, 
Blandford, 578.—contaminatus, n. s., 1381.—niponicus, n. s., 180. 
Cryphalus ewiguus, n. s., 82. 
Crypticus submaculatus, n. s., 362. 
Decialma erichsoni, n. s., 395. 
Diapus aculeatus, n. s., 139. 
Dicranocephalus adamsi and D. dabryi, from China, exhibited, viii. 
Diloboderus abderus, etc., from La Plata, exhibited, xx. 


(¢ Ixexyii: ») 


Diphyrrhyncus apicalis, n. s., 368.—ellipticus, n. s., 367. 

Dryocetes, new species described :—afinis, 93.—dinoderoides, 97.—luteus, 
94.—mestus, 95.—nobilis, 95.—pilosus, 92. 

Ectyche cerulea, n. s., 397. P 

Edylius, n. g., 353. EB. canescens, n. 8.5 854 

Eidophelus minutus, n. s., 88. 

Ennebeopsis, n. g., 375. E. pruimosus, n. s., 376. 

Ennebeus australis, n. s., 375. 

Genyocerus adustipennis, and other Japanese Coleoptera alluded to, 53-55. 

Glenea cantor, n. s., 488. 

Eelops swperbus, and a large number of other Corsican Coleoptera, alluded 
to, 229, 

Hylastes ambiguus, n. SAY (- 

Hylesinus, new species described :—cingulatus, 67.—costatus, 63.—lati- 
collis, 65.—nobilis, 64.—scutulatus, 67.—tristis, 66. 

Hyocis subparallela, n. s., 363. 

Hyorrhyncus, n. g., 58. H. lewisi, n. s., 60. 

Hypothenemus expers, un. 8.4 85.—peritus, n. s., 84. 

Japan, Rhynchophorous Coleoptera of (W. H. Blandford), species belonging 
to the following families and genera described or alluded to :— 
Acanthotomicus, n. g., 89.—Coccotrypes, 98.—Cosmoderes, 85.— 
Crossotarsus, 129.—Cryphalus, 82.—Crypturgus, 82.—Diapus, 139. 
—Dryocetes, 91.—Eidophelus, 88.—Genyocerus, 141.—Hylastes, 56.— 
Hylesinus, 63, Hyorrhyncus, n. g., 58._-Hypothenemus, 83.—Myelo- 
philus, 58.—Phleosinus, 68.—Pityophthorus, 87.—Platypus, 133.— 
Platypini, 127.—Polygraphus, 74.—Scolyto-platypini, 126.—Scolyto- 
platypus, 126.—Scolytus, 77.—Spherotrypes, n. g., 61.—Tomicus, 89. 
—Trypodendron, 124,—Xyleborus, 100. 

Longicorn Coleoptera obtained by Mr. J. J. Walker, during the voyage of 
H.M.S. ‘Penguin’ (C. J. Gahan); species belonging to the following 
families and genera described or alluded to:—Hgosoma, 481.— 
Miolesthes, 482.—Agapanthia, 485.—Apriona, 484,—Aristobia, 483.— 
Aromia, 482.—Batocera, 483.—Blepepheus, 433.—Cerambycide, 481. 
—Cerambyx, 482.—Ceresium, 482.—Chreonoma, 486.—Clytantus, 482. 
—Clytus, 482.—Coptops, 484.—Cyriocrates, 483.—Dere, 482.—Dorca- 
dida, 483.— Epepeotes, 483. — Erythrus, 482. — Eurypoda, 481.— 
Ezocentrus, 485.—Gerania, 484.—Glenea, 485.—Gnatholea, 482.— 
Gnoma, 484.—Hestima, 484.—Himantocera, 484.—Lamiide, 483.— 
Linda, 486.—Melanauster, 483.—Monohammus, 483.—Oberea, 486.— 
Olenecamptus, 484.—Philus, 481.—Phytecia, 485.—Polyzonus, 482.— 
Prionide, 481.—Psacothea, 483.—Pterolophia, 484.—Pyresthes, 482. 
—Rhytidodera, 482.—Ropica, 485.—Rosenbergia, 484.—Sternoplistes, 
483.—Sybra, 485.—Tetraglenes, 485.—Xenolea,483.—Xystrocera, 481. 

Lyphia tasmanica, n. s., 370. 

Megacephala regalis, alluded to, 447. 

Menephilus ruficornis, n. s., 391. 

Myrmecoptera, new species described :—angusticollis, 448.—formosa, n.s., 
451.—invicta, 452.—mashuna, 449.—marshalli, 150. 

Nyctozovlus sexcostatus, n. s., 388. 


( Ixxxvili ) 


Oberea walkeri, n. s., 487. 

Omolipus cyaneipennis, n. s., 394. 

Opatrum, new species described :—dispersum, 3861.—torridum, 359.— 
vagabundum, 361.—walkeri, 360. 

Paratovicum, n. g., 380. P. iridescens, n. s., 881. 

Phennis nigricornis, n. s., 382. 

Phleosinus, new species described :—dubius, 70.—minutus, 71.—pulchellus, 
69 —rudis, 73.—seriatus, 72. 

Pityophthorus jucundus, n. s., 87. 

Platydema bicinctum, nu. s., 873.—deplanatum, un. s., 374. 

Platypus, new species described :—calamus, 187.—hamatus, 188.—lewisi, 
134.—modestus, 133.—severini, 1386. 

Polygraphus miser, n. s., 76.—oblongus, n. s., 75.—provimus, n. s., 75. 

Pteroheleus nigricornis, n. s., 382. 

Rhina barbirostris, from Honduras, exhibited, xviii. 

Ropica squamosa, n.s., 486. 

Saragus bicarinatus, n. s., 385.—intricatus, n. s., 386. 

Scolyto-platypus shogun, n. s., 126. 

Scolytus, new species described :—agnatus, 78.—aratus, 79.—claviger, 80.— 
esuriens, 77.—frontalis, 79. 

Spherotypes, n.g., 61. 8S. globulus, n. s., 63.—pila, n.s., 62. 

Synapius filiformis, and other Coleoptera, exhibited, xxxii. 

Titena tasmanica, n. s., 396. 

Trypodendron pubipenne, n. s., 125.—sordidum, n. 8., 577. 

Velleius dilatatus, from the New Forest, exhibited, xxix. 

Xyleborus, new species described :—adumbratus, 115.—amputatus, 575.— 
apicalis, 105.—aquilus, 109.—attenwatus, 114.—bicolor, 118.—con- 
cisus, 107.—cucullatus, 121.—defensus, 118.—exesus, 119.—galeatus, 
123.—germanus, 106.—interjectus, 576.—lewisi, 104.—minutus, 116.— 
muticus, 112.—mutilatus, 103.—orbatus, 128.—praevius, 110.—schau- 
fussi, 117.—seriatus, 111. 


DIPTERA. 


Mallota eristaloides, from the New Forest, exhibited, xxv, 


HEMIPTERA. 


Bolbonota auro-sericea, n. s., 417.—rufo-notata, n. s., 416. 
Dorydium westwoodi, notes on, 411. 

Enchotype concinna, n. s., 419. 

Helopeltis theivora, description of male, 31. 

Hemiptera from Corsica, alluded to, 243. 

Hille ecuadorensis, n.s., 420. 

Hoplophora unicolor, n. s., 419. 

TLecanium prunastri, exhibited, xviii. 

Lucilla intermedia, n.s., 421. 

Orthocephalus championi, n. s., 247. 

Paradarnoides, n. g., 422.—P. ignipes, nu. s., 424.—severini, n. s., 423. 
Potnia concinna, n. 8., 419.—perobtusa, n. s., 419. 


( izemix”) 


Serinetha augur, exhibited, xv. 

Telamona prealta, n.s., 420.—ruficarinata, n. s., 421. 
Triquetra obtusa, n. 8., 417. 

Tropidoscyta transiens, n. s., 415. 


HYMENOPTERA. 


Allea, 504. A. contracta, 505. 

Alysia, 507. A. atra, 5138.—fuscipennis, 511.—incongrua, 516.—loripes, 
511.—lucicola, 516.—mandibulator, 513.—manducator, 508.—rufidens, 
510.—similis, 512.—sophia, 515.—soror, 510.—tipulae, 514.—trun- 
cator, 514. 

Aphaerete, 522. A. cephalotes, 528. 

Clasmodon, 500. C. apterus, 500. 

Cratospila, 506. C. circe, 506. 

Formica smaragdina, alluded to, xxxiii. 

Idiasta, 521. I. maritima, 521.—nephele, 522. 

Panerema, 501. P. inops, 501. = 

Pentaplewra, 519. P. angustula, 520.—fuliginosa, 520.—pumilio, 520. 

Phenocarpa, 524. FP. conspurcator, 531.—eugenia, 5380.—eunice, 532.— 
flavipes, 533.—galatea, 530.—livida, 533.—maria, 529.—nina, 582.— 
picinervis, 527.—pullata, 526.—punctigera, 526.—pratellae, 531.—rufi« 
ceps, 528.—tabida, 527. 

Pimpla bridgmani, n. s., from Devonshire, exhibited, xxxii. 

Praon absinthii, n. s., from Devonshire, exhibited, xxxii. 

Synerasis, 502. S. fucicola, 502.—halidati, 503. 

Tanycarpa, 517. T. ancilla, 518.—gracilicornis, 517.—rufinotata, 518. 

Trachyusa, 508. T. aurora, 504. 


LEPIDOPTERA. 


Addaea probolopis, n. 8., 477.—syndesma, n. 8., 478. 

Adela satrapodes, n. s., 26. 

Agrotis alexandriensis, n. s., 37.—nili, n. 8., 38. 

Alexandria, Lepidoptera from the neighbourhood of (G. T. Bethune-Baker), 
species belonging to the following genera described or alluded to :— 
Acherontia, 34.—Acidalia, 44.—Agdistis, 50.—Agrotis, 37.—Alucita, 50. 
— Ancylolomia, 48.—Anthocharis, 33.—Anthophilodes, 47.—Arrarde, 
44.—Bombyz, 36.—Botys, 47.—Bryophila, 47.—Caradrina, 42.—Cero- 
cala, 43.—Chilo, 48.—Cidaria, 45.—Cletthara, 35.—Colias, 33.—Cossus, 
36.—Crambus, 48.—Crocidosema, 50.—Cucullia, 42.—Danais, 34.— 
Deilephila, 35.—Deiopeia, 35.—Deudorix, 33.—Duponchelia, 48.— 
Earias, 35.—Emnatheudes, 50.—Erastria, 43.—Eriopus, 41.—Eromene, 
49.—Eupithecia, 45.—Eurychreon, 47.—Fidonia, 45.—Grammodes, 43. 
—Hapsifera, 50.— Heliophobus, 40. — Heliothis, 42.—Hellula, 47.— 
Hesperia, 34.—Hypena, 44.—Hypotia, 4}.—Leucanitis, 43.—Lycena, 
33.—Macaria, 44.—Mamestra, 39.—Murgarodes, 47.—Megasoma, 37.— 
Melissoblaptes, 50.—Metasia, 47.—Miana, 41.—Mnesivena, 45.—Nenio- 
ria, 44.—Nola, 35.—Nymphula, 48.—Ocnogyna, 36.—Odontia, 47.— 
Orgyia, 36.—Orobena, 47.—Pericyma, 43.—Plusia, 42.—Prodenia, 42. 


CG x6%) 


Pseudophia, 43.—Psyche, 36.—Scardia, 50.—Schenobius, 48.—Sesamia, 
42.—Spintherops, 44.—Sphinw, 34.—Sterrha, 45.—Synclera, 47.—Talis, 
47.—Thalpochares, 43.—Thestor, 33.—Tinea, 50.—Vanessa, 34. 

Ammatho disticha, n. s.,2.—epieantha, n.:s., 2. 

Anacampsis scutata, n. s., 14.—wollastoni, n. s., 545. 

Analthes pyrrhocoma, n. s., 460. : 

Anisoctena, n. g., 455. A. synomotis, n. s., 456. 

Anisodes dotilla, n. s., 179.—heydena, n. s., 178. 

Antiolopha, n. g., 25. A. hemiconis, n.s., 25. 

Anydraula wanthobathra, n. s., 474. 

Aporandria, n. g., 174. 

Arctia menthrastt, vars. of, from North Scotland, exhibited, xxvii. 

Arctoscelis, n. g.,22. A. epinyctia, n. s., 28. 

Argynnis aglaia, var. charlotta, from the New Forest, exhibited, iv., xvi. 

Aripana glaucias,n. s., 459. 

Arrarde parva,n. s., 44. ' 

Balanotis euryptera, n. s., 476. 

Blastobasis decolorella, 548.—desertarum vayr., radiata, n. v., 550.—laver- 
nella, n. s., 547.—lignea, n. s., 550.—lignea, var., adustella, n. v., 551. 

Boarmia ratotaria, n. s., 216. 

Botys atlanticum, n. s., 584.—maderensis, n. 8., 584, 

Bryophila fraudatricula var., pallida, n. v., 37. 

Cacamoda, n. g., 172. 

Cacoecia podana, exhibited, xix. 

Callocasta, n. g., 218. , 

Caradrina mediterraneae, n. s., 42. 

Carige zalska, n. s., 185. 

Cataclysta mesorphna, n. s., 10.—nyctopis, n. s., 475. 

Chelonia caja, var. from Abbots’ Wood, exhibited, xvi. 

Chilo dubia, n. s., 48. 

Cidaria bareconia, n. s., 189. 

Cistidia luoifera, n. s., 211. 

Cladodes arotrea, n. s., 15. 

Cletthara littora, n. s., 33. 

Comocritis, n. g., 24. C. olympia, n.s., 24. 

Cossus L-nigrwm, n.s., 36. 

Crambus afra, n. s., 49.—alerandriensis, n. s., 48.—atlanticus, 585. 

Crasigenes, n. g., 476. C. nicrospila, n. s., 477. 

Crocallis heydena, n. s., 208. 

Crypstiptya, n. g., 463. 

Dasyscopa, n. g., 464. D. homogenes, n. s., 464. 

Deilinia luteifrons, n. s., 193. 

Dithalma mecysma, n. s., 179. 

Donacaula chlorosema, n. s., 11. 

Ectropis planaria, n. s., 22i1.—simplaria, n. s., £21. 

Emydia cribrim, exhibited, xxix. 

Endographis, n. g., 464. HE. acrochlora, n.s., 455. 

Endotricha orthotis, n. s., 476. 

Epiplema ocusta, n. s., 165.—sreapa, n. s., 166. 

Hpistetus, n. g., 552. EH. divisus, n. s., 552.—pica, n. 8., 558. 


( xc) 


Eriocephala calthella, larva of, alluded to, 335. 

Eucrostes kafebera, n. 8., 177. 

Eupterote avesta, n. s., 157.—calandra, n. s., 157. 

Fascellina cyanifera, n. s., 201. 

Fidonia megiaria var., obscwraria, n. v., 40. 

Gastropacha quercifolia, larve of, exhibited, xvi. 

Geoglada, n. g., 184. G. helicola, n. s., 184. 

Gracilaria chalcanthes, n.s., 25. 

Gyadroma, n. g., 220. 

Heliophobus marsdeni, n. s., 40. 

Herculia psamathopis, n. s., 11. 

Hierangela, n. g.,14. H. erythrogramma, n. s., 15. 

Homosaces, n. g., 20. H. anthocoma, n. 8., 21. 

Hyalobathra dialychna, n. s., 7. 

Hydrocampa oxygona, n. s., 470. 

Hypochrosis venusa, n. s., 197. 

Hypochrysops scintillans, female described, 143. 

Hyponomeuta bakeri, n. s., 542. 

Hypotia bilinea, n. s., 46. 

Hypulia, n. g., 205. 

Icterodes fasciata, n. s., 211. 

Iotaphora, n. g., 168. 

Ischnoscopa, n. g., 464. I. chalcozona, n.8., 464. 

Khasia Hills, Lepidoptera from the (Col. Swinhoe) ; species belonging to 
the following families and genera described or alluded to :—Abragas, 
212.—Abrazinez, 211.—Absala, 170.—Acherontia, 146.—Acherontiine, 
146.—Achrosis, 198. — Acidalia, 180. — Acidaliide, 178.—Acosmeryz, 
147.—Actias, 152.—Agathia, 173.—Alex, 184.—Amana, 164.—Ambly- 
chia, 215.—Ambulycine, 147.—Ambulyz, 147.— Amraica, 213.— 
Anisodes, 178.—Anonychia, 222.—Antherea, 153.—Antheua, 160.— 
Anthyperythra, 204.—Anthyria, 191.—Anticyra, 160.—Apha, 156.— 
Apocalypsis, 151.—Aporandria, n. g., 174.—Ardonis, 186.—Arichanna, 
222.—Aropteris, 163.—Ascotis, 221.—Astacauda, 220.—Asthena, 192.— 
Attacus, 152.—Azurea, 164.—Bapta, 198.—Baradesa, 158.—Bardanes, 
J87.—Berta, 177.—Besaia, 161.—Boarmia, 216.—Boarmiide, 192.— 
Bombycide, 154.—Brahmexa, 154.—Brahmzide, 154.—Buzura, 214.— 
Caberodes, 195.—Cacamanda, 172.—Calcula, 212.—Callocasta, n. g., 
218.—Calymnia, 147.—Cambogia, 187.—Carige, 185.—Cassyma, 210.— 
Catascia, 218.—Catoria, 218.—Ceira, 160.—Celeraena, 168.—Ceph- 
nodes, 152.—Cerura, 160.—Chexrocampa, 148.—Chzrocampine, 147.— 
Chatamla, 164.—Chlorodontoptera, 171.—Chorodna, 214.—Chrysocras- 
peda, 180.—Cidaria, 189.—Cistidia, 211.—Collix, 185.—Corymica, 206. 
Craspedia, 181. — Cricula, 154. — Crisiala, 214, — Crocallis, :02.— 
Crocota, 204.—Cryptoloba, 188.—Cymatophoride, 162.—Dalima, 197. 
Daphnis, 150.—Darisa, 219.—Decetia, 164.—Defoa, 182.—Deilinia, 
193.—Deiliniine, 193.—Dirades, 166.—Dithalma, 179.—Dreata, 155.— 
Dudusa, 158.—Dysethia, 187.—Ectropis, 221.—Eilicrinia, 199.—Elibia, 
150.—Elphos, 215.—Ennominez, 197.—Epiceia, 162.—Epicoptide, 
162.—Epione, 205.—Epiplema, 165.—Epiplemide, 164.—Epipristis, 
171. — Epirrhoe, 188. — Eptsophthalma, 172. — Erebomorpha, 214,— 


Fs e bs 
(xem 4 

Erithrolophus, 180.—Eubyja, 213.—Euchloris, 174.—Eucrostes, 176.— 
Eumelia, 167.—Eupithecia, 186.—Eupterote, 156.—Eupterotidx, 155.— 
Euschema, 169.—Eustroma, 190.—Fascellina, 141.—Fentonia, 141.— 
Gangarides, 155.—Gandaritis, 191.—Ganisa, 156.—Gareus, 202.— 
Gargetta, 158.—Gaurena, 162.—Gelasma, 175.—Geoglada, n. g., 184.— 
Geometridx, 169.—Gonanticlea, 184.—Gurelca, 151.—Gyadroma, n. g., 
220.—Habrosyne, 162.—Hemerophila, 219.—Herochroma, 171.—Hetero- 
locha, 203.—Heteromiza, 199.—Heterostegane, 195.—Hirasa, 219.— 
Hyalinetta, 202.—Hypereschra, 160.—Hyperythra, 204.—Hypochrosis, 
197.—Hyposidra, 202.—Hypulia, n. g., 205.—Hyria, 182.—Ichthyura, 
161.—Icterodes, 211.—Iotaphora, n. g., 168.—Krananda, 210.—Laci- 
niodes, n. g., 188.—Langlia, 146.—Larentiide, 184.—Lassaba, 219.— 
Lissolica, 172.—Lobophora, 187.—Loepa, 153.—Lophopteryx, 161.— 
Lovochila, 174.—Luwiaria, 206.—Lygranoa, 187.—Macaria, 207.— 
Macariine, 206.—Macroglossa, 151.—Macroglossine, 151.—Mavzates, 
172.—Medasina, 217.—Megalochlora, 174.—Menocteniidx, 1(6.—Meso- 
trophe, 178.—Metaschalis, 161.—Metrocampa, 203.—Micronia, 163.— 
Micronidia, 163.—Microniidx,163.—Micronissa, 195.—Milionia, 213.— 
Myrteta, 194.—Nagadara, 199.—Naza, 168.—Nemoria, 176.—Nephele, 
151.—Netria, 161.—Nisaga, 157.—Noreia, 167.—Notodontide, 158.— 
Nyctalemon, 162.—Ocinara, 154.—C@nospila, 176.—Ophthalmodes, 215. 
—Orbasia, n. g., 222.—Orsonoba, 201.—Orthostyxidx, 167.—Orudiza, 
.166.—Ozola, 167.—Pachyodes, 169.—Palirisa, 155.—Paradecetia, 164. 
—Paralophia, 190.—Paricterodes, 211.—Peratophyga, n. g., 204.— 
Percnia, 212.—Pericallia, 205.—Perixera, 178.—Petelia, 194.—Phalera, 
158. — Phibalapteryx, 186. — Photoscotosia, 191. — Plemyria, 188. — 
Plutodes, 195. — Polynesia, 188.—Polyploca, 162.—Pomasia, 191.— 
Porstca, 159. — Prionia, 198. — Problepsis, 183. — Protoparce, 150. — 
Pseudangerona, 218.—Pseuderythra, n. g., 204.—Pseudodolbina, 151.— 
Pseudojana, 156.—Pseudomicronia, 163.—Pseudopterna, 170.—Pseudo- 
sphinz, 150.— Psilaleis, 222. — Psilocambogia, 192.— Psyra, 202. — 
Pydna, 159.—Racotis, .217.—Rambara, 168.—Remodes, 185.—Rhodo- 
metra, 192.—Rodostrophia, 181.—Rhopalopsyche, 151.—Runeca, 183.— 
Rutterellona, 220.—Salassa, 153.—Sarcinodes, 166.—Sataspes, 152.— 
Saturnia, 153.—Saturniidae, 152.—Scardamia, 204 —Scotopteryx, 22°. 
—Scotosia, 191.—Sirinopteryzx, 193.—Smerinthine, 146.—Somatina, 182. 
—Somera, 160.—Spatalia, 161.—Sphingide, 146.—Sphingine, 150.— 
Spilopera, 199.—Stegania, 195.—Streptopteron, 180.—Strophidia, 163. 
—Synegia, 205.—Synegiodes, 180.—Tagora, 155.—Tanaorhinus, .73.— 
Tarsolepis, 158.—Tasta, 194.—Tephrina, 210.—Thalera, 176.—Thale- 
rura, n. g., 175.—Theretra, 150.—Thinopteryx, 193.—Thyatira, 162.— 
Timandra, 177.—Tosaura, 169.—Trirachopoda, 178.—Trygodes, 184.— 
Turnaca, 159.—Uraniidx, 162.—Urapteroides, 163.—Urapterygine, 192. 
—Urapteryx, 192.—Xandrames, 217.—Xanthorrhoé, 188.—Xeropteryx, 
193.—Zamarada, 210.—Zeheba, 210.—Ziridava, 186. 

Taciniodes, n. g., 128. 

Laverna rebeli, n. s., 554. 

Lecithocera pachyntis, n. s., 17. 

Iimacodes testudo, larva of, alluded to, 345. 

Lissolica, n. g., 172. 


(- ‘xen *) 


Loxocorys, n. g., 6. 

Lycaena adonis and corydon, vars. exhibited, xv.—egyptiaca, n. s., 33. 

Macaria oliva, n. s., 207. 

Madeira Islands, Pterophoride, Tortricide, and Tineide of (Lord Walsing- 
ham); species belonging to the following families and genera described 
or alluded to:—Alucita, 539.—Anacampsis, 544.—Asychna, 555.— 
Bactra, 540.—Bedellia, 542.—Blastobasine, 546.—Blastobasis, 547.— 
Bryotropha, 544.—Carpocapsa, 540.—Carposina, 540.—Chauliodus, 554. 
—Choreutis, 545.—Conchylidine, 540.—Cosmopterygine, 555.—Cos- 
mopteryx, 555.—Depressaria, 546.—Depressariane, 546.—Endrosis, 546. 
—Rpistetus, n. g., 552.—Gelechiane, 543.—Glyphipterygine, 545.— 
Glyphipteryx, 545.—Gracilaria, 555.—Gracilarianez, 555.—Grapholitha 
540.—Grapholithine, 540.—Hypatima, 554.—Hyponomeuta, 542.— 
Hyponomeutine, 542.—Laverna, 554.—Luvernine, 554.—Lita, 5438.— 
Lithocolletis, 555.—Luonetiane, 555.—Ccophora, 546.—Q@cophorine, 
546.—G@nophila, 542.—Ozxyptilus, 538.—Paedisca, 540.—Phyllobrostis, 
555.—Plutella, 543.—Plutelline, 543.—Platyptilia, 538.—Pterophoride, 
538.—Siganorosis, 546.—Simethis, 545.—Sitotroga, 544,—Steganop- 
tycha, 541.—Tinea, 541.—Tineidx, 541.—Tineine, 641.—Tineola, 542. 
—Tortricide, 5389.—Tortriv, 589.—Trichophaga, 541. 

Madeira Islands, Pyralids, Crambidew, and Phycide of (G. T. Bethune- 
Baker) ; species belonging to the following genera described or alluded 
to: — Acheraea, 586.— Aglossa, 581.— Asopia, 581.—Botys, 583.— 
Crambus, 585.— Diasemia, 585. — Ephestia, 583. — Eromene, 586.— 
Galeria, 586. — Hellula, 583. — Margarodes, 585. — Myelois, 586. — 
Nomophila, 585.—Scoparia, 581. 

Malay Archipelago, Pyralidina from the (KE. Meyrick); species belonging to 
the following families and genera described or alluded to :—Addza, 477. 
—tholivx, 461.—Agathodes, 467.—Agrotera, 461.—Analthes, 460.— 
Anisoctena, n. g., 455.—Anydraula, 474.—Archernis, 467.—Aripana, 
459.—Balanotis, 476.—Caprinia, 468.—Cataclysta, 475.—Cnaphalo- 
crocis, 467.—Cometura, 468.—Conogethes, 459.—Coptobasis, 457.— 
Crambide, 480.—Crambus, 480.—Crasigenes, n. g., 476.—Crypsiptya, 
n. g., 463.—Cymoriza, 470.—Dasyscopa, n. g., 464.—Diasemia, 467.— 
Diathrausta, 467.—Dichocrocis, 461.—Diplopleustis, 477.—Endogra- 
phis, n. g., 454.—Endotricha, 476.—Ephestia, 479.—Galleriadxe, 479.— 
Hydriris, 468.—Hydrocampa, 470.—Ischnoscopa, nu. g., 464.—Isocen- 
tris, 464.—Margaronia, 456.—Margarosticha, 474.—Meroctena, 460.— 
Merotoma, n. g., 460.—Metasia, 465.—Metasiodes, n. g., 465.—Meto- 
portha, n. g., 463.—Mivophyla, 470.—Myelois, 479.—Nausinoé, 468.— 
Nistra, 468.—Nosophora, 459.—Notarcha, 461.—Nymphula, 470.— 
Oligernis, n. g., 470.—Orphnophanes, 469.—Pagyda, 467.—Perisyn- 
trocha, n. g., 469.—Phlyctaenia, 463.—Phycitide, 479.—Pinacia, 4 8. 
—Platamonia, 468.—Pleonectusa, 468.—Pleuroptya, 463.—Prasinoxena, 
n. g., 479.—Pterygisus, 470.—Ptilaeola, 467.—Pyralidide, 476.— 
Pyralis, 476.—Pyrausta, 463.—Pyraustide, 455.—Rehimena, 459.— 
Sagariphora, n. g., 460.—Salebria, 479.—Saroscelis, n. g., 461.— 
Schoenobius, 475.—Scirpophaga, 475.—Siculodes, 478.—Siculodide, 
477.—Siriocauta, 468.—Stegothyris, 467.—Stenia, 465.—Symmoracma, 


( :xeiv }) 


n. g., 469.—Syngamia, 467.—Syntomodora, un. g., 467.—Tabidia, 467. 
—Tylostega, n. g., 457. 

Mamestra abbas, n. s., 40.—afra, n. s., 39. 

Margaronia callizina, n. s., 5.—sphenoscoma, n. s., 406. 

Melasina ochrocoma, n. s., 26. 

Merotoma, n. g., 460. 

Metasia hilarodes, n. s., 465.—zanclogramma, n. s., 8. 

Metasiodes achromantias, n. s., 465.—calliophis, n. s., 466.—heliaula, n. 8., 
8.—eazalis, 466.—tholeropa, n.s., 466. 

Metoportha, n. g., 463. 

Miana trilinea, n. s., 41. 

Diltochrista callinoma, n. s., 3.—celidopa, n. s., 3.—eccentropis, n.8., 3.— 
geodetis, 4, 

Mnesivena bella, n. s., 46.—quadripunctata, n. s., 45. 

Myelois cinerella, 586. 

Nausinoé conchylia, n. s., 468. 

Nosophora scotaula, n. s., 459. 

Notarcha compsogramma, n. s., 461.—homomorpha, n. s., 462.—iophanes, 
n. s., 462.—pyranthes, n. s., 462. 

Nymphula fusco-marginata, n. s., 48. 

Ocinara cyproba, n. s., 154. 

(Enetus mirabilis, from Queensland, exhibited, xv. 

Oligernis, new species of, described :—argyropis, 473.—basilissa, 474.— 
ceratucha, 471.—endophthalma, 470.—endosaris, 472.—idiotis, 472.— 
leucochrysa, 471.—melanodes, 473.—orthoteles, 472. 

Omiodes erythrias, n. s., 457.—hiracia, n. s., 457. 

Onebala, genus redescribed, 15. 

Orbasia, n. g., 222. 

Ornithoptera andromache, from Borneo, exhibited, xv.—O. paradisea, from 
New Guinea, exhibited, xv.—O. trojana, from Palawan, exhibited, xv. 

Papilio homerus, description of larva and pupa of, 409.—telearchus, female 
of, from Cherra Punji, exhibited, xxxi. 

Pararge megaera, and other Lepidoptera, abnormal forms of, exhibited, xxxi. 

Pelosia tetrasema, n. s., 2. 

Peratophyga, n. g., 204. 

Percnia coryneta, n. s., 213. 

Periacma, n. g., 21. P. chlorodesma, n. 8., 22.—ferialis, n. s., 21.— 
orthiodes, n. s., 22. 

Perisyntrocha, n. g., 469. 

Phaeosaces torrida, n. s., 20. 

Placoptila, n. g., 23. P. electrica, n.s., 23. 

Platamonia ptochura, n. s., 468. 

Plusia moneta, from Christchurch, Hants, exhibited, ii. 

Plutodes lamisca, n. s., 196. 

Prasinowena, n. g., 479. P. hemisema, n. s., 480.—monospila, n. s., 480. 

Prosaris, n. g., 12.—P. pernigralis, n.s., 12. 

Pseuderythra, n. g., 204. 

Pseudophia benenotata, female described, 43. 

Psyche hampsoni, n. s., 36. 

Ptochorytis, n. g.,19. P. eremopa, n. 8., 19. 


( -xer®) 


Pydna bela, n. s., 159. 

Rambara efila, n. s., 168. 

Ruttellerona, n. g., 220. 

Sagariphora, n. g., 460. 8. heliochlena, n.s., 461. 

Salassa megastica, n. s., 158. 

Saridoscelis, n. g., 28. 8S. sphenias, n. s., 28. 

Saroscelis, n. g., 461. 

Scardia mediterranex, n. s., 30.—tholerodes, 27. 

Schenobius celidias, n. s., 475. 

Scirpophaga mnesidora, n. s., 475. 

Scoparia decorella, 581.—scoriella, 582.—stenota, 582.—wollastont, n. 8., 
583. 

Siculodes ancylosema, n.s., 478.—chalcosidera, n. s., 478.—platyntis, n. 8., 
479.—mochlias, n. s., 479. 

Somatina rosacea, n. s., 182. 

Spilopera hepaticata, n. s., 200. 

Stegania peralba, n. s., 195. 

Steganoptycha pygmaeana, from Norfolk, exhibited, xx. 

Sterrha sacraria, from Mudeford, exhibited, xxviii. 

Symmoracina, n. g., 469. S. spodinopa, n. s., 469. 

Synegia gopterana, n. s., 206. 

Syntomodora, n. g., 467. 

Tabidia craterodes, n. s., 457. 

Talis afra, n. s., 47. 

Tephrina atmala, n. s., 210. 

Temperature experiments on species of Vanessa and other Lepidoptera- 
(F. Merrifield and F. A. Dixey) ; species alluded to :—Araschnia levana, 
427.—Cidaria silaceata, 427.—Pararge egeria, 426.—Vanessa antiopa, 
436.—atalanta, 432, 439.—C-album, 4385, 441.—i0, 485, 440.—poly- 
chloros, 429, 441. 

Thalerura, n. g., 175. 

Thisizima, genus redescribed, 26. ; 

Tinea liomorpha, nu. s., 28.—platyntis, n. s., 28. 

Tipha, genus redescribed, 18.—T. helioclina, n. s., 19. 

Torodora, n. g., 16. T. ancylota,n. s., 17.—characteris, n. s., 16.—paral- 
lactis, n. s., 17. 

Trichernis, n. g., 20. T. centrias, n. s., 20. 

Turnaca delinewena, n. s., 159. 

Tylostega, n. g., 457. T. chrysanthes, n.s., 458.—mesodora, m. s., 458.— 
photias, n. s., 458.—schematias, n. s., 458. 

Upper Burma, Lepidoptera from (E. Meyrick); species belonging to the fol- 
lowing families and genera described or alluded to :—Acharana, 7.— 
Adela, 26.—Agrotera, 6.—Ammatha, 2.—Anacanvpsis, 14.—Ancylomia, 
14.—Antiolopha, n. g., 25—Aquita, 14.—Arctiade, 2.—Arctoscelis, 
n. g., 25.—Bizone, 2.—Blabophanes, 27.—Blastoblaside, 22.—Boc- 
choris, 9.—Botyodes, 6.—Canthelea, 18.—Cataclysta, 10.—Cerase, 24. 
—COladodes, 15.—Cnaphalocrocis, 9.—Comocritis, n. g., 24.—Cono- 
gethes, 6.—Cosmopteryx, 23.—Crambostenia, 11.—Critonia, 13.— 
Crobylophora, 29.—Deiopeia, 4.—Depressaridx, 20.—Dera, 13.—Dia- 
semia, 9.—Dolichostica, 9.—Donacaula, 11.—Elachistide, 13.—Etiella, 


( xtvi .) 


13.—Filodes, 10.—Galleriade, 14.—Calechiade, 14.—Gracilaria, 25.— 
Gracilaride, 25.—Hellula, 8.—Herculia, 11.—Heteroglypta, 138.— 
Hierangela, n. g., 14.—Homosaces, n. g., 20.—Hyalobathra, 7.— 
Hyboloma, 13.—Hypomeutide, 24.—Isocentris, 7.—Lamoria, 14,— 
Lebena, 4,—Lecithocera, 17.—Lozocorys, n. g., 6.—Margaronia, 5.— 
Melasina, 26.—Metasia, 8.—Metasiodes, n. g., 8.—Miltochrista, 3.— 
Mixophyla, 11.—Myelois, 13.—Nacoleia, 9.—Notarcha, 6.—Nymphula, 
10. — Heophoride, 22.— Omiodes, 6.— Onebala, 15.—Oxacne, 4.— 
Pagyda, 10.—Pelosia, 2.—Periacma, n. g., 21.—Phzxosaces, 20.— 
Phissama, 4.—Phlyctenia, 6.— Phycitide, 13.—Piesmopoda, 18.—Pla- 
coptila, n. g., 23.—Plutella, 24.—Plutellide, 24.—Prosaris, n. g., 12.— 
Psecadia, 22.—Ptochorytis, n. g., 19.—Ptychopseustis, 14.—Pyralide, 
11.—Pyralis, 12.—Pyrausta, 7.—Pyraustide, 5.—Rajendra, 4,— 
Rhodophza, 138.—Sameodes, 9.—Saridoscelis, 98,—Scardia, 27.— 
Schenobius, 11.—Scirpophaga, 11.—Siculodes, 18.—Siriocauta, 10.— 
Spilosoma, 4.—Stegothyris, 9.—Striglina, 18.—Surratha, 14.—Thisi- 
zima, 26.—Tinea, 28.—Tineide, 26.—Tipha, 18.—Titanio, 7.—Toro- 
dora, n. g., 16.—Trichernis, n. g., 20.—Xylorictide, 19.—Zalithia, 
n. 2, 18: 
Zalithia, n. g., 18.—uranopis, n.s., 18. 
Zygena filipendule, &c., pupx of, alluded to, 349. 


NEUROPTERA. 

Calotermes flavicollis, alluded to, vii. 

Leuctra, 559. 

Nemoura, 560. N. avicularis, n. s., 562.—borealis, n. s., 571.—cambrica, 
563.—cinerea, 571.—dubitans, n. s., 565.—inconspicua, 572.—lacustris 
570.—lateralis, 564.—maracandica, 567.—marginata, 569.—meyeri, 568. 
—ornata, 567.—praecow, n. s., 566.—variegata, 561. 

Panorpa gibberosa, alluded to, 498, xxiii. 

Polycentropus excisus, n. s., 491. 

Raphidia rhodopica, n. s., 489. 

Rhyacophila obtusa, n. s., 492. 

Taeniopteryx, 559. 

Termes lucifugus, alluded to, vii. 

Tinodes wnidentata, n. s., 491. 


ORTHOPTERA. 
Acridium septemfasciatum, exhibited, xxxii. 
Decticus albifrons, alluded to, xxix. 
Periplaneta australasiae, from Brighton, exhibited, xxxii. 


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